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Lifespan, longevity, and ageing

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Maximum longevity: 20.1 years (wild)
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Biology

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During the breeding season, pairs form which may last for a number of seasons. The spectacular courtship display involves the male flying in circles at a great height over the breeding area before falling close to the ground, performing elaborate sequences of tumbles. Occasionally the female may join him, and the pair lock talons and tumble through the air together. As the season progresses the male may be seen dropping food into the female's talons in mid-air (4). The nest, which may measure up to 80cm in diameter, is constructed on the ground with grass, reeds and sticks by the female. 3-8 eggs are laid from late April, and both parents contribute to feeding the chicks (2). Hunting occurs at a height of 2-6 meters above ground; when prey is located the bird suddenly drops down with its talons outstretched. Food taken includes small mammals and birds, carrion and sometimes insects, frogs and fish (2).
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Conservation

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The marsh harrier will benefit from the protection and creation of reedbed habitat (2). Egg collecting, nest disturbance and illegal poisoning must be controlled. Prevention of scrub encroachment and the maintenance of suitable water levels will maintain appropriate habitat and deter foxes, the main predator. As this harrier nests in close association with intensive agricultural land, levels of pesticides must be monitored. Research is being conducted on the breeding biology, food and hunting behaviour of the species in Britain and the Netherlands (3).
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Description

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Marsh harriers are recognised as harriers by their long, narrow tails, long legs and wings held in a 'V' in flight. This species is the largest of the harriers and has broad, rounded wings. Members of the genus Circus possess an owl-like 'ruff' of facial feathers that disguises very large ear openings, which enable the bird to detect prey species by the noises they produce. The plumage colour is variable, but mainly brown. Females and immature individuals have creamy coloured crowns and throats; mature males have pale-grey wings with a dark brown body and wings. Calls include a lapwing-like mewing and a 'chattering' alarm call (2).
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Habitat

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Marsh harriers are usually associated with wetlands, as the common name would suggest (2). Breeding occurs amongst reeds or sedges, usually in large reed beds. However there are records of some individuals breeding in cereal fields, or within small areas of reeds located amongst arable land or saltmarsh. Hunting generally occurs over agricultural land or open habitat containing aquatic vegetation (3).
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Range

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A scarce summer visitor to Britain, marsh harriers are also found in Europe, the Middle East, Central and northern Asia, and parts of Africa. In Britain the species mainly breeds in East Anglia, but can be seen on the south and east coast as far north as Scotland whilst it is migrating. It can also occasionally be seen in Wales and Ireland. Small numbers of individuals over-winter in East Anglia, Kent and south Wales (3). There are records of marsh harriers in Britain that date from the Iron Age, about 3000 years ago (2).
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Status

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Listed under Annex 1 of EC Birds Directive; Appendix II of the Bern Convention and the Birds of Conservation Concern Red List. Protected in the UK under Schedule 1 of the Wildlife and Countryside Act 1981 (3).
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Threats

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The species underwent a decline in numbers in the 18th and 19th centuries due to the effects of habitat drainage and persecution for predating upon game species. The population declined to just a single pair between 1959 and 1971, mainly as a result of egg-shell thinning caused by organochloride pesticides such as DDT. After 1972 the population began to recover; in 1994 there were 129 pairs, and numbers have continued to increase. Current threats include the deliberate disturbance of nesting sites, egg collecting, illegal poisoning and predation by foxes. The small size of reedbed habitats may result in an increased impact of disturbance (3).
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Status in Egypt

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Regular passage visitor, winter visitor and resident breeder?

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Brief Summary

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Marsh harriers in the air are easiest to recognize by their long wings held in a shallow V as they fly low along the open space. They make large circles as they watch for prey, mostly gliding with occasional powerful wingbeats. Sometimes, they force water fowl to take cover under water so often that they become exhausted and are easily grabbed with their long legs. Although they also nest on the Wadden Islands, they go south for the winter, staying anywhere between the Flevopolders and Africa.
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Zoologger: The only cross-dressing bird of prey

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Marsh harriers are one of only two bird species – and the only bird of prey – where some of the males mimic females...

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Western marsh harrier

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The western marsh harrier (Circus aeruginosus) is a large harrier, a bird of prey from temperate and subtropical western Eurasia and adjacent Africa. It is also known as the Eurasian marsh harrier. Formerly, a number of relatives were included in C. aeruginosus, which was then known as "marsh harrier". The related taxa are now generally considered to be separate species: the eastern marsh harrier (C. spilonotus), the Papuan harrier (C. spilothorax) of eastern Asia and the Wallacea, the swamp harrier (C. approximans) of Australasia and the Madagascar marsh harrier (C. maillardi) of the western Indian Ocean islands.

The western marsh harrier is often divided into two subspecies, the widely migratory C. a. aeruginosus which is found across most of its range, and C. a. harterti which is resident all-year in north-west Africa.

Taxonomy

The western marsh harrier was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae under the binomial name Falco aeruginosus.[2] Linnaeus specified the locality as Europe but restricted this to Sweden in 1761.[3][4] The western marsh harrier is now placed in the genus Circus that was introduced by the French naturalist Bernard Germain de Lacépède in 1799.[5][6] The genus name Circus is derived from the Ancient Greek kirkos, referring to a bird of prey named for its circling flight (kirkos, "circle"), probably the hen harrier. The specific aeruginosus is Latin for "rusty".[7]

Two subspecies are recognised:[6]

  • C. a. aeruginosus (Linnaeus, 1758) – breeding in Europe and the Middle East to central Asia south Europe, Africa, south Asia; wintering in south Europe, Africa and south Asia
  • C. a. harterti Zedlitz, 1914 – resident in northwest Africa

Description

A fairly pale adult female (note brown remiges and yellow eye) winters near Hodal (Faridabad district, Haryana, India)
Adult male (front), juvenile (behind) and adult female (back), illustration from 1899

The western marsh harrier is 48 to 56 cm (19 to 22 in) in length, has a wingspan of 115 to 130 cm (45 to 51 in) and a weight of 400 to 650 g (14 to 23 oz) in males and 550 to 800 g (19 to 28 oz) in females.[8] It is a large, bulky harrier, larger than other European harriers, with fairly broad wings, and is sexually dimorphic. The male's plumage is mostly a cryptic reddish-brown with lighter yellowish streaks, which are particularly prominent on the breast. The head and shoulders are mostly pale greyish-yellowish. The rectrices and the secondary and tertiary remiges are pure grey, the latter contrasting with the brown forewing and the black primary remiges at the wingtips. The upperside and underside of the wing look similar, though the brown is lighter on the underwing. Whether from the side or below, flying males appear characteristically three-colored brown-grey-black.[9] The legs, feet, irides and the cere of the black bill are yellow.[10]

The female is almost entirely chocolate-brown. The top of the head, the throat and the shoulders have of a conspicuously lighter yellowish colour; this can be clearly delimited and very contrasting, or (particularly in worn plumage) be more washed-out, resembling the male's head colours. But the eye area of the female is always darker, making the light eye stand out, while the male's head is altogether not very contrastingly coloured and the female lacks the grey wing-patch and tail. Juveniles are similar to females, but usually have less yellow, particularly on the shoulders.[9]

There is a rare hypermelanic morph with largely dark plumage. It is most often found in the east of the species' range. Juveniles of this morph may look entirely black in flight.

Distribution and habitat

The male is characterised by the very clear chestnut brown mantle and the grey secondaries and black outer primaries

This species has a wide breeding range from Europe and northwestern Africa to Central Asia and the northern parts of the Middle East. It breeds in almost every country of Europe but is absent from mountainous regions and subarctic Scandinavia. It is rare but increasing in Great Britain where it has spread as far as eastern Scotland.[11] In the Middle East there are populations in Turkey, Iraq, and Iran, while in Central Asia the range extends eastwards as far as north-west China, Mongolia, and the Lake Baikal region of Siberia.

Most populations of the western marsh harrier are migratory or dispersive. Some birds winter in milder regions of southern and western Europe, while others migrate to the Sahel, Nile basin and Great Lakes region in Africa, or to Arabia, the Indian subcontinent, and Myanmar. The all-year resident subspecies harterti inhabits Morocco, Algeria, and Tunisia.

Vagrants have reached Iceland, the Azores, Malaysia, and Sumatra. The first documented (but unconfirmed) record for the Americas was one bird reportedly photographed on 4 December 1994 at Chincoteague National Wildlife Refuge in Accomack County, Virginia. Subsequently, there were confirmed records from Guadeloupe (winter of 2002/2003), from Laguna Cartagena National Wildlife Refuge in Puerto Rico (early 2004 and January/February 2006)[12][13][14] and in Bermuda (December 2015).[15]

The female usually is identifiable by the rather dark plumage except the creamy crown, nape, and throat

Like the other marsh harriers, it is strongly associated with wetland areas, especially those rich in common reed (Phragmites australis). It can also be met with in a variety of other open habitats, such as farmland and grassland, particularly where these border marshland. It is a territorial bird in the breeding season, and even in winter it seems less social than other harriers, which often gather in large flocks.[16] But this is probably simply due to habitat preferences, as the marsh harriers are completely allopatric while several of C. aeruginosus grassland and steppe relatives winter in the same regions and assemble at food sources such as locust outbreaks. Still, in Keoladeo National Park of Rajasthan (India) around 100 Eurasian marsh harriers are observed to roost together each November/December; they assemble in tall grassland dominated by Desmostachya bipinnata and vetiver (Chrysopogon zizanioides), but where this is too disturbed by human activity they will use floating carpets of common water hyacinth (Eichhornia crassipes) instead – the choice of such roost sites may be to give early warning of predators, which will conspicuously rustle through the plants if they try to sneak upon the resting birds[17]

Behaviour and ecology

Breeding

Egg, Collection Museum Wiesbaden, Germany

The start of the breeding season varies from mid-March to early May. Western marsh harrier males often pair with two and occasionally three females. Pair bonds usually last for a single breeding season, but some pairs remain together for several years.

The ground nest is made of sticks, reeds and grasses. It is usually built in a reedbed, but the species will also nest in arable fields. There are between three and eight eggs in a normal clutch. The eggs are oval in shape and white in colour, with a bluish or greenish tinge when recently laid. The eggs are incubated for 31–38 days and the young birds fledge after 30–40 days.[18]

Food and feeding

It hunts in typical harrier fashion, gliding low over flat open ground on its search for prey, with its wings held in a shallow V-shape and often with dangling legs. It feeds on small mammals, small birds, insects, reptiles, and frogs.[19][20]

Status and conservation

Wintering female hunting near Kolkata (West Bengal, India)
Western marsh harrier in Estonia
Circus aeruginosus by Jos Zwarts

The western marsh harrier declined in many areas between the 19th and the late 20th centuries due to persecution, habitat destruction and excessive pesticide use. It is now a protected species in many countries. In Great Britain, the population was likely extinct by the end of the 19th century. A single pair in Horsey, Norfolk bred in 1911, and by 2006, the Rare Breeding Birds Panel had recorded at least 265 females rearing 453 young. It made a comeback in Ireland as well, where it had become extinct in 1918.[21]

It still faces a number of threats, including the shooting of birds migrating through the Mediterranean region. They are vulnerable to disturbance during the breeding season and also liable to lead shot poisoning. Still, the threats to this bird have been largely averted and it is today classified as Species of Least Concern by the IUCN.[1]

References

  1. ^ a b BirdLife International (2019). "Circus aeruginosus". IUCN Red List of Threatened Species. 2019: e.T22695344A155490248. Retrieved 6 November 2021.
  2. ^ Linnaeus, Carl (1758). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis (in Latin). Vol. 1 (10th ed.). Holmiae (Stockholm): Laurentii Salvii. p. 91.
  3. ^ Linnaeus, Carl (1761). Fauna svecica : sistens animalia sveciae regni: mammalia, aves, amphibia, pisces, insecta, vermes, distributa per classes & ordines, genera & species, cum differentiis specierum, synonymis auctorum, nominibus incolarum, locis natalium, descriptionibus insectorum (in Latin) (2nd ed.). Stockholmiae: Sumtu & Literis Direct. Laurentii Salvii. p. 23.
  4. ^ Mayr, Ernst; Cottrell, G. William, eds. (1979). Check-list of Birds of the World. Volume 1 (2nd ed.). Cambridge, Massachusetts: Museum of Comparative Zoology. p. 319.
  5. ^ Lacépède, Bernard Germain de (1799). "Tableau des sous-classes, divisions, sous-division, ordres et genres des oiseux". Discours d'ouverture et de clôture du cours d'histoire naturelle (in French). Paris: Plassan. p. 4. Page numbering starts at one for each of the three sections.
  6. ^ a b Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (January 2021). "Hoatzin, New World vultures, Secretarybird, raptors". IOC World Bird List Version 11.1. International Ornithologists' Union. Retrieved 10 June 2021.
  7. ^ Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 34, 109. ISBN 978-1-4081-2501-4.
  8. ^ Cramp 1980, pp. 105, 115.
  9. ^ a b Cramp 1980, p. 105.
  10. ^ Cramp 1980, p. 115.
  11. ^ Holling, Mark (September 2010). Rare Birds Breeding Panel. "Rare breeding birds in the United Kingdom in 2008" (PDF). British Birds. 103: 482–538.
  12. ^ American Ornithologists' Union 2000
  13. ^ Banks et al. 2005
  14. ^ Merkord, Rodríguez & Faaborg 2006
  15. ^ "Birders Spot New Species at Annual Count". The Royal Gazette. The Royal Gazette.
  16. ^ Clarke et al. 1998
  17. ^ Verma 2002
  18. ^ "Avibirds.com". Avibirds.com. Retrieved 2021-08-31.
  19. ^ "Marsh harrier". RSPB's Birds by Name. Royal Society for the Protection of Birds.
  20. ^ "Marsh Harrier". BBC Nature.
  21. ^ Parkin, David; Knox, Alan (2008). The Status of Birds in Britain and Ireland. A & C Black. p. 112. ISBN 9781408125007. Retrieved 2018-06-07.
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Western marsh harrier: Brief Summary

provided by wikipedia EN

The western marsh harrier (Circus aeruginosus) is a large harrier, a bird of prey from temperate and subtropical western Eurasia and adjacent Africa. It is also known as the Eurasian marsh harrier. Formerly, a number of relatives were included in C. aeruginosus, which was then known as "marsh harrier". The related taxa are now generally considered to be separate species: the eastern marsh harrier (C. spilonotus), the Papuan harrier (C. spilothorax) of eastern Asia and the Wallacea, the swamp harrier (C. approximans) of Australasia and the Madagascar marsh harrier (C. maillardi) of the western Indian Ocean islands.

The western marsh harrier is often divided into two subspecies, the widely migratory C. a. aeruginosus which is found across most of its range, and C. a. harterti which is resident all-year in north-west Africa.

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