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Ptychadena amharensis Smith, Noonan & Colston 2017
collect
overview
data
names
Scientific Names
Preferred Names
Ptychadena amharensis Smith, Noonan & Colston 2017
Species recognized by
EOL Dynamic Hierarchy April 2022
Reference taxon from Integrated Taxonomic Information System (ITIS)
Ptychadena amharensis
Smith, Noonan & Colston 2017
Species recognized by
EOL Dynamic Hierarchy 1.1
,
EOL Dynamic Hierarchy 1.1
, and
Frost Darrel R 2018 Amphibian Species of the World
Ptychadena amharensis
Species recognized by
wikidata_hierarchy tar gz
Ptychadena amharensis Smith, Noonan & Colston 2017
Species recognized by
Catalogue of Life 26 March 2024
,
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
,
GBIF data coverage
,
Dynamic Hierarchy Version 2.2.3
, and
EOL Dynamic Hierarchy Version 2.2.4
Common Names
There are no common names associated with this taxon.
Curated hierarchies for Ptychadena amharensis Smith, Noonan & Colston 2017
EOL Dynamic Hierarchy April 2022
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Deuterostomia
Chordata
Vertebrata
Gnathostomata
Osteichthyes
Sarcopterygii
Tetrapoda
Lissamphibia
Anura
Ptychadenidae Dubois 1987
Ptychadena Boulenger 1917
Ptychadena amharensis Smith, Noonan & Colston 2017
(this page)
Ptychadena aequiplicata (Werner 1898)
Ptychadena anchietae (Bocage 1868)
Ptychadena ansorgii (Boulenger 1905)
Ptychadena arnei Perret 1997
Ptychadena baroensis Smith, Noonan & Colston 2017
Ptychadena bibroni (Hallowell 1845)
Ptychadena boettgeri (Pfeffer 1893)
Ptychadena broadleyi Stevens 1972
Ptychadena bunoderma (Boulenger 1907)
Ptychadena christyi (Boulenger 1919)
Ptychadena chrysogaster Laurent 1954
Ptychadena cooperi (Parker 1930)
Ptychadena erlangeri (Ahl 1924)
Ptychadena gansi Laurent In Gans, Laurent & Pandit 1965
Ptychadena goweri Smith, Noonan & Colston 2017
Ptychadena grandisonae Laurent 1954
Ptychadena guibei Laurent 1954
Ptychadena harenna Largen 1997
Ptychadena hylaea Schmidt & Inger 1959
Ptychadena ingeri Perret 1991
Ptychadena keilingi (Monard 1937)
Ptychadena levenorum Smith, Noonan & Colston 2017
Ptychadena longirostris (Peters 1870)
Ptychadena mahnerti Perret 1996
Ptychadena mapacha Channing 1993
Ptychadena mascareniensis (Duméril & Bibron 1841)
Ptychadena mossambica (Peters 1854)
Ptychadena mutinondoensis Channing & Willem 2018
Ptychadena nana Perret 1980
Ptychadena neumanni (Ahl 1924)
Ptychadena newtoni (Bocage 1886)
Ptychadena nilotica (Seetzen 1855)
Ptychadena nuerensis Smith, Noonan & Colston 2017
Ptychadena obscura (Schmidt & Inger 1959)
Ptychadena oxyrhynchus (Smith 1849)
Ptychadena perplicata Laurent 1964
Ptychadena perreti Guibé & Lamotte 1958
Ptychadena porosissima (Steindachner 1867)
Ptychadena pujoli Lamotte & Ohler 1997
Ptychadena pumilio (Boulenger 1920)
Ptychadena retropunctata (Angel 1949)
Ptychadena schillukorum (Werner 1908)
Ptychadena stenocephala (Boulenger 1901)
Ptychadena straeleni (Inger 1968)
Ptychadena submascareniensis (Guibé & Lamotte 1953)
Ptychadena subpunctata (Bocage 1866)
Ptychadena superciliaris (Günther 1858)
Ptychadena taenioscelis Laurent 1954
Ptychadena tellinii (Peracca 1904)
Ptychadena tournieri (Guibé & Lamotte 1955)
Ptychadena trinodis (Boettger 1881)
Ptychadena upembae (Schmidt & Inger 1959)
Ptychadena uzungwensis (Loveridge 1932)
Ptychadena wadei Largen 2000
Catalogue of Life 26 March 2024
Biota
Animalia
Chordata
Vertebrata
Gnathostomata
Osteichthyes
Sarcopterygii
Tetrapoda
Amphibia
Anura
Ptychadenidae Dubois 1987
Ptychadena Boulenger 1917
Ptychadena amharensis Smith, Noonan & Colston 2017
(this page)
Ptychadena aequiplicata (Werner 1898)
Ptychadena anchietae (Bocage 1868)
Ptychadena ansorgii (Boulenger 1905)
Ptychadena arnei Perret 1997
Ptychadena baroensis Smith, Noonan & Colston 2017
Ptychadena bibroni (Hallowell 1845)
Ptychadena boettgeri (Pfeffer 1893)
Ptychadena broadleyi Stevens 1972
Ptychadena bunoderma (Boulenger 1907)
Ptychadena christyi (Boulenger 1919)
Ptychadena chrysogaster Laurent 1954
Ptychadena cooperi (Parker 1930)
Ptychadena erlangeri (Ahl 1924)
Ptychadena gansi Laurent In Gans, Laurent & Pandit 1965
Ptychadena goweri Smith, Noonan & Colston 2017
Ptychadena grandisonae Laurent 1954
Ptychadena guibei Laurent 1954
Ptychadena harenna Largen 1997
Ptychadena hylaea Schmidt & Inger 1959
Ptychadena ingeri Perret 1991
Ptychadena keilingi (Monard 1937)
Ptychadena levenorum Smith, Noonan & Colston 2017
Ptychadena longirostris (Peters 1870)
Ptychadena mahnerti Perret 1996
Ptychadena mapacha Channing 1993
Ptychadena mascareniensis (Duméril & Bibron 1841)
Ptychadena mossambica (Peters 1854)
Ptychadena mutinondoensis Channing & Willem 2018
Ptychadena nana Perret 1980
Ptychadena neumanni (Ahl 1924)
Ptychadena newtoni (Bocage 1886)
Ptychadena nilotica (Seetzen 1855)
Ptychadena nuerensis Smith, Noonan & Colston 2017
Ptychadena obscura (Schmidt & Inger 1959)
Ptychadena oxyrhynchus (Smith 1849)
Ptychadena perplicata Laurent 1964
Ptychadena perreti Guibé & Lamotte 1958
Ptychadena porosissima (Steindachner 1867)
Ptychadena pujoli Lamotte & Ohler 1997
Ptychadena pumilio (Boulenger 1920)
Ptychadena retropunctata (Angel 1949)
Ptychadena schillukorum (Werner 1908)
Ptychadena stenocephala (Boulenger 1901)
Ptychadena straeleni (Inger 1968)
Ptychadena submascareniensis (Guibé & Lamotte 1953)
Ptychadena subpunctata (Bocage 1866)
Ptychadena superciliaris (Günther 1858)
Ptychadena taenioscelis Laurent 1954
Ptychadena tellinii (Peracca 1904)
Ptychadena tournieri (Guibé & Lamotte 1955)
Ptychadena trinodis (Boettger 1881)
Ptychadena upembae (Schmidt & Inger 1959)
Ptychadena uzungwensis (Loveridge 1932)
Ptychadena wadei Largen 2000
Dynamic Hierarchy Version 2.2.3
Ptychadena amharensis Smith, Noonan & Colston 2017
(this page)
Ptychadena aequiplicata (Werner 1898)
Ptychadena anchietae (Bocage 1868)
Ptychadena ansorgii (Boulenger 1905)
Ptychadena arnei Perret 1997
Ptychadena baroensis Smith, Noonan & Colston 2017
Ptychadena bibroni (Hallowell 1845)
Ptychadena boettgeri (Pfeffer 1893)
Ptychadena broadleyi Stevens 1972
Ptychadena bunoderma (Boulenger 1907)
Ptychadena christyi (Boulenger 1919)
Ptychadena chrysogaster Laurent 1954
Ptychadena cooperi (Parker 1930)
Ptychadena erlangeri (Ahl 1924)
Ptychadena gansi Laurent In Gans, Laurent & Pandit 1965
Ptychadena goweri Smith, Noonan & Colston 2017
Ptychadena grandisonae Laurent 1954
Ptychadena guibei Laurent 1954
Ptychadena harenna Largen 1997
Ptychadena hylaea Schmidt & Inger 1959
Ptychadena ingeri Perret 1991
Ptychadena keilingi (Monard 1937)
Ptychadena levenorum Smith, Noonan & Colston 2017
Ptychadena longirostris (Peters 1870)
Ptychadena mahnerti Perret 1996
Ptychadena mapacha Channing 1993
Ptychadena mascareniensis (Duméril & Bibron 1841)
Ptychadena mossambica (Peters 1854)
Ptychadena mutinondoensis Channing & Willem 2018
Ptychadena nana Perret 1980
Ptychadena neumanni (Ahl 1924)
Ptychadena newtoni (Bocage 1886)
Ptychadena nilotica (Seetzen 1855)
Ptychadena nuerensis Smith, Noonan & Colston 2017
Ptychadena obscura (Schmidt & Inger 1959)
Ptychadena oxyrhynchus (Smith 1849)
Ptychadena perplicata Laurent 1964
Ptychadena perreti Guibé & Lamotte 1958
Ptychadena porosissima (Steindachner 1867)
Ptychadena pujoli Lamotte & Ohler 1997
Ptychadena pumilio (Boulenger 1920)
Ptychadena retropunctata (Angel 1949)
Ptychadena schillukorum (Werner 1908)
Ptychadena stenocephala (Boulenger 1901)
Ptychadena straeleni (Inger 1968)
Ptychadena submascareniensis (Guibé & Lamotte 1953)
Ptychadena subpunctata (Bocage 1866)
Ptychadena superciliaris (Günther 1858)
Ptychadena taenioscelis Laurent 1954
Ptychadena tellinii (Peracca 1904)
Ptychadena tournieri (Guibé & Lamotte 1955)
Ptychadena trinodis (Boettger 1881)
Ptychadena upembae (Schmidt & Inger 1959)
Ptychadena uzungwensis (Loveridge 1932)
Ptychadena wadei Largen 2000
EOL Dynamic Hierarchy 1.1
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Deuterostomia
Chordata
Vertebrata
Gnathostomata
Osteichthyes
Sarcopterygii
Tetrapoda
Lissamphibia
Anura
Ptychadenidae Dubois 1987
Ptychadena Boulenger 1917
Ptychadena amharensis Smith, Noonan & Colston 2017
(this page)
Ptychadena aequiplicata (Werner 1898)
Ptychadena anchietae (Bocage 1868)
Ptychadena ansorgii (Boulenger 1905)
Ptychadena arnei Perret 1997
Ptychadena baroensis Smith, Noonan & Colston 2017
Ptychadena bibroni (Hallowell 1845)
Ptychadena boettgeri (Pfeffer 1893)
Ptychadena broadleyi Stevens 1972
Ptychadena bunoderma (Boulenger 1907)
Ptychadena christyi (Boulenger 1919)
Ptychadena chrysogaster Laurent 1954
Ptychadena cooperi (Parker 1930)
Ptychadena erlangeri (Ahl 1924)
Ptychadena gansi Laurent 1965
Ptychadena goweri Smith, Noonan & Colston 2017
Ptychadena grandisonae Laurent 1954
Ptychadena guibei Laurent 1954
Ptychadena harenna Largen 1997
Ptychadena hylaea Schmidt & Inger 1959
Ptychadena ingeri Perret 1991
Ptychadena keilingi (Monard 1937)
Ptychadena levenorum Smith, Noonan & Colston 2017
Ptychadena longirostris (Peters 1870)
Ptychadena mahnerti Perret 1996
Ptychadena mapacha Channing 1993
Ptychadena mascareniensis (Duméril & Bibron 1841)
Ptychadena mossambica (Peters 1854)
Ptychadena mutinondoensis Channing & Willem 2018
Ptychadena nana Perret 1980
Ptychadena neumanni (Ahl 1924)
Ptychadena newtoni (Bocage 1886)
Ptychadena nilotica (Seetzen 1855)
Ptychadena nuerensis Smith, Noonan & Colston 2017
Ptychadena obscura (Schmidt & Inger 1959)
Ptychadena oxyrhynchus (Smith 1849)
Ptychadena perplicata Laurent 1964
Ptychadena perreti Guibé & Lamotte 1958
Ptychadena porosissima (Steindachner 1867)
Ptychadena pujoli Lamotte & Ohler 1997
Ptychadena pumilio (Boulenger 1920)
Ptychadena retropunctata (Angel 1949)
Ptychadena schillukorum (Werner 1908)
Ptychadena stenocephala (Boulenger 1901)
Ptychadena straeleni (Inger 1968)
Ptychadena submascareniensis (Guibé & Lamotte 1953)
Ptychadena subpunctata (Bocage 1866)
Ptychadena superciliaris (Günther 1858)
Ptychadena taenioscelis Laurent 1954
Ptychadena tellinii (Peracca 1904)
Ptychadena tournieri (Guibé & Lamotte 1955)
Ptychadena trinodis (Boettger 1881)
Ptychadena upembae (Schmidt & Inger 1959)
Ptychadena uzungwensis (Loveridge 1932)
Ptychadena wadei Largen 2000
EOL Dynamic Hierarchy Version 2.2.4
Ptychadena amharensis Smith, Noonan & Colston 2017
(this page)
Ptychadena aequiplicata (Werner 1898)
Ptychadena anchietae (Bocage 1868)
Ptychadena ansorgii (Boulenger 1905)
Ptychadena arnei Perret 1997
Ptychadena baroensis Smith, Noonan & Colston 2017
Ptychadena bibroni (Hallowell 1845)
Ptychadena boettgeri (Pfeffer 1893)
Ptychadena broadleyi Stevens 1972
Ptychadena bunoderma (Boulenger 1907)
Ptychadena christyi (Boulenger 1919)
Ptychadena chrysogaster Laurent 1954
Ptychadena cooperi (Parker 1930)
Ptychadena erlangeri (Ahl 1924)
Ptychadena gansi Laurent In Gans, Laurent & Pandit 1965
Ptychadena goweri Smith, Noonan & Colston 2017
Ptychadena grandisonae Laurent 1954
Ptychadena guibei Laurent 1954
Ptychadena harenna Largen 1997
Ptychadena hylaea Schmidt & Inger 1959
Ptychadena ingeri Perret 1991
Ptychadena keilingi (Monard 1937)
Ptychadena levenorum Smith, Noonan & Colston 2017
Ptychadena longirostris (Peters 1870)
Ptychadena mahnerti Perret 1996
Ptychadena mapacha Channing 1993
Ptychadena mascareniensis (Duméril & Bibron 1841)
Ptychadena mossambica (Peters 1854)
Ptychadena mutinondoensis Channing & Willem 2018
Ptychadena nana Perret 1980
Ptychadena neumanni (Ahl 1924)
Ptychadena newtoni (Bocage 1886)
Ptychadena nilotica (Seetzen 1855)
Ptychadena nuerensis Smith, Noonan & Colston 2017
Ptychadena obscura (Schmidt & Inger 1959)
Ptychadena oxyrhynchus (Smith 1849)
Ptychadena perplicata Laurent 1964
Ptychadena perreti Guibé & Lamotte 1958
Ptychadena porosissima (Steindachner 1867)
Ptychadena pujoli Lamotte & Ohler 1997
Ptychadena pumilio (Boulenger 1920)
Ptychadena retropunctata (Angel 1949)
Ptychadena schillukorum (Werner 1908)
Ptychadena stenocephala (Boulenger 1901)
Ptychadena straeleni (Inger 1968)
Ptychadena submascareniensis (Guibé & Lamotte 1953)
Ptychadena subpunctata (Bocage 1866)
Ptychadena superciliaris (Günther 1858)
Ptychadena taenioscelis Laurent 1954
Ptychadena tellinii (Peracca 1904)
Ptychadena tournieri (Guibé & Lamotte 1955)
Ptychadena trinodis (Boettger 1881)
Ptychadena upembae (Schmidt & Inger 1959)
Ptychadena uzungwensis (Loveridge 1932)
Ptychadena wadei Largen 2000
Frost Darrel R 2018 Amphibian Species of the World
Amphibia
Anura
Ptychadenidae Dubois, 1987
Ptychadena Boulenger, 1917
Ptychadena amharensis Smith, Noonan, and Colston, 2017
(this page)
Ptychadena "mascareniensis"
Ptychadena aequiplicata (Werner, 1898)
Ptychadena anchietae (Bocage, 1868)
Ptychadena ansorgii (Boulenger, 1905)
Ptychadena arnei Perret, 1997
Ptychadena baroensis Smith, Noonan, and Colston, 2017
Ptychadena bibroni (Hallowell, 1845)
Ptychadena boettgeri (Pfeffer, 1893)
Ptychadena broadleyi Stevens, 1972
Ptychadena bunoderma (Boulenger, 1907)
Ptychadena christyi (Boulenger, 1919)
Ptychadena chrysogaster Laurent, 1954
Ptychadena cooperi (Parker, 1930)
Ptychadena erlangeri (Ahl, 1924)
Ptychadena gansi Laurent, 1965
Ptychadena goweri Smith, Noonan, and Colston, 2017
Ptychadena grandisonae Laurent, 1954
Ptychadena guibei Laurent, 1954
Ptychadena harenna Largen, 1997
Ptychadena hylaea Schmidt and Inger, 1959
Ptychadena ingeri Perret, 1991
Ptychadena keilingi (Monard, 1937)
Ptychadena levenorum Smith, Noonan, and Colston, 2017
Ptychadena longirostris (Peters, 1870)
Ptychadena mahnerti Perret, 1996
Ptychadena mapacha Channing, 1993
Ptychadena mascareniensis (Duméril and Bibron, 1841)
Ptychadena mossambica (Peters, 1854)
Ptychadena mutinondoensis Channing and Willem, 2018
Ptychadena nana Perret, 1980
Ptychadena neumanni (Ahl, 1924)
Ptychadena newtoni (Bocage, 1886)
Ptychadena nilotica (Seetzen, 1855)
Ptychadena nuerensis Smith, Noonan, and Colston, 2017
Ptychadena obscura (Schmidt and Inger, 1959)
Ptychadena oxyrhynchus (Smith, 1849)
Ptychadena perplicata Laurent, 1964
Ptychadena perreti Guibé and Lamotte, 1958
Ptychadena porosissima (Steindachner, 1867)
Ptychadena pujoli Lamotte and Ohler, 1997
Ptychadena pumilio (Boulenger, 1920)
Ptychadena retropunctata (Angel, 1949)
Ptychadena schillukorum (Werner, 1908)
Ptychadena stenocephala (Boulenger, 1901)
Ptychadena straeleni (Inger, 1968)
Ptychadena submascareniensis (Guibé and Lamotte, 1953)
Ptychadena subpunctata (Bocage, 1866)
Ptychadena superciliaris (Günther, 1858)
Ptychadena taenioscelis Laurent, 1954
Ptychadena tellinii (Peracca, 1904)
Ptychadena tournieri (Guibé and Lamotte, 1955)
Ptychadena trinodis (Boettger, 1881)
Ptychadena upembae (Schmidt and Inger, 1959)
Ptychadena uzungwensis (Loveridge, 1932)
Ptychadena wadei Largen, 2000
Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Animalia
Bilateria
Deuterostomia
Chordata
Vertebrata
Gnathostomata
Tetrapoda
Amphibia
Anura
Ptychadenidae Dubois 1987
Ptychadena Boulenger 1917
Ptychadena amharensis Smith, Noonan & Colston 2017
(this page)
Ptychadena aequiplicata (Werner 1898)
Ptychadena anchietae (Bocage 1868)
Ptychadena ansorgii (Boulenger 1905)
Ptychadena arnei Perret 1997
Ptychadena baroensis Smith, Noonan & Colston 2017
Ptychadena bibroni (Hallowell 1845)
Ptychadena boettgeri (Pfeffer 1893)
Ptychadena broadleyi Stevens 1972
Ptychadena bunoderma (Boulenger 1907)
Ptychadena christyi (Boulenger 1919)
Ptychadena chrysogaster Laurent 1954
Ptychadena cooperi (Parker 1930)
Ptychadena erlangeri (Ahl 1924)
Ptychadena gansi Laurent In Gans, Laurent & Pandit 1965
Ptychadena goweri Smith, Noonan & Colston 2017
Ptychadena grandisonae Laurent 1954
Ptychadena guibei Laurent 1954
Ptychadena harenna Largen 1997
Ptychadena hylaea Schmidt & Inger 1959
Ptychadena ingeri Perret 1991
Ptychadena keilingi (Monard 1937)
Ptychadena levenorum Smith, Noonan & Colston 2017
Ptychadena longirostris (Peters 1870)
Ptychadena mahnerti Perret 1996
Ptychadena mapacha Channing 1993
Ptychadena mascareniensis (Duméril & Bibron 1841)
Ptychadena mossambica (Peters 1854)
Ptychadena mutinondoensis Channing & Willem 2018
Ptychadena nana Perret 1980
Ptychadena neumanni (Ahl 1924)
Ptychadena newtoni (Bocage 1886)
Ptychadena nilotica (Seetzen 1855)
Ptychadena nuerensis Smith, Noonan & Colston 2017
Ptychadena obscura (Schmidt & Inger 1959)
Ptychadena oxyrhynchus (Smith 1849)
Ptychadena perplicata Laurent 1964
Ptychadena perreti Guibé & Lamotte 1958
Ptychadena porosissima (Steindachner 1867)
Ptychadena pujoli Lamotte & Ohler 1997
Ptychadena pumilio (Boulenger 1920)
Ptychadena retropunctata (Angel 1949)
Ptychadena schillukorum (Werner 1908)
Ptychadena stenocephala (Boulenger 1901)
Ptychadena straeleni (Inger 1968)
Ptychadena submascareniensis (Guibé & Lamotte 1953)
Ptychadena subpunctata (Bocage 1866)
Ptychadena superciliaris (Günther 1858)
Ptychadena taenioscelis Laurent 1954
Ptychadena tellinii (Peracca 1904)
Ptychadena tournieri (Guibé & Lamotte 1955)
Ptychadena trinodis (Boettger 1881)
Ptychadena upembae (Schmidt & Inger 1959)
Ptychadena uzungwensis (Loveridge 1932)
Ptychadena wadei Largen 2000
wikidata_hierarchy tar gz
Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Deuterostomia
Chordata
Olfactores
Craniata
Vertebrata
Gnathostomata
Osteichthyes
Sarcopterygii
Dipnotetrapodomorpha
Tetrapodomorpha
Tetrapoda
Amphibia
Lissamphibia
Anura
Ptychadenidae
Ptychadena
Ptychadena amharensis
(this page)
Ptychadena aequiplicata
Ptychadena anchietae
Ptychadena ansorgii
Ptychadena arnei
Ptychadena baroensis
Ptychadena beka
Ptychadena bibroni
Ptychadena boettgeri
Ptychadena broadleyi
Ptychadena bunoderma
Ptychadena christyi
Ptychadena chrysogaster
Ptychadena cooperi
Ptychadena delphina
Ptychadena doro
Ptychadena erlangeri
Ptychadena filwoha
Ptychadena gansi
Ptychadena goweri
Ptychadena grandisonae
Ptychadena guibei
Ptychadena harenna
Ptychadena hylaea
Ptychadena ingeri
Ptychadena keilingi
Ptychadena largeni
Ptychadena levenorum
Ptychadena longirostris
Ptychadena mahnerti
Ptychadena mapacha
Ptychadena mascareniensis
Ptychadena mossambica
Ptychadena nana
Ptychadena neumanni
Ptychadena newtoni
Ptychadena nilotica
Ptychadena nuerensis
Ptychadena obscura
Ptychadena oxyrhynchus
Ptychadena perplicata
Ptychadena perreti
Ptychadena porosissima
Ptychadena pujoli
Ptychadena pumilio
Ptychadena retropunctata
Ptychadena robeensis
Ptychadena schillukorum
Ptychadena stenocephala
Ptychadena straeleni
Ptychadena submascareniensis
Ptychadena subpunctata
Ptychadena superciliaris
Ptychadena taenioscelis
Ptychadena tellinii
Ptychadena tournieri
Ptychadena trinodis
Ptychadena upembae
Ptychadena uzungwensis
Ptychadena wadei