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Canyon Liveforever

Dudleya cymosa subsp. cymosa

Comprehensive Description

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Dudleya gigantea Rose, Bull. N. Y. Gard. 3 : 23. 1903
Acaulescent or nearly so, with a thick short rootstock. Basal leaves in a dense rosette, very glaucous, oblanceolate, 5-7 cm. long, drying very thin ; flowering stems rather stout, about 3 cm. long ; inflorescence paniculate, with numerous usually erect branches ; pedicels rather slender, 5-10 mm. long, erect ; calyx-lobes broadly ovate, obtuse or acutish ; corolla drying deep-red, 9-10 mm. long, the segments erect, acute, united at base into a short tube 1.5 mm. long.
Type locality : New York Falls, Amador County, California. Distribution : Known only from the type locality.
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bibliographic citation
John Kunkel SmaII, George Valentine Nash, Nathaniel Lord Britton, Joseph Nelson Rose, Per Axel Rydber. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(1). New York Botanical Garden, New York, NY
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Comprehensive Description

provided by North American Flora
Dudleya nevadensis (S. Wats.) Britton & Rgse, Bull. N. Y
Bot. Gard. 3 : 20. 1903,
Cotyledon nevadensis S. Wats. Bot. Calif. 1 : 212. 1876.
Acaulescent, green, scarcely glaucous. Basal leaves obovate to oblanceolate-spatulate,
acute or acuminate, 6-10 cm. long, 3-5 cm. wide, narrowest at the base; leaves of flowering
branches lanceolate to oblong-lanceolate, 1-3 cm. long, acute, sessile or slightly clasping;
flowering branches 1-2 dm. high; cyme compound, rather dense, 5-8 cm. broad; pedicels
ra,ther stout, 6-12 mm. long, mostly longer than the flowers ; flower-buds short-ovoid, acute,
little longer than thick ; calyx-segments triangular-ovate-lanceolate, acute, 3^ mm. long,
glaucous ; corolla 10-12 mm. long, yellow tinged with red, cleft to the middle or below.
Type locality : Sonora, California. Distribution : Mountains of middle California.
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bibliographic citation
John Kunkel SmaII, George Valentine Nash, Nathaniel Lord Britton, Joseph Nelson Rose, Per Axel Rydber. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(1). New York Botanical Garden, New York, NY
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Comprehensive Description

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Dudleya angustiflora Rose, Bull. N. Y. Bot. Gard. 3 : 14. 1903
Acaulescent. Leaves forming a rosette, lanceolate, 3-4 cm. long, acute or acuminate, very glaucous; flowering stems slender, 1.5-2 dm. high; inflorescence a somewhat spreading panicle with 2-4 elongated branches ; pedicels slender, 4-12 mm. long ; calyx glaucous, deeply 5-parted ; its lobes broadly lanceolate, acute, 3 mm. long ; corolla very narrow, 10-12 mm. long, reddish (at least drying so), its segments united into a tube 2 mm. long ; stamens about two-thirds as long as the corolla, borne on its tube ; carpels erect.
Type locality : Daunt, Tulare County, California. Distribution : Known only from the type locality.
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bibliographic citation
John Kunkel SmaII, George Valentine Nash, Nathaniel Lord Britton, Joseph Nelson Rose, Per Axel Rydber. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(1). New York Botanical Garden, New York, NY
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Comprehensive Description

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Dudleya plattiana
Add the synonym: Echeveria Plattiana Nelson & Macbr. Bot. Gaz. 56: 477. 1913.
Add: Illustrations: Meehan, Nat. Fl. II. 2: pi. 27; Armstrong, Field Book W. Wild Fl. 195.
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bibliographic citation
Per Axel Rydberg. 1918. ROSACEAE (conclusio). North American flora. vol 22(6). New York Botanical Garden, New York, NY
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Comprehensive Description

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Dudleya plattiana (Jepson) Britton & Rose, Bull. N. Y
Bot. Gard. 3 : 20. 1903.
Cotyledon Plattiana Jepson, Fl. West. Middle Calif. 267. 1901.
Acaulescent. Leaves somewhat glaucous, oblanceolate, 7-10 cm. long, narrowed at base, acute or acuminate; flowering stems 12-20 cm. long; stem-leaves lanceolate, rather narrow at base ; inflorescence rather short, more or less flat-topped ; pedicels slender, 4-10 mm. long; calyx-lobes triangular, acute or acuminate; corolla-segments lanceolate, acute, 8-10 mm. long, reddish or dr5dng so.
Type locality : Vaca Mountains, California. Distribution : Inner coast range,, central California.
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cc-by-nc-sa-3.0
bibliographic citation
John Kunkel SmaII, George Valentine Nash, Nathaniel Lord Britton, Joseph Nelson Rose, Per Axel Rydber. 1905. ROSALES, PODOSTEMONACEAE, CRASSULACEAE, PENTHORACEAE and PARNASSIACEAE. North American flora. vol 22(1). New York Botanical Garden, New York, NY
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Dudleya cymosa subsp. cymosa

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Dudleya cymosa subsp. cymosa is a species of succulent perennial plant in the family Crassulaceae native to California. It is the autonymous subspecies for Dudleya cymosa, and is known by the common name canyon liveforever. It is native to the California Coast Ranges, the Sierra Nevada and the Santa Monica Mountains. It is characterized by bright-yellow, orange or red flowers and broad, wide leaves. This plant is commonly found growing on rocky outcrops, talus slopes, and in shaded canyons.

Description

A solitary or few-branched rosette forming succulent with broad leaves and bright-yellow, orange, or red flowers.

Vegetative morphology

Each basal rosette is formed on the apex of a stem, known as a caudex. The caudex is short, usually less than 5 cm long, and is 1 to 3.5 cm in diameter. It is typically unbranched, forming a solitary rosette, but may less commonly branch into a few rosettes. Each of the basal rosette is 6 to 20 cm in diameter, composed of 6 to 25 leaves. The leaves are typically 3 to 17 cm long, 10 to 60 mm wide, and 1 to 5 mm thick. The leaves are shaped ovate or deltate to oblanceolate or spatulate (shaped like a spoon). The margin (edge) of the leaves is often folded upward at the widest point, and the tip of the leaf is more or less recurved and is shaped acuminate to mucronate.[1][2]

Reproductive morphology

Detail of the flowers

The inflorescence is typically asymmetrical radially, because the pedicels turn to the sun or away from the cliff the plant is on. The peduncle (floral stem) is 5 to 45 cm tall, 2 to 8 mm wide, and ascending. On the inflorescence are 7 to 20 bracts, positioned horizontally to ascending, and shaped ovate to triangular-lanceolate, with the tips acute to acuminate. The lowermost bracts are 0.5 to 3 cm long and 5 to 15 mm wide. The inflorescence branches 2 to 4 times, and then may subsequently branch 0 to 3 times. The terminal branches are circinate when young (spiral-shaped, like a fern frond unfurling), but are ascending in age, 1 to 17 cm long and with 2 to 10 flowers. Suspending the flowers are the pedicels, which are erect, the lowermost 5 to 15 mm long.[1][2]

The calyx of the flowers is 3 to 7 mm long, and 2.5 to 6 mm wide. The calyx lobes (equivalent to sepals) are triangular to triangular ovate, 1.5 to 5 mm long. The corolla is cylindrical in anthesis, with the tips of the petals spreading from 45 to 90 degree angles. The petals are colored yellow, orange, or red, and may be glaucous along their midrib. The petals are shaped elliptic to narrowly lanceolate, with an acute tip, 7 to 15 mm long and 2 to 4 mm wide, connate (fused to form a tube) 1 to 2.5 mm.[1][2]

Taxonomy

Cladogram of Dudleya cymosa subsp. cymosa Virens clade

D. abramsii subsp. setchellii

D. cymosa subsp. paniculata

D. cymosa subsp. cymosa

Taxonomic history

The original description is from 1858 by Charles Antoine Lemaire, in the 10th issue of L'Illustration Horticole, as Echeveria cymosa. The type locality is probably California, as the description simply states "Californie? Třes distincte!" As there was no authentic type specimen, the neotype for the species is an illustration by John Gilbert Baker of Cotyledon cymosa in Refugio Botanicum.[1]

The illustration by John Gilbert Baker which defines the species Dudleya cymosa.

In their work on North American Crassulaceae, Nathaniel Lord Britton and Joseph Nelson Rose placed Echeveria cymosa as Dudleya cymosa. With the recognition of infraspecific taxa, it became Dudleya cymosa subsp. cymosa. Reid Moran in the 1950s recognized a larger subspecies of Dudleya cymosa known as subsp. gigantea based on its larger size, but additional data proved this subspecies invalid, as the supposed larger size was not evident after numerous collections and data from cultivation.[1]

Phylogenetics

This plant is a diploid, with a chromosome number of n = 17, the base number for Dudleya. Phylogenetic analysis has placed this plant with the closely related to Dudleya cymosa subsp. paniculata and Dudleya abramsii subsp. setchellii. The other subspecies of Dudleya cymosa are not closely related.[3]

Distribution and habitat

In habitat

This species is endemic to California. It is distributed throughout the California Coast Ranges, reaching a northern extent in Humboldt County, also distributed in the Sierra Nevada, and in the Santa Monica Mountains. It is found in rocky outcroppings, talus slopes, and shaded canyon slopes.[2]

References

  1. ^ a b c d e Nakai, Kei M. (1987). "Some New and Reconsidered California Dudleya (Crassulaceae)". Madroño. 34 (4): 335–338.
  2. ^ a b c d McCabe, Stephen W. (2012). "Dudleya cymosa subsp. cymosa". Jepson eFlora. Jepson Flora Project. Archived from the original on 2017-08-29. Retrieved 30 December 2021.
  3. ^ Yost, J. M.; Bontrager, M.; McCabe, S. W.; Burton, D.; Simpson, M. G.; Kay, K. M.; Ritter, M. (2013). "Phylogenetic relationships and evolution in Dudleya (Crassulaceae)" (PDF). Systematic Botany. 38 (4): 1096–1104. doi:10.1600/036364413X674760. S2CID 15715233.
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Dudleya cymosa subsp. cymosa: Brief Summary

provided by wikipedia EN

Dudleya cymosa subsp. cymosa is a species of succulent perennial plant in the family Crassulaceae native to California. It is the autonymous subspecies for Dudleya cymosa, and is known by the common name canyon liveforever. It is native to the California Coast Ranges, the Sierra Nevada and the Santa Monica Mountains. It is characterized by bright-yellow, orange or red flowers and broad, wide leaves. This plant is commonly found growing on rocky outcrops, talus slopes, and in shaded canyons.

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