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Sand Sturgeon

Scaphirhynchus platorynchus (Rafinesque 1820)

Behavior

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Shovelnose sturgeons search for food by manipulating their large barbels which have taste buds and electrorecptors. These barbels hang below the mouth and feel the bottom as the sturgeon swims. This allows them to sense when food or other obstacles are below.

Communication Channels: tactile

Perception Channels: tactile ; chemical

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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William Fink, University of Michigan-Ann Arbor
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Associations

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Other fishes, such as catfishes and burbots, probably eat the young shovelnose sturgeons. The main predator of adult shovelnose sturgeons is man. Presently, about 25 tons of shovelnose sturgeon are harvested annually. Sixty percent come from the Mississippi River upstream of St. Louis, Missouri. They are harvested in late fall and early winter for both meat and highly valued roe. The alteration of large rivers, and construction of locks and dams for navigational purposes has contributed significantly to the decline of the species by blocking access to ancestral spawning grounds, and by eliminating its required habitat.

Known Predators:

  • burbots (Lota lota)
  • yellow bullheads (Ameiurus natalis)
  • brown bullheads (Ameiurus nebulosus)
  • black bullheads (Ameiurus melas)
  • catfish (Ictalurus)
  • humans (Homo sapiens)
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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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Morphology

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Shovelnose sturgeons have an elongated body with five rows of sharply keeled body plates: 14-19 dorsal plates, 38-47 lateral, 10-14 ventrolateral. They are olive to yellowish-brown in color on the top with lighter sides, and white on the bottom. The head contains bony plates with short spines at the tip of the snout and anterior to the eye. The snout is rounded and pointed up. The caudal peduncle is long, depressed, and fully armored. The tail is heterocercal, with the upper lobe containing long, thread-like filaments. All four barbels are evenly spaced under the jaw, unlike the related pallid sturgeon, which has barbels unevenly spaced under the jaw. The most readily distinguishing characteristic of the shovelnose is its small size and dark color. They rarely exceed 5 lbs (2.3 kg) or over 28 inches (70 cm)in length.

Range mass: .050 to 8.170 kg.

Average mass: 2.500 kg.

Range length: 6.0 to 106.2 cm.

Average length: 91.0 cm.

Other Physical Features: ectothermic ; heterothermic ; bilateral symmetry

Sexual Dimorphism: female larger

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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Life Expectancy

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The largest shovelnose sturgeon recorded weighed 13.72 lbs (6.2 kg). Most are not expected to live over 30 years of age.

Range lifespan
Status: wild:
61 (high) years.

Average lifespan
Status: captivity:
70 years.

Average lifespan
Status: wild:
30 years.

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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William Fink, University of Michigan-Ann Arbor
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Habitat

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Shovelnose sturgeons are primarily bottom dwellers, preferring high turbidity in large waters, and are usually found in pools downstream of sandbars or along the main channel border. The flowing freshwater is between 18-20 deg C. During low water or in the warm summer months, these sturgeons seek cooler, deep channel areas.

Average elevation: known to inhabit low elevations m.

Range depth: 2 to 7 m.

Average depth: 4 m.

Habitat Regions: temperate ; freshwater

Aquatic Biomes: benthic ; rivers and streams

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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Distribution

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Shovelnose sturgeons are a freshwater species historically found in most portions of the Mississippi and Missouri river basins. This area ranges from Montana south to Louisiana, and from Pennsylvania west to New Mexico. However, shovelnose sturgeons are no longer found in Pennsylvania, New Mexico, and large parts of Kansas, Kentucky, and Tennessee where they were once abundant. The main cause for the restriction of range is the construction of dams on many of their native waterways.

Biogeographic Regions: nearctic (Native )

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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Trophic Strategy

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Scaphirhynchus platorynchus are opportunistic feeders, eating mostly aquatic insects, mussels, worms and crustaceans. Altered stream flow can affect the sturgeon’s ability to find food and also influences the abundance of prey organisms. The barbels detect prey, and then the sucker type mouth easily picks up the food organisms. Most food is found over sand and gravel substrates.

Animal Foods: insects; mollusks; terrestrial worms; aquatic or marine worms

Foraging Behavior: filter-feeding

Primary Diet: carnivore (Insectivore , Eats non-insect arthropods, Molluscivore , Vermivore)

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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Benefits

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Shovelnose sturgeons have many benefits to humans. They have been commercially harvested since 1987, and the meat is considered a delicacy, especially when smoked. They are also considered a sport fish in 12 of the 24 states in which they occur. The roe is used as an acceptable caviar and it has also been introduced as a potential aquarium fish.

Positive Impacts: pet trade ; food

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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William Fink, University of Michigan-Ann Arbor
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Benefits

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There are no known adverse affects of shovelnose sturgeons on humans.

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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William Fink, University of Michigan-Ann Arbor
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Life Cycle

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Eggs are deposited over cobble, gravel, or rock, becoming adhesive and attaching to the bottom substrates. After five to eight days, the eggs hatch and the young begin to search for food on the river bottom. Females grow significantly faster than males.

Development - Life Cycle: indeterminate growth

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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William Fink, University of Michigan-Ann Arbor
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Conservation Status

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Shovelnose sturgeons are considered extirpated in three states, fully protected in four, and rare, or of special concern in eight states as of 1996. However, they are not currently listed as federally threatened or endangered.

US Federal List: no special status

CITES: no special status

IUCN Red List of Threatened Species: vulnerable

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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William Fink, University of Michigan-Ann Arbor
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Untitled

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Rafinesque described this species as the 97th species described in his book, "Ichthyologia Ohiensis: Natural History of the Fishes Inhabiting the River Ohio and Its Tributary Streams," published in 1820.

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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William Fink, University of Michigan-Ann Arbor
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Renee Sherman Mulcrone
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Associations

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Scaphirhynchus platorynchus feed on invertebrates, stirring up and loosening bottom substrates. The shovelnose sturgeon harbors a newly discovered virus called the Missouri River Sturgeon Iridovirus or MRSIV. The shovelnose sturgeon is host to the hickory-nut Obovaria olivaria, and may possibly be hosts for other pearly mussels including commercial species. Lamprey also parasitize the fish.

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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William Fink, University of Michigan-Ann Arbor
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Renee Sherman Mulcrone
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Reproduction

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Shovelnose sturgeons migrate upstream, sometimes as far as 540 km if unimpeded, in a search for acceptable reproduction habitat and mates. Due to proximate cues, such as day length, water flow, and water temperature, all adults of reproducing age gather at spawning sites in large numbers. Although these fish are usually bottom dwellers, they rise to the surface during this time. Commonly, the same sites are used for spawning over the years. Once at the spawning area, mating begins, and fertilization of eggs occurs.

Not all females spawn every year. The frequency is determined by food supply and ability to store adequate fat to produce mature gametes. Males however, always spawn if physically possible.

Mating System: polygynandrous (promiscuous)

Breeding begins when females are around seven years of age and the males are approximately five. Mature shovelnose sturgeons travel upstream to spawn over rocky substrates in flowing water between 17-21 deg C, usually downstream from a dam in April to early July. The male and female swim side by side while they release eggs and sperm. Once the eggs are fertilized, they adhere to the rocky substrates. It is very important that there is a continuous stream of water flowing over them, and that the substrate is not too fine, so that it does not settle over the eggs and suffocate them. After 5 to 8 days the dark gray eggs, measuring 2 to 3 mm in diameter, hatch. Within approximately three months, when the young are 15 to 20 cm, they are independent, and leave their natal habitat.

Breeding interval: Females do not spawn every year, and frequency is influenced by food supply and fat storage. If spawning does occur, once at the breeding location, breeding lasts for a few days, and then eggs are deposited.

Breeding season: April to early July

Average number of offspring: 50,000 eggs at one time for each female.

Range gestation period: 5 to 8 days.

Average gestation period: 7 days.

Average time to independence: 3 months.

Average age at sexual or reproductive maturity (female): 7 years.

Average age at sexual or reproductive maturity (male): 5 years.

Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); fertilization (External ); broadcast (group) spawning; oviparous

These sturgeons have to use a great deal of energy to swim upstream for long distances to spawning areas. Once at the spawning areas, and fertilization of eggs has occurred the female deposits approximately 50,000 eggs per spawning season. The parental involvement is very limited after eggs have been deposited. Both male and female return downstream to their original home range, leaving un-hatched eggs attached to bottom substrates at spawning areas to develop independently.

Parental Investment: pre-fertilization (Provisioning); pre-hatching/birth (Provisioning: Female)

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Lord, K. 2004. "Scaphirhynchus platorynchus" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Scaphirhynchus_platorynchus.html
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Katie Lord, University of Michigan-Ann Arbor
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William Fink, University of Michigan-Ann Arbor
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Benefits

provided by FAO species catalogs
Sport fish in 12 states of the USA. Commercial harvest data about 25 tons each year (Keenlyne, 1997).

Brief Summary

provided by FAO species catalogs
Sholvenose sturgeon are primarily a bottom dwelling species.Usually found in association with sand substrate, often near rock or gravel; they live where there is a current of 20-40 cm /s being usually found in pools dowstream from sandbars in unchanalized rivers, along the main channel border, downstream from dams, or in association with wing dams in rivers navigational training structures (Keenlyne, 1997).Oportunistic feeder that prey on aquatic invertebrates, primarily on inmature insects. By frequency, the most important food items of this species are Diptera, Trichoptera, Ephemeroptera, Odonata, Plecoptera, other insects, other invertebrates and plants. Spawning habits of sholvenose sturgeon has not been described. Spawning is believed to occur over hard substrate in primary tributary streams to the main rivers, or along the border of the amin river channels. The capture of specimens in spawning condition indicates that this occur from late April to June. Egg are adhesive and requires current for proper development. Most males become sexually mature at age 5, while most females do not mature until age 7. In areas of poor food supply, males and females become sexually mature at a smaller size. Following sexual maturity is influence by food supply and ability to store adequate fat to produce mature gametes (mostly fom Keenlyne, 1997 and Mayden & Kuhajda, 1996). Few individuals probably exceed 12-15 years of age.

Size

provided by FAO species catalogs
Maximum size: 86 cm TL. Maximum weight: up to 7 Kg (Keenlyne, 1997). Usually less than 2.5 Kg.

Diagnostic Description

provided by FAO species catalogs
Spiracle absent. Snout flat, spadelike. Caudal peduncle depressed, long, and completely armed. Mouth small, transverse. Lower lip continuous, with 4 lobes. Branchiostegal connection narrow. Eye very small. Barbels with accesory fringe, evently placed. D:29-36; A:18-24 rays. 13-19 (common 14-18) dorsal scutes. 38-50 (42-46) lateral scutes, that are tilted forward anteriorly. 9-14 (11-14) ventral scutes. with 8-9 scutes posterior to both, dorsal and anal fins. 4-6 scutes between anus and anal fin. With preorbital spines and with 1-17 spines on the dorsal tip of the snout. Spines significantly larger that those present in homologous areas of S. suttkusi. Belly and breast of specimens larger than 20 cm, completely armored with subrhombic plates, becoming much smaller forward. Pale olive, darker above, where the color is often a yellowish brown. Belly whitish (Forbes & Richardson, 1908).

References

  • Bailey, R.M & F.B. Cross - 1954River sturgeons of the American genus Scaphirhynchus: characters, distribution and synonymy. Pap. Michigan Acad.Sci., Arts and Letters . 39: 169-208.
  • Carlson, D.M., W.L Phliegger, L. Trial & P.S. Haverland - 1985. Distribution, biology, and hybridization of Scaphirhynchus albus and S. platorynchus in the Missouri and Mississippi rivers. Environ. Biol. Fish. 14: 51-59.
  • Cross, F. B. - 1967. Handbook of fishes of Kansas. Mus. Nat. Hist., Univ. Kans., Miscel. Publ. 45:1-357.
  • Helms, D.R - 1974. Age and growth of the sholvenose sturgeon, Scaphirhynchus platorynchus (Rafinesque), in the Mississippi River. Proc. Iowa Acad. Sci. 81:73-75 .
  • Keenlyne, K.D. - 1997. Life history and status of the sholvenose sturgeon, Scaphirhynchus platorynchus. Environ. Biol. Fish. 48: 291-298.
  • Mayden, R.L. & B.R. Kuhajda - 1997. Threatened fishes of the world: Scaphirhynchus suttkusi Williams & Clemmer, 1991 (Acipenseridae). Environ. Biol. Fish. 48: 418-419.
  • Mayden, R.L. & B.R. Kuhajda - 1996. Systematics, taxonomy, and conservation status of the endangered Alabama sturgeon, Scaphirhynchus suttkusi Williams and Clemmer (Actinopterygii, Acipenseridae). Copeia . 1996(2): 241-273.
  • Modde, T. & J.C. Schmulbach - 1977. Food and feeding behavior of the shovelnose sturgeon, Scaphirhynchus platorynchus, in the unchannelized Missouri River, South Dakota. Trans. Am. Fish. Soc. 106(6):602-608.
  • Page, L.M & B.M. Burr - 1991. A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p.
  • Williams, J.D & G.H. Clemmer - 1991. Scaphirhynchus suttkusi , a new sturgeon from the Mobile Basin of Alabama and Mississippi. Bull. Alabama Mus. Nat. Hist. 10:17-31.

Trophic Strategy

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Feeds mainly on insects (Ref. 30604).
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Recorder
Pascualita Sa-a
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Migration

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Potamodromous. Migrating within streams, migratory in rivers, e.g. Saliminus, Moxostoma, Labeo. Migrations should be cyclical and predictable and cover more than 100 km.
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Recorder
Rainer Froese
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Diagnostic Description

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Bases of outer barbels in line with or ahead of inner barbels (Ref. 86798).
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Rodolfo B. Reyes
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Biology

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Occurs in the bottom of main channels and embayments of large turbid rivers; frequently in flowing water over sand and mixed with gravel and mud. Feeds primarily on larvae of aquatic insects (especially burrowing mayflies, caddis larvae and midge larvae (Ref. 10294)), worms, and crustaceans (Ref. 50937). Spawning occurs from April to June in large rivers in areas of swift current and coarse substrates (Ref. 10294).
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Importance

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gamefish: yes; aquarium: public aquariums
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Shovelnose sturgeon

provided by wikipedia EN

The shovelnose sturgeon (Scaphirhynchus platorynchus)[5] is the smallest species of freshwater sturgeon native to North America. It is often called hackleback, sand sturgeon, or switchtail. Switchtail refers to the long filament found on the upper lobe of the caudal fin (often broken off as adults). Shovelnose sturgeon are the most abundant sturgeon found in the Missouri River and Mississippi River systems, and were formerly a commercially fished sturgeon in the United States of America (Pflieger 1997). In 2010, they were listed as threatened under the U.S. Endangered Species Act due to their resemblance to the endangered pallid sturgeon (S. albus),[6] with which shovelnose sturgeon are sympatric.[6][7]

Description

A shovelnose sturgeon.

The sturgeons of the family Acipenseridae have bony scutes along the sides and back and four barbels on the underside of the rostrum. A total of 25 extant species of sturgeon are recognized, including 17 within the genus Acipenser. Sturgeon are distributed around the northern part of the Northern Hemisphere (holarctic distribution) and have marine, freshwater, and anadromous members. Sturgeons, including the shovelnose, are highly regarded for their flesh and their roe, from which premium grades of caviar are made (Barton 2007).

They can reach 1 metre (39 in) in length and up to 4.8 kilograms (11 lb) in weight but 50–85 centimetres (20–33 in) and 2.5 kilograms (5.5 lb) is more common.

The scientific name Scaphirhynchus, Greek, means "spade snout", and platorynchus, Greek, means "broad snout". The shovelnose sturgeon is characterized by a long slender filament on the upper lobe of the caudal fin. They have a flattened rostrum (modified snout) that is also shovel-shaped. There are four fringed barbels on the ventral side of the rostrum that can be found in a straight line, which is equidistant from the mouth opening to the tip of the snout, unlike pallid sturgeon. The belly of the shovelnose sturgeon is covered with scale-like plates, which is another distinguishing factor from pallid sturgeon who have primarily scaleless bellies. Coloration of the shovelnose sturgeon ranges from a light-brown to buff with a white belly (Pflieger 1997).

Distribution, habitat, and reproduction

Shovelnose sturgeon

The shovelnose sturgeon, as its distribution represents, is impacted very little by turbidity. The Missouri River and the Mississippi River systems tend to carry high sediment loads. Here the sturgeon inhabits the open channel or main channel areas of the large rivers. It lives on the bottom, often in areas with swift current and sand or gravel bottom. As with many riverine fish species, the shovelnose sturgeon does not have a restricted home range and may travel long distances (Pflieger 1997).

The shovelnose sturgeon feeds on the bottom, using its highly protrusible mouth to suck up its food. The diet is mainly aquatic insect larvae, consisting principally of mayflies, true flies (Diptera), and caddisflies. The sturgeon also feed on crustaceans, worms, and small fish (Carlson et al. 1985;Held 1969). As a result, to bottom feeding it is a host to the glochidia (larvae) of several species of freshwater mollusc, including Quadrula pustulosa (pimpleback), Obovaria olivaria (hickorynut) and Lampsilis teres (yellow sandshell). It is the only known host of the hickorynut mussel.[7]

A typical sturgeon's life history includes a migration from feeding grounds to breeding grounds in large rivers. During spawning, behavior changes and swimming near the surface occurs. Females do not spawn every year and spawning chronology is not readily evident. Spawning takes place over gravel in fairly swift water. Eggs hatch after 3 to 5 days, and the larvae—about 1 cm long—drift downstream to suitable rearing areas in the river (Barton 2007). Carlson et al. (1985) found that the growth of the shovelnose sturgeon is relatively slow, reaching 21.3 inches (54 cm) in five years.

Conservation

The roe of the shovelnose sturgeon was marketed as "hackleback" caviar. As old-world sources of Caspian and Black Sea sturgeon caviar became overfished, especially in the wake of the collapse of the Soviet Union, roe from shovelnose sturgeon became commercially important.[7] The flesh of the sturgeon is widely considered a delicacy, especially smoked sturgeon. Poaching of the shovelnose sturgeon is becoming a problem, as they must be 8–10 years old before spawning can occur, and females do not become gravid every year. There has some interest in marketing the shovelnose sturgeon as an aquarium species.[7] In 2010, the shovelnose sturgeon was listed as threatened under the U.S. Endangered Species Act due to its resemblance to the endangered pallid sturgeon (Scaphirhynchus albus),[6] with which shovelnose sturgeon are sympatric.[6][7]

References

  1. ^ Phelps, Q.; Webb, M. (2022). "Scaphirhynchus platorynchus". IUCN Red List of Threatened Species. 2022: e.T19943A81762958. doi:10.2305/IUCN.UK.2022-1.RLTS.T19943A81762958.en. Retrieved 6 March 2023.
  2. ^ "Appendices | CITES". cites.org. Retrieved 14 January 2022.
  3. ^ Froese, R.; Pauly, D. (2017). "Acipenseridae". FishBase version (02/2017). Retrieved 18 May 2017.
  4. ^ "Acipenseridae" (PDF). Deeplyfish- fishes of the world. Retrieved 18 May 2017.
  5. ^ a b Rafinesque, C. S. (1820). Ichthyologia Ohiensis : or, Natural history of the fishes inhabiting the river Ohio and its tributary streams, preceded by a physical description of the Ohio and its branches. Lexington, Ky: Printed for the author by W. G. Hunt.
  6. ^ a b c d 75 FR 53598
  7. ^ a b c d e "Shovelnose Sturgeon (Scaphirhynchus platorynchus)" (PDF). Onalaska, Wisconsin: La Crosse Fishery Resource Office, U.S. Fish and Wildlife Service. 16 May 2001. Archived from the original (PDF) on 25 January 2022.
  • Barton, M. 2007. Bond's Biology of Fishes, 3rd edition. Thompson Brooks/Cole, Belmont, California.
  • Carlson, D.M., W.L. Pflieger, L. Trial, and P.S. Haverland. 1985. Distribution, biology, and hybridization of Scaphirhynchus albus and S. platorynchus in the Missouri and Mississippi rivers. Env. Biol. Fishes 14(1):51-59.
  • Held, J.W. 1969. Some early summer foods of the shovelnose sturgeon in the Missouri River. Trans. Am. Fish. Soc. 98(3):514-517.
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Shovelnose sturgeon: Brief Summary

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The shovelnose sturgeon (Scaphirhynchus platorynchus) is the smallest species of freshwater sturgeon native to North America. It is often called hackleback, sand sturgeon, or switchtail. Switchtail refers to the long filament found on the upper lobe of the caudal fin (often broken off as adults). Shovelnose sturgeon are the most abundant sturgeon found in the Missouri River and Mississippi River systems, and were formerly a commercially fished sturgeon in the United States of America (Pflieger 1997). In 2010, they were listed as threatened under the U.S. Endangered Species Act due to their resemblance to the endangered pallid sturgeon (S. albus), with which shovelnose sturgeon are sympatric.

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