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Ptychoptera sculleni Alexander 1943

Scientific Names

Preferred Names

Ptychoptera sculleni Alexander 1943
Species recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from Systema Dipterorum in Species 2000 & ITIS Catalogue of Life
Ptychoptera sculleni
Species recognized by wikipedia NL and United States Species List
Ptychoptera sculleni Alexander 1943
Species recognized by EOL Dynamic Hierarchy 1.1, EOL Dynamic Hierarchy 1.1, and GBIF classification
Ptychoptera sculleni
Species recognized by wikidata_hierarchy tar gz and wikipedia EN
Ptychoptera sculleni
Recognized by BHL data coverage
Ptychoptera sculleni Alexander
Recognized by NMNH type specimens
Ptychoptera sculleni Alexander 1943
Species recognized by Catalogue of Life 26 March 2024, Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy, GBIF data coverage, and Dynamic Hierarchy Version 2.2.3 - Test

Common Names

There are no common names associated with this taxon.

Curated hierarchies for Ptychoptera sculleni Alexander 1943

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Diptera
Ptychopteridae
Ptychoptera
Ptychoptera sculleni Alexander 1943 (this page)
Ptychoptera africana Alexander 1920
Ptychoptera agnes Krzeminski & Zwick 1993
Ptychoptera albimana (Fabricius 1787)
Ptychoptera alexanderi Hancock, Marcos-Garcia & Rotheray 2006
Ptychoptera alina Krzeminski & Zwick 1993
Ptychoptera annandalei Brunetti 1918
Ptychoptera bellula Alexander 1937
Ptychoptera byersi Alexander 1966
Ptychoptera camerounensis Alexander 1921
Ptychoptera capensis Alexander 1917
Ptychoptera chalybeata Alexander 1956
Ptychoptera clitellaria Alexander 1935
Ptychoptera coloradensis Alexander 1937
Ptychoptera contaminata (Linnaeus 1758)
Ptychoptera daimio Alexander 1921
Ptychoptera delmastroi Zwick & Stary 2002
Ptychoptera distincta Brunetti 1911
Ptychoptera formosensis Alexander 1924
Ptychoptera garhwalensis Alexander 1959
Ptychoptera gutianshana Yang & Chen 1995
Ptychoptera handlirschi (Czizek 1919)
Ptychoptera helena (Peus 1958)
Ptychoptera hugoi Tjeder 1968
Ptychoptera ichitai Nakamura & Saigusa 2009
Ptychoptera ichneumonoidea Alexander 1946
Ptychoptera japonica Alexander 1913
Ptychoptera javensis Alexander 1937
Ptychoptera kosiensis Stuckenberg 1983
Ptychoptera kyushuensis Nakamura & Saigusa 2009
Ptychoptera lacustris Meigen 1830
Ptychoptera lenis Osten Sacken 1877
Ptychoptera lenta (Harris 1776)
Ptychoptera longicauda (Tonnoir 1919)
Ptychoptera longwangshana Yang & Chen 1998
Ptychoptera madagascariensis Alexander 1937
Ptychoptera malaisei Alexander 1946
Ptychoptera matongoensis Alexander 1958
Ptychoptera metallica Walker 1848
Ptychoptera minor Alexander 1920
Ptychoptera minuta Tonnoir 1919
Ptychoptera monoensis Alexander 1947
Ptychoptera obscura (Peus 1958)
Ptychoptera osceola Alexander 1959
Ptychoptera pallidicostalis Nakamura & Saigusa 2009
Ptychoptera paludosa Meigen 1804
Ptychoptera pauliani Alexander 1957
Ptychoptera pectinata Macquart 1834
Ptychoptera pendula Alexander 1937
Ptychoptera perbona Alexander 1946
Ptychoptera persimilis Alexander 1947
Ptychoptera peusi Joost 1974
Ptychoptera praescutellaris Alexander 1946
Ptychoptera quadrifasciata Say 1824
Ptychoptera ressli Theischinger 1978
Ptychoptera robinsoni Alexander 1957
Ptychoptera scutellaris Meigen 1818
Ptychoptera sikkimensis Alexander 1965
Ptychoptera silvicola Zwyrtek & Rozkosny 1967
Ptychoptera subscutellaris Alexander 1921
Ptychoptera sumatrensis Alexander 1936
Ptychoptera surcoufi (Seguy 1925)
Ptychoptera takeuchii Tokunaga 1938
Ptychoptera tibialis Brunetti 1911
Ptychoptera townesi Alexander 1943
Ptychoptera uelensis Alexander 1928
Ptychoptera uta Alexander 1947
Ptychoptera yamato Nakamura & Saigusa 2009
Ptychoptera yasumatsui Tokunaga 1939

Catalogue of Life 26 March 2024

Biota
Animalia
Arthropoda
Insecta
Diptera
Ptychopteridae
Ptychoptera
Ptychoptera sculleni Alexander 1943 (this page)
Ptychoptera
Ptychoptera africana Alexander 1920
Ptychoptera agnes Krzemiński & Zwick 1993
Ptychoptera albimanus (Fabricius 1787)
Ptychoptera alexanderi Hancock, Marcos-García & Rotheray 2006
Ptychoptera alina Krzemiński & Zwick 1993
Ptychoptera annandalei Brunetti 1918
Ptychoptera bannaensis Kang, Yao & Yang 2019
Ptychoptera bellula Alexander 1937
Ptychoptera byersi Alexander 1966
Ptychoptera camerounensis Alexander 1921
Ptychoptera capensis Alexander 1917
Ptychoptera chalybeata Alexander 1956
Ptychoptera circinans Kang, Xue & Zhang 2019
Ptychoptera clitellaria Alexander 1935
Ptychoptera coloradensis Alexander 1937
Ptychoptera contaminata (Linnaeus 1758)
Ptychoptera cordata Zhang & Kang 2021
Ptychoptera cordata Zhang & Tang 2021
Ptychoptera daimio Alexander 1921
Ptychoptera delmastroi Zwick & Starý 2003
Ptychoptera distincta Brunetti 1911
Ptychoptera emeica Kang, Xue & Zhang 2019
Ptychoptera formosensis Alexander 1924
Ptychoptera garhwalensis Alexander 1959
Ptychoptera gutianshana Yang & Chen 1995
Ptychoptera handlirschi (Czižek 1919)
Ptychoptera hekouensis Kang, Gao & Zhang 2022
Ptychoptera helena (Peus 1958)
Ptychoptera hugoi Tjeder 1968
Ptychoptera ichitai Nakamura & Saigusa 2009
Ptychoptera ichneumonoidea Alexander 1946
Ptychoptera japonica Alexander 1913
Ptychoptera javensis Alexander 1937
Ptychoptera kosiensis Stuckenberg 1983
Ptychoptera kyushuensis Nakamura & Saigusa 2009
Ptychoptera lacustris Meigen 1830
Ptychoptera lenis Osten Sacken 1877
Ptychoptera lenta (Harris 1776)
Ptychoptera lii Kang, Yao & Yang 2019
Ptychoptera longa Kang, Gao & Zhang 2022
Ptychoptera longicauda (Tonnoir 1919)
Ptychoptera longwangshana Yang & Chen 1998
Ptychoptera lucida Kang, Xue & Zhang 2019
Ptychoptera lushuiensis Kang, Yao & Yang 2019
Ptychoptera madagascariensis Alexander 1937
Ptychoptera malaisei Alexander 1946
Ptychoptera matongoensis Alexander 1958
Ptychoptera mesozoica Kalugina 1989
Ptychoptera metallica Walker 1848
Ptychoptera minor Alexander 1920
Ptychoptera minuta Tonnoir 1919
Ptychoptera miocenica (Cockerell 1910)
Ptychoptera monoensis Alexander 1947
Ptychoptera obscura (Peus 1958)
Ptychoptera osceola Alexander 1959
Ptychoptera pallidicostalis Nakamura & Saigusa 2009
Ptychoptera paludosa Meigen 1804
Ptychoptera pauliani Alexander 1957
Ptychoptera pectinata Macquart 1834
Ptychoptera pendula Alexander 1937
Ptychoptera perbona Alexander 1946
Ptychoptera persimilis Alexander 1947
Ptychoptera peusi Joost 1974
Ptychoptera praescutellaris Alexander 1946
Ptychoptera qinggouensis Kang, Yao & Yang 2019
Ptychoptera quadrifasciata Say 1824
Ptychoptera ressli Theischinger 1978
Ptychoptera robinsoni Alexander 1957
Ptychoptera scutellaris Meigen
Ptychoptera separata Kang, Xue & Zhang 2019
Ptychoptera sikkimensis Alexander 1966
Ptychoptera silvicola Zwyrtek & Rozkošný 1967
Ptychoptera subscutellaris Alexander 1921
Ptychoptera sumatrensis Alexander 1936
Ptychoptera surcoufi (Séguy 1925)
Ptychoptera takeuchii Tokunaga 1938
Ptychoptera tianmushana Shao & Kang 2021
Ptychoptera tibialis Brunetti 1911
Ptychoptera townesi Alexander 1943
Ptychoptera uelensis Alexander 1928
Ptychoptera uta Alexander 1947
Ptychoptera wangae Kang, Yao & Yang 2019
Ptychoptera xiaohuangshana Kang, Gao & Zhang 2022
Ptychoptera xinglongshana Yang 1996
Ptychoptera yamato Nakamura & Saigusa 2009
Ptychoptera yasumatsui Tokunaga 1939
Ptychoptera yunnanica Zhang & Kang 2021
Ptychoptera yunnanica Zhang & Tang 2021

Dynamic Hierarchy Version 2.2.3 - Test

Ptychoptera sculleni Alexander 1943 (this page)
Ptychoptera africana Alexander 1920
Ptychoptera agnes Krzeminski & Zwick 1993
Ptychoptera albimana (Fabricius 1787)
Ptychoptera alexanderi Hancock, Marcos-Garcia & Rotheray 2006
Ptychoptera alina Krzeminski & Zwick 1993
Ptychoptera annandalei Brunetti 1918
Ptychoptera bellula Alexander 1937
Ptychoptera byersi Alexander 1966
Ptychoptera camerounensis Alexander 1921
Ptychoptera capensis Alexander 1917
Ptychoptera chalybeata Alexander 1956
Ptychoptera clitellaria Alexander 1935
Ptychoptera coloradensis Alexander 1937
Ptychoptera contaminata (Linnaeus 1758)
Ptychoptera daimio Alexander 1921
Ptychoptera delmastroi Zwick & Stary 2002
Ptychoptera distincta Brunetti 1911
Ptychoptera formosensis Alexander 1924
Ptychoptera garhwalensis Alexander 1959
Ptychoptera gutianshana Yang & Chen 1995
Ptychoptera handlirschi (Czizek 1919)
Ptychoptera helena (Peus 1958)
Ptychoptera hugoi Tjeder 1968
Ptychoptera ichitai Nakamura & Saigusa 2009
Ptychoptera ichneumonoidea Alexander 1946
Ptychoptera japonica Alexander 1913
Ptychoptera javensis Alexander 1937
Ptychoptera kosiensis Stuckenberg 1983
Ptychoptera kyushuensis Nakamura & Saigusa 2009
Ptychoptera lacustris Meigen 1830
Ptychoptera lenis Osten Sacken 1877
Ptychoptera lenta (Harris 1776)
Ptychoptera longicauda (Tonnoir 1919)
Ptychoptera longwangshana Yang & Chen 1998
Ptychoptera madagascariensis Alexander 1937
Ptychoptera malaisei Alexander 1946
Ptychoptera matongoensis Alexander 1958
Ptychoptera metallica Walker 1848
Ptychoptera minor Alexander 1920
Ptychoptera minuta Tonnoir 1919
Ptychoptera monoensis Alexander 1947
Ptychoptera obscura (Peus 1958)
Ptychoptera osceola Alexander 1959
Ptychoptera pallidicostalis Nakamura & Saigusa 2009
Ptychoptera paludosa Meigen 1804
Ptychoptera pauliani Alexander 1957
Ptychoptera pectinata Macquart 1834
Ptychoptera pendula Alexander 1937
Ptychoptera perbona Alexander 1946
Ptychoptera persimilis Alexander 1947
Ptychoptera peusi Joost 1974
Ptychoptera praescutellaris Alexander 1946
Ptychoptera quadrifasciata Say 1824
Ptychoptera ressli Theischinger 1978
Ptychoptera robinsoni Alexander 1957
Ptychoptera scutellaris Meigen 1818
Ptychoptera sikkimensis Alexander 1965
Ptychoptera silvicola Zwyrtek & Rozkosny 1967
Ptychoptera subscutellaris Alexander 1921
Ptychoptera sumatrensis Alexander 1936
Ptychoptera surcoufi (Seguy 1925)
Ptychoptera takeuchii Tokunaga 1938
Ptychoptera tibialis Brunetti 1911
Ptychoptera townesi Alexander 1943
Ptychoptera uelensis Alexander 1928
Ptychoptera uta Alexander 1947
Ptychoptera yamato Nakamura & Saigusa 2009
Ptychoptera yasumatsui Tokunaga 1939

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Diptera
Ptychopteridae
Ptychoptera
Ptychoptera sculleni Alexander 1943 (this page)
Ptychoptera africana Alexander 1920
Ptychoptera agnes Krzeminski & Zwick 1993
Ptychoptera albimana (Fabricius 1787)
Ptychoptera alexanderi Hancock, Marcos-Garcia & Rotheray 2006
Ptychoptera alina Krzeminski & Zwick 1993
Ptychoptera annandalei Brunetti 1918
Ptychoptera bellula Alexander 1937
Ptychoptera byersi Alexander 1966
Ptychoptera camerounensis Alexander 1921
Ptychoptera capensis Alexander 1917
Ptychoptera chalybeata Alexander 1956
Ptychoptera clitellaria Alexander 1935
Ptychoptera coloradensis Alexander 1937
Ptychoptera contaminata (Linnaeus 1758)
Ptychoptera daimio Alexander 1921
Ptychoptera delmastroi Zwick & Stary 2002
Ptychoptera distincta Brunetti 1911
Ptychoptera formosensis Alexander 1924
Ptychoptera garhwalensis Alexander 1959
Ptychoptera gutianshana Yang & Chen 1995
Ptychoptera handlirschi (Czizek 1919)
Ptychoptera helena (Peus 1958)
Ptychoptera hugoi Tjeder 1968
Ptychoptera ichitai Nakamura & Saigusa 2009
Ptychoptera ichneumonoidea Alexander 1946
Ptychoptera japonica Alexander 1913
Ptychoptera javensis Alexander 1937
Ptychoptera kosiensis Stuckenberg 1983
Ptychoptera kyushuensis Nakamura & Saigusa 2009
Ptychoptera lacustris Meigen 1830
Ptychoptera lenis Osten Sacken 1877
Ptychoptera lenta (Harris 1776)
Ptychoptera longicauda (Tonnoir 1919)
Ptychoptera longwangshana Yang & Chen 1998
Ptychoptera madagascariensis Alexander 1937
Ptychoptera malaisei Alexander 1946
Ptychoptera matongoensis Alexander 1958
Ptychoptera metallica Walker 1848
Ptychoptera minor Alexander 1920
Ptychoptera minuta Tonnoir 1919
Ptychoptera monoensis Alexander 1947
Ptychoptera obscura (Peus 1958)
Ptychoptera osceola Alexander 1959
Ptychoptera pallidicostalis Nakamura & Saigusa 2009
Ptychoptera paludosa Meigen 1804
Ptychoptera pauliani Alexander 1957
Ptychoptera pectinata Macquart 1834
Ptychoptera pendula Alexander 1937
Ptychoptera perbona Alexander 1946
Ptychoptera persimilis Alexander 1947
Ptychoptera peusi Joost 1974
Ptychoptera praescutellaris Alexander 1946
Ptychoptera quadrifasciata Say 1824
Ptychoptera ressli Theischinger 1978
Ptychoptera robinsoni Alexander 1957
Ptychoptera scutellaris Meigen 1818
Ptychoptera sikkimensis Alexander 1965
Ptychoptera silvicola Zwyrtek & Rozkosny 1967
Ptychoptera subscutellaris Alexander 1921
Ptychoptera sumatrensis Alexander 1936
Ptychoptera surcoufi (Seguy 1925)
Ptychoptera takeuchii Tokunaga 1938
Ptychoptera tibialis Brunetti 1911
Ptychoptera townesi Alexander 1943
Ptychoptera uelensis Alexander 1928
Ptychoptera uta Alexander 1947
Ptychoptera yamato Nakamura & Saigusa 2009
Ptychoptera yasumatsui Tokunaga 1939

GBIF classification

Animalia
Arthropoda
Insecta
Diptera
Ptychopteridae
Ptychoptera Meigen 1803
Ptychoptera sculleni Alexander 1943 (this page)
Bittacomorpha miocenica (Cockerell 1910)
Ptychoptera africana Alexander 1920
Ptychoptera agnes Krzeminski & Zwick 1993
Ptychoptera albimana (Fabricius 1787)
Ptychoptera alexanderi Hancock, Marcos-Garcia & Rotheray 2006
Ptychoptera alina Krzeminski & Zwick 1993
Ptychoptera annandalei Brunetti 1918
Ptychoptera bannaensis
Ptychoptera bellula Alexander 1937
Ptychoptera byersi Alexander 1966
Ptychoptera camerounensis Alexander 1921
Ptychoptera capensis Alexander 1917
Ptychoptera chalybeata Alexander 1956
Ptychoptera clitellaria Alexander 1935
Ptychoptera coloradensis Alexander 1937
Ptychoptera contaminata (Linnaeus 1758)
Ptychoptera daimio Alexander 1921
Ptychoptera deleta Novak 1877
Ptychoptera delmastroi Zwick & Stary 2002
Ptychoptera distincta Brunetti 1911
Ptychoptera eocenica Podenas 2007
Ptychoptera formosensis Alexander 1924
Ptychoptera garhwalensis Alexander 1959
Ptychoptera gutianshana Yang & Chen 1995
Ptychoptera handlirschi (Czizek 1919)
Ptychoptera helena (Peus 1958)
Ptychoptera hugoi Tjeder 1968
Ptychoptera ichitai Nakamura & Saigusa 2009
Ptychoptera ichneumonoidea Alexander 1946
Ptychoptera japonica Alexander 1913
Ptychoptera javensis Alexander 1937
Ptychoptera kosiensis Stuckenberg 1983
Ptychoptera kyushuensis Nakamura & Saigusa 2009
Ptychoptera lacustris Meigen 1830
Ptychoptera lenis Osten Sacken 1877
Ptychoptera lenta (Harris 1776)
Ptychoptera lii
Ptychoptera longicauda (Tonnoir 1919)
Ptychoptera longwangshana Yang & Chen 1998
Ptychoptera lushuiensis
Ptychoptera madagascariensis Alexander 1937
Ptychoptera malaisei Alexander 1946
Ptychoptera matongoensis Alexander 1958
Ptychoptera mesozoica Kalugina 1989
Ptychoptera metallica Walker 1848
Ptychoptera minor Alexander 1920
Ptychoptera minuta Tonnoir 1919
Ptychoptera monoensis Alexander 1947
Ptychoptera obscura (Peus 1958)
Ptychoptera osccola Alexander 1959
Ptychoptera osceola Alexander 1959
Ptychoptera pallidicostalis Nakamura & Saigusa 2009
Ptychoptera paludosa Meigen 1804
Ptychoptera pauliani Alexander 1957
Ptychoptera pectinata Macquart 1834
Ptychoptera pendula Alexander 1937
Ptychoptera perbona Alexander 1946
Ptychoptera persimilis Alexander 1947
Ptychoptera peusi Joost 1974
Ptychoptera praescutellaris Alexander 1946
Ptychoptera qinggouensis
Ptychoptera quadrifasciata Say 1824
Ptychoptera ressli Theischinger 1978
Ptychoptera robinsoni Alexander 1957
Ptychoptera scutellaris Wiedemann 1818
Ptychoptera sikkimensis Alexander 1965
Ptychoptera silvicola Zwyrtek & Rozkosny 1967
Ptychoptera subscutellaris Alexander 1921
Ptychoptera sumatrensis Alexander 1936
Ptychoptera surcoufi (Seguy 1925)
Ptychoptera takeuchii Tokunaga 1938
Ptychoptera tibialis Brunetti 1911
Ptychoptera townesi Alexander 1943
Ptychoptera uelensis Alexander 1928
Ptychoptera uta Alexander 1947
Ptychoptera wangae
Ptychoptera yamato Nakamura & Saigusa 2009
Ptychoptera yasumatsui Tokunaga 1939

Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Diptera
Nematocera
Ptychopteromorpha
Ptychopteridae
Ptychopterinae
Ptychoptera Meigen 1803
Ptychoptera sculleni Alexander 1943 (this page)
Ptychoptera byersi Alexander 1966
Ptychoptera coloradensis Alexander 1937
Ptychoptera metallica Walker 1848
Ptychoptera minor Alexander 1920
Ptychoptera monoensis Alexander 1947
Ptychoptera osccola Alexander 1959
Ptychoptera pendula Alexander 1937
Ptychoptera quadrifasciata Say 1824
Ptychoptera townesi Alexander 1943
Ptychoptera uta Alexander 1947

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Holometabola
Diptera
Nematocera
Ptychopteridae
Ptychoptera
Ptychoptera sculleni (this page)
Paraptychoptera
Ptychoptera
Ptychoptera africana
Ptychoptera agnes
Ptychoptera albimana
Ptychoptera alexanderi
Ptychoptera alina
Ptychoptera annandalei
Ptychoptera bannaensis
Ptychoptera bellula
Ptychoptera byersi
Ptychoptera camerounensis
Ptychoptera capensis
Ptychoptera chalybeata
Ptychoptera circinans
Ptychoptera clitellaria
Ptychoptera coloradensis
Ptychoptera contaminata
Ptychoptera daimio
Ptychoptera delmastroi
Ptychoptera distincta
Ptychoptera emeica
Ptychoptera formosensis
Ptychoptera garhwalensis
Ptychoptera gutianshana
Ptychoptera handlirschi
Ptychoptera helena
Ptychoptera hugoi
Ptychoptera ichitai
Ptychoptera ichneumonoidea
Ptychoptera japonica
Ptychoptera javensis
Ptychoptera kosiensis
Ptychoptera kyushuensis
Ptychoptera lacustris
Ptychoptera lenis
Ptychoptera lenta
Ptychoptera lii
Ptychoptera longicauda
Ptychoptera longwangshana
Ptychoptera lucida
Ptychoptera lushuiensis
Ptychoptera madagascariensis
Ptychoptera malaisei
Ptychoptera matongoensis
Ptychoptera metallica
Ptychoptera minor
Ptychoptera minuta
Ptychoptera monoensis
Ptychoptera obscura
Ptychoptera osceola
Ptychoptera pallidicostalis
Ptychoptera paludosa
Ptychoptera pauliani
Ptychoptera pectinata
Ptychoptera pendula
Ptychoptera perbona
Ptychoptera persimilis
Ptychoptera peusi
Ptychoptera praescutellaris
Ptychoptera qinggouensis
Ptychoptera quadrifasciata
Ptychoptera ressli
Ptychoptera robinsoni
Ptychoptera scutellaris
Ptychoptera scutilifera
Ptychoptera separata
Ptychoptera sikkimensis
Ptychoptera silvicola
Ptychoptera subscutellaris
Ptychoptera sumatrensis
Ptychoptera surcoufi
Ptychoptera takeuchii
Ptychoptera tibialis
Ptychoptera townesi
Ptychoptera uelensis
Ptychoptera uta
Ptychoptera wangae
Ptychoptera yamato
Ptychoptera yankovskiana
Ptychoptera yasumatsui
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