Orobdella ketagalan Nakano & Lai 2012

Body firm, muscular, elongated, gaining regularly in width in caudal direction, dorso-ventral depressed, sides nearly parallel from mid-length to point just anterior to caudal sucker, BL 70.9 mm, BW 6.4 mm (Fig. 2). Caudal sucker ventral, oval, diameter smaller than BW (Figs 2B, 3D). Color faded in preservative (Fig. 2). Somite I completely merged with prostomium (Fig. 3A). Somite II uniannulate, not separated from I (Fig. 3A). Somites III and IV uniannulate (Fig. 3A). Somite V biannulate, (a1 + a2) = a3, a3 forming posterior margin of oral sucker (Fig. 3A, B). Somites VI and VII triannulate, a1 = a2 = a3 (Fig. 3A, B). Somites VIII–XXV quadrannulate, a1 = a2 = b5 = b6 (Fig. 3A–E); b5 of X being first annulus on clitellum, a2 of XIII being last annulus of clitellum (Fig. 3E). Somite XXVI triannulate, a1 > a2 > a3, a3 being last complete annulus on venter (Fig. 3C, D). Somite XXVII incomplete uniannulate with slight furrow (Fig. 3C); anus behind it with no post-anal annulus (Fig. 3C). Anterior ganglionic mass in VI a2 and a3. Ganglia VIII–XXI in a2 of each somite (Fig. 4A). Ganglion XIII in a2 and b5 (Fig. 4A). Ganglia XIV–XXIII in a2 of each somite (Fig. 4A). Ganglia XXIV and XXV in a1 and a2 of each somite. Ganglion XXVI in b6 of somite XXV. Posterior ganglionic mass in XXVI a1–a3. Eyes, three pairs, first pair dorsally in furrow of II/III, second and third pairs dorsolaterally on posterior margin of V (a1 + a2) (Fig. 3A). Nephridiopores, 17 pairs, ventrally at posterior margin of a1 of each somite of VIII–XXIV (Fig. 3B, E). Papillae numerous, minute, hardly visible, one row on every annulus. Pharynx agnathous, euthylaematous, reaching to XV a1 (Fig. 3F). Crop tubular, acecate, in XV a1 to XXI a2. Gastropore conspicuous, ventral, located middle of XIII a1 (Fig. 3E, G). Gastroporal duct narrow, simple tubular, joining with crop in XIV/XV (Fig. 3F). Intestine tubular, acecate, in XXI a2 to XXIV b5/b6. Rectum tubular, thin-walled. Male gonopore located at middle of XI b6 (Fig. 3E). Female gonopore at middle of XIII a1, inconspicuous, located behind gastropore (Fig. 3E, G). Gonopores separated by 1/2 + 4 + 1/2 annuli (Fig. 3E). Testisacs multiple, one or two testisacs on each side in each annulus, in XV a2 to XXV b5 (Fig. 4A). Sperm ducts in XI b5 to XV a2, coiled in XIII b5 to XV a1 (Fig. 4A): small paired sperm duct bulbs in XV a1 (Fig. 4A, B). Epididymides absent. Ejaculatory bulbs absent. Paired atrial cornua in XI b5 and b6, undeveloped, coniform (Fig. 4A, C). Atrium body short, muscular, globular in XI b5 and b6 (Fig. 4A, C–E). Penis sheath and penis absent. Ovisacs, one pair, thin-walled, globular, in XIII a2 and b5 (Fig. 4A, F). Oviducts thin-walled, right oviduct crossing ventrally beneath nerve cord, both oviducts converging into common oviduct in XIII a1/a2 (Fig. 4A, F). Common oviduct thin-walled, very short, directly ascending to female gonopore (Fig. 4F).

Known from Yangmingshan National Park and adjacent areas in northern Taipei City, Taiwan (Fig. 1).