Leptonema crassum Ulmer 1905

Leptonema crassum Ulmer

Leptonema crassum Ulmer, 1905a:58.—Mosely, 1933:12.—Fischer, 1963:168.

This species has been widely reported from Latin America, being known from central Mexico south throughout Central and South America to northern Argentina. Since it appears generally to breed in the larger rivers I suspect that the examples taken at Rancho Grande represent windblown specimens.

MATERIAL.—VENEZUELA, ARAGUA: Rancho Grande, 10–21 Feb 1969, 1 1. BARINAS: Río Santo Domingo, Barinas, 17 Feb 1976, 16 1. Barinitas, 22–26 Feb 1969, 10 6. Puente Parangula, 8 km S Barinitas, 18 Feb 1976, 4. MERIDA: El Vigia, 2 Jun 1976, 1.

Leptonema crassum Ulmer

Leptonema crassum Ulmer, 1905a:58, 59, Pl. II: figs. 52, 53 [holotype , in ZSZMH. destroyed 1943].—Fischer, 1963:168; 1972:156.—Weidner, 1964:84.—Flint. 1966:5; 1972:235 [Argentina]; 1978:385.

MAP 10.—Distributions of Leptonema guyanense, new species. L. crassum Ulmer, and L. divaricatum, new species.

Leptonema radiale Navás, 1927:42 [holotype , in DEI].—Flint. 1978:385 [synonymy].

Leptonema grisolinum Navás, 1933a:312, 313, fig. 23 [holotype , in MNHNP] [new synonymy].

TYPE-LOCALITIES.—Of crassum, Espirito Santo, Brazil. Of radiale, Minas Gerais, Brazil. Of grisolinum, Caucagua, Edo. Miranda, Venezuela.

DISTRIBUTION.—Argentina, Brazil, Colombia, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Venezuela.

MATERIAL EXAMINED.—ARGENTINA, Pcia. Misiones, Puerto Rico, 4–8 Apr 1971, C.M. and O.S. Flint, Jr., 2, 3; Arroyo Saura, 9 km N L.N. Alem, 20 Nov 1973, O.S. Flint, Jr., 4; Río Iguazú, Camp Nañdu, 25 Nov 1973, O.S. Flint, Jr., 7, 3; Iguazú, 30Jan-13 Mar 1945, Hayward et al., 5; Puerto Libertad, 24 Nov 1973, O.S. Flint, Jr., 24, 24; Arroyo Piray Guazú, N San Pedro, 22 Nov 1973, O.S. Flint, Jr., 1, 1; Arroyo Liso, 8 km W General Güemes, 19 Nov 1973, O.S. Flint, Jr., 1. Pcia. Entre Rios, Río Uruguay, Salto Grande, 16 Nov 1973, O.S. Flint, Jr., 1. BRAZIL, Madeira-Mamore R.R. Co. Camp 43, Mann and Baker, 2. Edo. Mato Grosso, Aripuana Dist., 10°11′S, 59°48′W, 17 Mar 1977, N.D. Penny, 1; Buriti, Chapada dos Guimaraes, 1200 m, 17–28 Jan 1972, E.G., I., and E.A. Munroe, 1. 1. Edo. Goias, Fazenda Nova Orlandia, Jatai, Jan 1964, Martins et al., 1. Edo. Minas Gerais, Serra do Cipó, km 110, 29 Oct 1974, C.G. Froehlich (#397), 20, 5. Edo. São Paulo, Piracicaba, 12 Oct 1965, C.A. Triplehorn, 1, 2. COLOMBIA, Dpto. Meta, Restrepo, 500 m, 7–10 Aug 1936, M. Bates, 1, 3.

COSTA RICA, [Pcia. Limon], Cairo, 21 Apr 1944, P. Knight, 1; Hamburg Farm, 28 Mar, C.W. Dodge, 2, 1. [Pcia. Alajuela], San Carlos, 1. [Pcia. Guanacaste], 10 mi NW Liberia, 25 Jul 1965, P.J. Spangler, 2. Pcia. Heredia, Puerto Viejo, 6 Apr 1969, D.C. Rentz, 1; Finca La Selva, near Puerto Viejo, 30 Jun 1967, D.R. Paulson, 2, 1; same, but 22–31 Jul 1969, D.H. Janzen, 1. GUATEMALA, [Dpto. Izabal], Los Amates, Kellerman, 4, 5; Cayuga, Sep 1915, Wm. Schaus, 1. HONDURAS, [Dpto. Atlántida], Lancetilla, Tela, 2 May, M. Bates, 2. [Dpto. Colón], Puerto Castilla, 21 Jun 1926, R.H. Painter, 1. Dpto. Comayagua, Las Limas, 2000′, 20–13 May, J.B. Edwards, 4, 2. MEXICO, Edo. Oaxaca, Río Valle Nacional, Chilte-pec, 25 May 1981, C.M. and O.S. Flint, Jr., 21, 3; Palomares, 16–21 Sep 1961, R. and K. Dreisbach, 1; San Mateo Yetla, 3 km S Valle Nacional, 25 May 1981, C.M. and O.S. Flint, Jr., 1, 1; Arroyo Choapan, Bethania, 31 km S Tuxtepec, 24 May 1981, C.M. and O.S. Flint, Jr., 2; Tuxtepec, 7 Nov 1934, J. Camelo G., 1; Uxpanapa, 27 Sep 1977, J. Bueno S., 1O; Monte Flor, 19 May 1956, 1; Camelia Roja, 2 May 1975, J. Bueno S., 3. Finca San Carlos, Colonia Agricola Progresso, Municipio Matias Romero, 28 May 1959, F. Medellín, 2. Edo. Chiapas, Cascada Misjolha. 20 km S Palenque, 17–18 May 1981, C.M. and O.S. Flint, Jr., 19; Río Lacanjá, 22 km N Ocosingo, 19 May 1981, C.M. and O.S. Flint, Jr., 1; Bonampak, 3 May 1978, H. Brailovsky, 1; Sta. Elena, 20 km beyond Lagunas de Montebello, 8 Apr 1979, J. Bueno S., 1. Edo. Tabasco, Río Puyacatengo, E Teapa, 28–29 Jul 1966, Flint and Ortiz, 1.

NICARAGUA, [Dpto. Nueva Segova], Río Coco, Ocotal, 31 Jul 1967, O.S. Flint, Jr., 1. PANAMA, [Pcia. Panamá], Taboga Island, 18 Feb 1912, A. Busck, 1. [Pcia. Colón], Alajuela [Alhajuela], 10–17 Apr 1911, A. Busck, 3, 12; Chilibre, 14 May, N. Banks, 1. [Canal Zone], Tabernilla, 4 May 1907, A. Busck, 1, 1. Pcia. Veraguas, Río Jorones, Jorones, 5 Jun 1983, P.J. Spangler, 2. Pcia. Chiriquí, Río San Felix, Las Lajas, 28 May 1983, P.J. Spangler, 1; Lino, Fassl, 1. Pcia. Darién, Río Tuira at Boca de Cupe, 18 Feb 1985, J. Louton, 6, 7; Río Tuira at Río Pucuro, 16–17 Feb 1985, J. Louton, 12, 3. Com. San Blas, Río Coatí Grande near coast, 2 Mar 1985, Flint and Louton, 42, 28. PARAGUAY, Bella Vista, Nov, F. Schade, 1. PERU, [Dpto. Cuzco], Quince Mil, 20–30 Oct 1962, L.E. Peña G., 1. VENEZUELA, Edo. Barinas, Barinitas, 22–23 Feb 1969, P. and P. Spangler, 5. Edo. Miranda, Caucagua, Oct 1923, M. Grisol, holotype grisolinum; valley Río Guira, SW Panaquire, 10°12′N, 66°17′W, 280 m, 18–21 Sep 1979, Fernandez and Clavijo, 3, 2. Edo. Zulia, Dist. Baralt, Río Paují at Rt 3, between San Juan and San Antonio, 9–11 Oct 1979, H.M. Savage, 13, 4. Material in CNC, CUI, DEI, IBUNAM, IML, INHS, IZAM, MCZ, MNHNP, USNM, and USP.