Satyria orquidiensis Pedraza

Epiphytic shrub, lianoid; stems brown-grey, terete, glabrous, bark smooth; twigs terete, smooth, glabrous. Axillary buds compressed; prophylls 2, valvate, lanceolate, inconspicuous, 2.5–4.5 mm long, margin eciliate, apex acuminate, abaxially puberulous, the hairs inconspicuous, ca. <0.2 mm long, eglandular and unicellular (all indumentum composed of this type of hairs except when indicated). Leaves alternate but apparently pseudoverticillate, originating in clusters of (2–)5–8 leaves separated by leafless sections 11.5–17.5 cm long; petiole terete, pulvinate, 3–6 mm long, glabrate; lamina coriaceous, elliptic, sometimes oblanceolate, (14–)18.7–28 × (3–)5–8 cm, base subcordate, margin entire and eciliate, apex acuminate, adaxially glabrous, abaxially glabrate (but appearing glabrous), the hairs inconspicuous, (< 0.2 mm long), caducous but with persistent bases that are red and swollen, apparently multicellular and glandular; laminar glands inconspicuous, basal, rounded; venation plinerved (acrodromous), suprabasal, with 3–4 visible orders (consistently well-marked up to 3th order) in dry specimens, prominent lateral veins 2(–3) per side, subopposite or alternate, concentrated in the basal third, ascending, brochidodromous, midvein and secondaries adaxially impressed and abaxially raised. Inflorescence a 4–6-flowered raceme, more than one raceme arising from an axillary pad (pincushion like), often cauliflorous; inflorescence and floral bracts alike, persistent, chartaceous, ovate, 0.5–2.6 × 0.8–1.2 mm, margin entire and ciliolate, the hairs inconspicuous and eglandular, apex acute to acuminate, adaxially glabrous, abaxially glabrate, venation obscure; rachis orange, 4–8.5 mm long, glabrate, warts abundant and minute; pedicel dark or lightly orange, articulate with calyx, 1.9–2.3 cm long (3 cm when fruiting), basally (1–1.2 mm diam.) less than half the diameter of apex (3–4.5 mm) (in vivo 2 mm basally, 5–7 mm apically), with the apex becoming discoid and almost as wide as the calyx tube, glabrate, minute warts present at very base both in vivo and when dry; bracteoles 2, basal, supopposite to opposite, chartaceous, ovate, 2–2.4 × 0.6–0.8 mm, margin entire and ciliolate, the hairs inconspicuous, caducous and eglandular, apex acuminate, glabrous on both sides, venation obscure. Flowers 5-merous (some calyces 6-merous), actinomorphic, diplostemonous. Calyx dark or light orange, campanulate, with a marked transversal constriction between limb and tube, inconspicuously 5-angled, the angles alternating with lobes, conspicuously ribbed in the tube and with softer ribs in the limb, 4.1–5.8 mm long (6–7.2 mm in vivo), glabrate, the hairs inconspicuous, <0.1 mm long; tube oblate, 2–2.2 mm long (2.5–3 mm in vivo); limb slightly flaring, 2.1–3.6 mm long (3.5–4.7 mm in vivo); lobes deltate, 1–2 × 3.5–4.5 mm long (1.6–2 × 5 mm in vivo), margin entire, eglandular, and eciliate, apex acute; sinuses practically flat (broadly U-shaped in vivo); aestivation valvate. Corolla basal two thirds orange, apical third green, fleshy, not bistratose, obconic, 3.2–4 cm long (3.4–4.4 cm in vivo), 1–1.2 cm diam. (1–1.1 cm in vivo), 3 mm wide at throat (4.7–5.5 mm in vivo), inconspicuously 5-angled, glabrate without, the hairs inconspicuous, <0.1 mm long, glabrous within; lobes ovate, 1.5–2 × 1–2 mm, but sinuses often further tearing toward end of anthesis and the lobes then oblong and 4.8–7 × 1–2 mm, apex acute, not strongly reflexed at maturity; aestivation valvate. Stamens 10, dimorphic, staminal cycles with different anther lengths and dehiscence orientation, included, not adherent to corolla. Long stamens 10–11.2 mm long; filaments connate into a tube, straight, 3.5–4 mm long, glabrous or distally glabrate on the abaxial side, the hairs a handful, inconspicuous, multicellular and apparently eglandular; anthers 9–9.5 mm long, slightly prognathous, narrowing at base and widening at apex, without a clear distinction between tubules and thecae; thecae 6–6.5 mm long, without basal appendage, minutely papillate at least basally; tubules 2, free, turned inwards like bull’s horns, 3 mm long, smooth, dehiscing by latrorse elliptical slits 3 mm long, distal margin ornamented with small and irregular lobes. Short stamens 9–10 mm long, same shape, indumentum and features as long stamens except when indicated; filaments 3.5–4 mm long; anthers 8–8.5 mm long; thecae 5.5 mm long; tubules pointing upwards and without space between them, 2.5–3 mm long, dehiscing by introrse elliptical slits 2.5–3 mm long. Nectary not pulvinate, slightly concave to flat, glabrous; style white, 2.6–3.7 cm long, included; stigma punctiform. Berry cream-colored and ribbed when immature, turning purple at maturity.