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Estòmids ( Catalan; Valencian )

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Els estòmids (Stomiidae) són una família de peixos marins pertanyent a l'ordre dels estomiformes.[1]

Alimentació

Són depredadors que mengen peixos més petits[2] a les zones abissals.

Distribució geogràfica

Es troba als oceans Índic, Atlàntic i Pacífic.[3]

Gèneres i espècies

Referències


Bibliografia


Enllaços externs

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Estòmids: Brief Summary ( Catalan; Valencian )

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Světlonošovití ( Czech )

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Světlonošovití (Stomiidae) je čeleď dravých paprskoploutvých ryb, obývajících oceány po celém světě v hloubkách od tří set do pěti tisíc metrů. Zahrnuje šest podčeledí, 27 rodů a asi 270 druhů. Světlonošovité ryby mají vesměs protáhlé tělo převážně černé barvy bez šupin a relativně velkou hlavu s dlouhými ostrými zuby, dosahují maximální délky okolo půl metru. Jsou vybaveny fotofory, které jim umožňují v temném prostředí přilákat kořist. Díky volnému spojení týlní kosti s prvním obratlem mohou otevřít ústa pod úhlem až 120 stupňů, což jim umožňuje spolknout i značně velké živočichy. Jejich hlavním predátorem je vorvaň obrovský.

Seznam rodů

  • Aristostomias (pestrosvítivec) Zugmayer, 1913
  • Astronesthes (svítivec) Richardson, 1845
  • Bathophilus (svítivka) Giglioli, 1882
  • Borostomias (svítivec) Regan, 1908
  • Chauliodus (zubatka) Bloch & Schneider, 1801
  • Chirostomias Regan & Trewavas, 1930
  • Echiostoma Lowe, 1843
  • Eupogonesthes Parin & Borodulina, 1993
  • Eustomias Vaillant, 1888
  • Flagellostomias Parr, 1927
  • Grammatostomias Goode & Bean, 1896
  • Heterophotus Regan & Trewavas, 1929
  • Idiacanthus Peters, 1877
  • Leptostomias Gilbert, 1905
  • Malacosteus (pestrosvítivec) Ayres, 1848
  • Melanostomias Brauer, 1902
  • Neonesthes Regan & Trewavas, 1929
  • Odontostomias Norman, 1930
  • Opostomias Günther, 1887
  • Pachystomias Günther, 1887
  • Photonectes Günther, 1887
  • Photostomias (pestrosvítivec) Collett, 1889
  • Rhadinesthes Regan & Trewavas, 1929
  • Stomias Cuvier, 1816
  • Tactostoma Bolin, 1939
  • Thysanactis Regan & Trewavas, 1930
  • Trigonolampa Regan & Trewavas, 1930

Externí odkazy

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Světlonošovití: Brief Summary ( Czech )

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 src= Pestrosvítivec černý  src= Zubatka Sloanova  src= Svítivec čertovský

Světlonošovití (Stomiidae) je čeleď dravých paprskoploutvých ryb, obývajících oceány po celém světě v hloubkách od tří set do pěti tisíc metrů. Zahrnuje šest podčeledí, 27 rodů a asi 270 druhů. Světlonošovité ryby mají vesměs protáhlé tělo převážně černé barvy bez šupin a relativně velkou hlavu s dlouhými ostrými zuby, dosahují maximální délky okolo půl metru. Jsou vybaveny fotofory, které jim umožňují v temném prostředí přilákat kořist. Díky volnému spojení týlní kosti s prvním obratlem mohou otevřít ústa pod úhlem až 120 stupňů, což jim umožňuje spolknout i značně velké živočichy. Jejich hlavním predátorem je vorvaň obrovský.

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Barten-Drachenfische ( German )

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Die Barten-Drachenfische oder Schuppendrachenfische (Stomiidae) sind eine mit über 270 Arten in 28 Gattungen in allen Ozeanen, einschließlich des Mittelmeers, verbreitete Familie von Tiefseefischen. Zu ihnen gehören (nach traditioneller Auffassung) die bekannten Viperfische (Chauliodus).

Merkmale

Die Tiere haben einen langgestreckten Körper, Kopf und Maul sind groß. Die ungleichen Zähne sind dolchförmig, rückwärts gekrümmt und ragen bei geschlossenem Maul vor. Die Schuppen fallen leicht ab und sind dunkel, braun oder schwarz. Die Tiere besitzen Leuchtorgane, teilweise auch in der Mund- und Kiemenhöhle. Die Tiere werden 4 bis 50 cm lang. Sie können auch (im Verhältnis zu ihren eigenen Ausmaßen) sehr große Beute, z. B. Sardinen, überwältigen. Die Gattungen Aristostomias, Eustomias, Malacosteus, Pachystomias und Photostomias können dazu ihren Kopf weiter zurücklegen als alle anderen Knochenfische und ihr Maul bis zu einem Winkel von 120° aufreißen. Ermöglicht wird dies durch ein zusätzliches, knorpeliges Gelenk zwischen Hinterkopf und dem ersten Wirbel.[1]

Nach aktuellen phylogenetischen Untersuchungen[2] sind die Barten-Drachenfische kein Monophylum, denn die Viperfische sind näher mit der Familie Tiefsee-Beilfische verwandt als mit den übrigen untersuchten Barten-Drachenfischen. Auch die hier vorgenommene Unterteilung in Unterfamilien scheint in dieser Form keinen Bestand zu haben.

Literatur

Einzelnachweise

  1. N. K. Schnell, G. D. Johnson: Evolution of a Functional Head Joint in Deep-Sea Fishes (Stomiidae). In: PLOS ONE. Band 12, Nr. 2, Februar 2017, Art. e0170224. doi:10.1371/journal.pone.0170224
  2. Ricardo Betancur-R, Edward O. Wiley, Gloria Arratia, Arturo Acero, Nicolas Bailly, Masaki Miya, Guillaume Lecointre, Guillermo Ortí: Phylogenetic classification of bony fishes. In: BMC Evolutionary Biology. Juli 2017, S. 162. doi:10.1186/s12862-017-0958-3
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Barten-Drachenfische: Brief Summary ( German )

provided by wikipedia DE

Die Barten-Drachenfische oder Schuppendrachenfische (Stomiidae) sind eine mit über 270 Arten in 28 Gattungen in allen Ozeanen, einschließlich des Mittelmeers, verbreitete Familie von Tiefseefischen. Zu ihnen gehören (nach traditioneller Auffassung) die bekannten Viperfische (Chauliodus).

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Stomiidae

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Stomiidae is a family of deep-sea ray-finned fish, including the barbeled dragonfishes. They are quite small, usually around 15 cm, up to 26 cm. These fish are apex predators and have enormous jaws filled with fang-like teeth.[1] They are also able to hinge the neurocranium and upper-jaw system, which leads to the opening of the jaw to more than 100 degrees.[1] This ability allows them to consume extremely large prey, often 50% greater than their standard length.[1]

Habitat

The family Stomiidae can be found in all oceans. They also exist at a wide range of depths between the surface and thousands of meters deep into the bathypelagic zone,[2] depending on the water's ideal feeding and breeding conditions. There is also some evidence that certain species within the family Stomiidae exhibit migratory behavior. Temperature, salinity, oxygen, and fluorescence profiles of an area can impact some species' (like Sloane's viperfish Chauliodus sloani) preferred habitat changes from day to night with DVM.[3]

Brian Coad, ichthyologist from the Canada Museum of Nature once observed that there are "64 [species of Dragonfishes] reported from Canada, 5 of which reach the Arctic ''. These species are most commonly found in the mesopelagic to bathypelagic regions at a depth of 1000m-4000m, and in the Arctic, most samples of these species have been captured along the Davis Strait. The average temperature in these waters is approximately 3-4°C [4] Some examples of species discovered in that region are: Astronesthes cf. richardsoni; Borostomia antarcticus; Chauliodus sloani; Malacosteus niger; Rhadinesthes decimus; Stomias boa. [4]

Species of Antarctic dragonfish are found in the Southern Ocean. There are 16 species in the Antarctic, all belonging to the suborder Notothenioidei.[2] Two species in this region that are currently generating interest in further scientific study are sister species Acanthodraco dewitti and Psilodraco breviceps.[2]

Features

It is one of the many species of deep-sea fish that can produce their own light through a chemical process known as bioluminescence.[5] A special organ known as a photophore helps produce this light. The deep-sea dragonfishes have large heads, and mouths equipped with many sharp fang-like teeth. They have a long stringlike structure known as barbel, with a light-producing photophore at the tip, attached to their chin. They also have photophores attached along the sides of their body. A specific species of Stomiidae, the Chauliodus, cannot luminescence longer than 30 minutes without adrenaline. However, in presence of adrenaline, it can produce light for many hours.[6] They produce blue-green light, the wavelengths of which can travel the farthest in the ocean. The deep-sea dragonfish waves its barbel back and forth and produces flashing lights on and off to attract prey and potential mates. Many of the species they prey upon also produce light themselves, which is why they have evolved to have black stomach walls to keep the lights concealed while digesting their meal in order to stay hidden from their predators.

Jaw Morphology

The jaw of members in the stomiidae family is adapted extremely well for survival and predation in the deep sea. Although small in size, the dragonfish jaw is adapted to capture large prey that are up to 50% the body mass of themselves.[7] The long "loosejaw" of the dragonfish exhibits increased resistive forces to lower jaw adduction compared to fish with shorter jaws; however, due to decreased surface are of the lower jaw, dragonfish are able to lower the mechanical advantage of adduction and increase adduction velocity through the reduction of resistive forces. Additionally, it is seen that the adductor mass of the lower jaw of deep-sea dragonfish is significantly decreased, allowing for increased ability to attain high adduction velocity.[8] This makes the deep-sea dragonfish significantly more competitive when hunting for prey due to its ability to capture large prey quickly and efficiently.

An important distinction in jaw morphology between an adult dragonfish and its larvae is the shape of the mouth. The adult fish have a elongated snout-like face with a protruding jaw, while the larvae have a rounder shaped mouth and a lower jaw that does not protrude.[9]

Additionally, members of this family have a unique head joint that contribute to its ability to open its 'loosejaw' so wide. Deep-sea dragonfish have a flexible connection between the base of the skull and first vertebrae called the occipito-vertebral gap where only the flexible notochord is present.[10][11][12] This gap is a result of either vertebral reduction or notochord elongation.[11] Functionally, the gap allows deep-sea dragonfish to pull back their cranium and open their mouths up to 120°, which is significantly farther than other taxa that lack such a head joint.[10] This is what allows deep-sea dragonfish to engulf such large prey, resulting in improved survive through the ability to consume more organisms in an extremely food limited environment.

On top of an extremely well adapted jaw, members of the stomiidae family also have teeth that are adapted for hunting in deep sea. Their teeth are sharp, hard, stiff, and transparent when wet,[7][13] making their teeth dangerous weapons as these teeth become basically invisible in the light absent deep sea. This means the refraction index of their teeth is nearly identical to that of the sea water they inhabit.[7] The transparency is due to a nanoscale structure of hydroxyapatite and collagen, while the tips of the transparent teeth of deep-sea dragonfish were found to emit more red light in seawater[13] which further contributes to its transparency as red light is close to invisible at the depths that the deep-sea dragonfish reside due to a lack of light penetration.

Evolution of Sensory Organs

The deep-sea dragonfishes are part of the stomiidae family, making up a clade of 28 genera and 290 species. The dragonfish possess unique adaptations to help them thrive in the deepest parts of the ocean. This family species have been discovered to use certain long-wave and short-wave bioluminescence to communicate, lure prey, distract predators, and camouflage themselves.[14] The stomiidae family has many unique adaptations to their sensory organs for the deep sea. Most deep-sea organisms have only a single visual pigment sensitive to the absorbance ranges of 470-490 nm.[15] This type of optical system is commonly found in the stomiidae family. However, three genera of dragonfish evolved the ability to produce both long-wave and short-wave bioluminescence.[16] In addition, deep-sea dragon fishes evolved retinas with far-red emitting photophores and rhodopsins.[14] These far-red emitting properties produce long-wave bioluminescence greater than 650 nm. This unique evolutionary trait was first seen around 15.4 Ma and had a single evolutionary origin within the stomiidae family.[14]

Reproductive Features

Dragonfish females exhibit two distinct cohorts oocytes, one which is a white cream color during the first growing stage and the other which is orange-reddish in vitellogenesis. The orange-reddish ovaries are released in the current spawning season, while the other batch is in the growing stage.[17] Stomiids are gonochoristic, allowing them to increase their reproductive fitness by using their energy to produce gametes instead of reconfiguring the reproductive system. The female adult stomiids are also larger than the males.[18]

Behavior

Dragonfish are a type of teleost fish that inhabit the deep sea and use bioluminescence to detect prey and communicate with potential mates. They possess far-red emitting photophores and rhodopsins that are sensitive to long-wave emissions greater than 650 nm, and have adapted to the unique light conditions of the deep-sea environment.[14]

Reproductive Behavior

Egg-laying, which predominantly occurs in October, is preceded by a distinctive whirling behavior driven by the male prodding the side of the female's abdomen.[9] Additionally, dragonfish possess a unique adaptation of being able to see using chlorophyll in their eyes, which may allow them to detect the weak bioluminescence of their prey and navigate their dark habitats more effectively. This research sheds light on the reproductive behavior and early life stages of the naked dragonfish and contributes to our understanding of the ecology and behavior of dragonfish species.

Dragonfish also display a parental care behavior, where they guard their nest, staying within close proximity and resting on its pelvic fins. This guarding behavior has been documented in all the major clades of Antarctic notothenoids, except Artedidraconidae.[19]

Evolution and Adaptations of the Visual System

One study focuses on the stomiid family, which includes loosejaws and dragonfishes, analyzing the genetic makeup of the visual pigments in these fish and how they have adapted to the unique light conditions of the deep-sea environment. The research helps us understand how dragonfish behavior and vision have evolved to allow them to thrive in the deep sea. Dragonfish use far-red emitting photophores and rhodopsins to detect prey and navigate their habitats.[14] Additionally, dragonfish use chlorophyll in their eyes to detect the weak bioluminescence of their prey, which is an unusual adaptation for a vertebrate.[20]

Visual Communication and Behavior

Teleost fishes exhibit a wide range of visual signals, including color, texture, form, and motion, that are used to find mates, establish dominance, defend territory, and coordinate group behavior. Dragonfish have specialized bioluminescent organs that produce red light to communicate with potential mates and prey.[21] Understanding the visual communication and behavior of teleost fishes is essential to understanding the behavior of dragonfish in their natural habitats.

Overview of Bioluminescence in Stomiidae

Dragonfish of the Stomiidae family are largely characterized by their bioluminescent barbels, which act as lures for prey and are a species-specific structure.[22] These barbels extend anteriorly off the bottom jaw, and prey attracted to its bioluminescence include lanternfish and bristlemouths.[5] It is proposed that the specificity of bioluminescent barbel structure to certain species allows for advantageous same-species recognition that promotes genetic isolation, in addition to allowing scientists to more easily identify distinct species due to anatomical barbel differences.[23] The diversity of Stomiidae species is exceptional for their clade age thanks largely to the species-specific barbels.[23] Further, sexual dimorphism of bioluminescence in dragonfish contributes to even greater diversity within the species, but the greater abundance of immature specimens within research collections makes studying sexual dimorphism challenging.[22]

A red photophore is visible in the suborbital region of this Malacosteus rendering.

In addition to a bioluminescent barbel, members of the Stomiidae family have a blue light emitting photophore in the postorbital region.[24] Some dragonfish, such as the Malacosteus niger, also have a unique red light emitting photophore in the suborbital region.[24] It is thought that the mechanism of red bioluminescence produced by the suborbital photophore is facilitated by energy transmission and is chemically similar to the blue bioluminescence of the barbel.[24] While suborbital photophores that emit red bioluminescence are particularly helpful for finding prey, since many organisms in the deep sea can only see blue light, it appears as though this red light emission by dragonfish is not directly associated with prey choice, and it is thus hypothesized that it may be used for intraspecific communication.[24] This raises an interesting question of to what extent the red bioluminescence determines dragonfish prey choice.

Lure Bioluminescence

Species of the Stomiidae family use blue bioluminescence for communication, camouflage, and as a luring mechanism.[25] They emit shortwave blue bioluminescence from postorbital photophores and from a long, slender appendage on the chin, called the barbel.[26] The shaft of the barbel is composed of cylindrical muscles, blood vessels and nervous fibers, and the bulb of the barbel has a single photophore.[27] The catecholamine adrenaline is found in the connective tissue within the stem.[28] One hypothesis regarding barbel control is that adrenaline innervation may control both the movement of the barbel and its production of bioluminescence. Data from a study performed on specimens of the Stomias boa species agree with this hypothesis because the barbels of the dragonfish produced light emissions following exposure to external adrenaline.[28]

The loose jaw dragonfishes, which include species from Aristostomias, Malacosteus, and Pachystomias, have the ability to detect and produce red bioluminescence.[29] This is made possible by far-red emitting photophores located under the eye and rhodopsins that are sensitive to long-wave emissions.[30] This red bioluminescence is used to illuminate prey and to detect other far-red dragonfishes, because it goes undetected by most other species.[30] The species with far-red emitting photophores differ in morphology and behavior from most other dragonfish species. For example, the barbels of these species are more simple in structure than those of other dragonfishes.[29] They also differ in foraging strategies. While most dragonfishes that produce shortwave blue bioluminescence undergo regular diel vertical migrations, this is not seen in those with far-red emissions. The foraging strategy they undergo involves remaining in the deep-sea and emitting far-red bioluminescence to illuminate a small area and search for prey.[29] Although Malacosteus, Pachystomias, and Aristostomias all have suborbital photophores that produce red bioluminescence, there are differences in the suborbital photophores between these three genera, in their shape, color, flash duration, and maximum emission.[31]

Representative species gallery

References

  1. ^ a b c Kenaley, Christopher P. (2012). "Exploring Feeding Behavior In Deep-sea Dragonfishes (Teleostei: Stomiidae): Jaw Biomechanics and Functional Significance of a Loosejaw". Biological Journal of the Linnean Society. 106 (1): 224–240. doi:10.1111/j.1095-8312.2012.01854.x.
  2. ^ a b c Desvignes, T.; Postlethwait, J. H.; Konstantinidis, P. (2020). "Biogeography of the antarctic dragonfishes Acanthodraco dewitti and Psilodraco breviceps with re-description of Acanthodraco dewitti (Notothenioidei: Bathydraconidae)". Polar Biology. 43 (5): 565–572.
  3. ^ Eduardo, L. N.; Lucena-Frédou, F.; Mincarone, M. M.; Soares, A.; Le Loc'h, F.; Frédou, T.; Ménard, F.; Bertrand, A. (2020). "Trophic ecology, habitat, and migratory behaviour of the viperfish Chauliodus sloani reveal a key mesopelagic player". Scientific Reports. 10 (1).
  4. ^ a b Coad, B.; Reist, J. (2017). Marine Fishes of Arctic Canada. Toronto: Toronto: University of Toronto Press. pp. Chapter 25 "Stomiidae: Dragonfishes".
  5. ^ a b Carmichael, Stephen W. (November 2019). "It Can Bite You Even if You Can't See It". Microscopy Today. 27 (6): 8–10. doi:10.1017/S1551929519001056. ISSN 1551-9295.
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  7. ^ a b c Gilbert, Pupa U. P. A.; Stifler, Cayla A. (2019-07-10). "See-Through Teeth, Clearly". Matter. 1 (1): 27–29. doi:10.1016/j.matt.2019.06.015. ISSN 2590-2385.
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  13. ^ a b Velasco-Hogan, Audrey; Deheyn, Dimitri D.; Koch, Marcus; Nothdurft, Birgit; Arzt, Eduard; Meyers, Marc A. (July 2019). "On the Nature of the Transparent Teeth of the Deep-Sea Dragonfish, Aristostomias scintillans". Matter. 1 (1): 235–249. doi:10.1016/j.matt.2019.05.010.
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  16. ^ Douglas, R. H.; Mullineaux, C. W.; Partridge, J. C. (2000-09-29). Collin, S.P.; Marshall, N.J. (eds.). "Long–wave sensitivity in deep–sea stomiid dragonfish with far–red bioluminescence: evidence for a dietary origin of the chlorophyll–derived retinal photosensitizer of Malacosteus niger". Philosophical Transactions of the Royal Society of London. Series B: Biological Sciences. 355 (1401): 1269–1272. doi:10.1098/rstb.2000.0681. ISSN 0962-8436. PMC 1692851. PMID 11079412.
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  18. ^ Marks, Alex D.; Kerstetter, David W.; Wyanski, David M.; Sutton, Tracey T. (2020). "Reproductive Ecology of Dragonfishes (Stomiiformes: Stomiidae) in the Gulf of Mexico". Frontiers in Marine Science. 7. doi:10.3389/fmars.2020.00101. ISSN 2296-7745.
  19. ^ Barrera-Oro, Esteban R.; Lagger, Cristian (2010-11-01). "Egg-guarding behaviour in the Antarctic bathydraconid dragonfish Parachaenichthys charcoti". Polar Biology. 33 (11): 1585–1587. doi:10.1007/s00300-010-0847-3. ISSN 1432-2056.
  20. ^ Douglas, R. H.; Partridge, J. C.; Dulai, K.; Hunt, D.; Mullineaux, C. W.; Tauber, A. Y.; Hynninen, P. H. (June 1998). "Dragon fish see using chlorophyll". Nature. 393 (6684): 423–424. doi:10.1038/30871. ISSN 1476-4687.
  21. ^ Marshall, N. J. "Communication Behavior: Visual Signals" (PDF).
  22. ^ a b Herring, Peter J. (August 2007). "REVIEW. Sex with the lights on? A review of bioluminescent sexual dimorphism in the sea". Journal of the Marine Biological Association of the United Kingdom. 87 (4): 829–842. doi:10.1017/S0025315407056433. ISSN 1469-7769.
  23. ^ a b Davis, Matthew P.; Holcroft, Nancy I.; Wiley, Edward O.; Sparks, John S.; Leo Smith, W. (2014-05-01). "Species-specific bioluminescence facilitates speciation in the deep sea". Marine Biology. 161 (5): 1139–1148. doi:10.1007/s00227-014-2406-x. ISSN 1432-1793. PMC 3996283. PMID 24771948.
  24. ^ a b c d Herring, Peter J.; Cope, Celia (2005-09-28). "Red bioluminescence in fishes: on the suborbital photophores of Malacosteus, Pachystomias and Aristostomias". Marine Biology. 148 (2): 383–394. doi:10.1007/s00227-005-0085-3. ISSN 0025-3162.
  25. ^ Kenaley, Christopher P.; DeVaney, Shannon C.; Fjeran, Taylor T. (2014-01-30). "THE COMPLEX EVOLUTIONARY HISTORY OF SEEING RED: MOLECULAR PHYLOGENY AND THE EVOLUTION OF AN ADAPTIVE VISUAL SYSTEM IN DEEP-SEA DRAGONFISHES (STOMIIFORMES: STOMIIDAE)". Evolution. 68 (4): 996–1013. doi:10.1111/evo.12322. ISSN 0014-3820.
  26. ^ "Splendor in the Dark". Discover Magazine. Retrieved 2023-04-12.
  27. ^ Herring, Peter J.; Cope, Celia (2005-12-01). "Red bioluminescence in fishes: on the suborbital photophores of Malacosteus, Pachystomias and Aristostomias". Marine Biology. 148 (2): 383–394. doi:10.1007/s00227-005-0085-3. ISSN 1432-1793.
  28. ^ a b Mallefet, Jérôme; Duchatelet, Laurent; Hermans, Claire; Baguet, Fernand (2019-01-01). "Luminescence control of Stomiidae photophores". Acta Histochemica. 121 (1): 7–15. doi:10.1016/j.acthis.2018.10.001. ISSN 0065-1281. PMID 30322809. S2CID 53505749.
  29. ^ a b c Kenaley, Christopher P.; DeVaney, Shannon C.; Fjeran, Taylor T. (2014-01-30). "THE COMPLEX EVOLUTIONARY HISTORY OF SEEING RED: MOLECULAR PHYLOGENY AND THE EVOLUTION OF AN ADAPTIVE VISUAL SYSTEM IN DEEP-SEA DRAGONFISHES (STOMIIFORMES: STOMIIDAE)". Evolution. 68 (4): 996–1013. doi:10.1111/evo.12322. ISSN 0014-3820.
  30. ^ a b "Splendor in the Dark". Discover Magazine. Retrieved 2023-04-12.
  31. ^ Herring, Peter J.; Cope, Celia (2005-12-01). "Red bioluminescence in fishes: on the suborbital photophores of Malacosteus, Pachystomias and Aristostomias". Marine Biology. 148 (2): 383–394. doi:10.1007/s00227-005-0085-3. ISSN 1432-1793.

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Stomiidae: Brief Summary

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Stomiidae is a family of deep-sea ray-finned fish, including the barbeled dragonfishes. They are quite small, usually around 15 cm, up to 26 cm. These fish are apex predators and have enormous jaws filled with fang-like teeth. They are also able to hinge the neurocranium and upper-jaw system, which leads to the opening of the jaw to more than 100 degrees. This ability allows them to consume extremely large prey, often 50% greater than their standard length.

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Stomiidae ( Spanish; Castilian )

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 src=
Astronesthes similus.
 src=
Cabeza de Chauliodus.
 src=
Cabeza de Photostomias.
 src=
Stomias nebulosus.

Los peces-demonio o estómidos son la familia Stomiidae de peces marinos incluida en el orden stomiformes, distribuidos por las aguas abisales del océano Pacífico, Atlántico e Índico.[1]​ Su nombre procede del griego: stoma (boca), por su característica enorme boca bajo la cabeza.[2]

Anatomía

Los adultos no tienen verdadero rastrillo en las branquias, tienen un solo hueso infraorbital y uno o ninguno supramaxilares; el cuerpo tiene fotóforos productores de luz; la mayoría tienen una larga barbilla asociado al aparato hioideo; algunas especies no poseen aletas pectorales; el color del cuerpo en la mayoría es muy oscuro.[1]

Subfamilias y géneros

Existen unas 299 especies agrupadas en los 26 géneros siguientes:[3]

Referencias

  1. a b Nelson, J.S. (1994). Fishes of the world (en inglés) (3ª edición). Nueva York: John Wiley & Sons, Inc. pp. 600.
  2. Romero, P. (2002). An etymological dictionary of taxonomy. Madrid: unpublished.
  3. "Stomiidae". En FishBase (Rainer Froese y Daniel Pauly, eds.). Consultada en mayo de 2010. N.p.: FishBase, 2010.

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Stomiidae: Brief Summary ( Spanish; Castilian )

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 src= Astronesthes similus.  src= Cabeza de Chauliodus.  src= Cabeza de Photostomias.  src= Stomias nebulosus.

Los peces-demonio o estómidos son la familia Stomiidae de peces marinos incluida en el orden stomiformes, distribuidos por las aguas abisales del océano Pacífico, Atlántico e Índico.​ Su nombre procede del griego: stoma (boca), por su característica enorme boca bajo la cabeza.​

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Stomiidae ( Basque )

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Stomiidae arrain hezurdunen familia da, Stomiiformes ordenakoa.[1]. Nahiko txikiak dira eta zaila da hamabost zentimetro baino gehiagoko luzera izatea.

Generoak

Familiak 284 espezie ditu, 26 generotan banaturik:

Erreferentziak

  1. Joseph S. Nelson Fishes of the World John Wiley & Sons 207-208 or. ISBN 0-471-54713-1.
(RLQ=window.RLQ||[]).push(function(){mw.log.warn("Gadget "ErrefAurrebista" was not loaded. Please migrate it to use ResourceLoader. See u003Chttps://eu.wikipedia.org/wiki/Berezi:Gadgetaku003E.");});
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Stomiidae: Brief Summary ( Basque )

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Stomiidae arrain hezurdunen familia da, Stomiiformes ordenakoa.. Nahiko txikiak dira eta zaila da hamabost zentimetro baino gehiagoko luzera izatea.

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Kitakalat ( Finnish )

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Kitakalat (Stomiidae) on sukasuukaloihin kuuluva heimo. Heimon lajeja tavataan syvistä vesistä kaikista lämpimistä ja lauhkeista valtameristä.

Lajit ja anatomia

Kitakalojen heimo jaetaan 5:een tai 6:een alaheimoon. Heimoon kuuluu 28 sukua ja lähteestä riippuen 228–292 lajia. Tyypillisiä piirteitä kaikille alaheimojen lajeille ovat hyvin pitkänomainen ruumiinmuoto, suuri pää ja suu, suuret piikkimäiset hampaat, leuan nauhamainen, uloke, jonka päässä on valoelimiä, vatsan valoelimet ja suomuttomuus. Selkäevän pituus ja sijainti vaihtelevat alaheimoittain. Eräillä lajeilla on rasvaevä ja eräiltä puuttuvat rinta- ja vatsaevät. Suurimmillaan kitakalojen heimon lajit saavuttavat noin 50 cm:n pituuden. Väritykseltään heimon kalat ovat tyypillisesti mustia tai tummanruskeita. Eräät lajit ovat hopeanharmaita tai vihertäviä.[1][2][3][4]

Levinneisyys ja elintavat

Kitakalalajeja tavataan Atlantista, Intian valtamerestä ja Tyynestämerestä. Ne elävät syvissä vesissä aina 4 500 metrin syvyyteen asti. Öisin ne liikkuvat lähempänä pintaa kuin päivisin. Heimon kalat ovat petokaloja, jotka saalistavat planktonäyriäisiä ja suuremmat lajit myös kaloja. Ne houkuttelevat saaliseläimiä luokseen valoelimiensä avulla.[2][3][4]

Lähteet

  1. a b Nelson, Joseph S.: Fishes of the world, s. 210-212. Chichester: John Wiley and Sons, 2006. ISBN 978-0-471-25031-9. Kirja Googlen teoshaussa (viitattu 04.06.2013). (englanniksi)
  2. a b Family Stomiidae (peilipalvelin) FishBase. Froese, R. & Pauly, D. (toim.). Viitattu 4.6.2013. (englanniksi)
  3. a b Family Stomiidae (PDF) FAO. (englanniksi)
  4. a b Dianne J. Bray: Stomiidae Fishes of Australia. Viitattu 04.06.2013. (englanniksi)
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Kitakalat: Brief Summary ( Finnish )

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Kitakalat (Stomiidae) on sukasuukaloihin kuuluva heimo. Heimon lajeja tavataan syvistä vesistä kaikista lämpimistä ja lauhkeista valtameristä.

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Stomiidae ( French )

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Dragons à écailles

Les Stomiidae (Stomiidés en français), communément appelés Dragons à écailles, sont une famille de poissons téléostéens vivant dans les abysses.

Liste des sous-familles

 src=
Dessin de Malacosteus niger en chasse.
 src=
Un Chauliodus observé dans les abysses au large des îles Samoa (expédition Okeanos Explorer).

Selon ITIS (6 mars 2014)[2] & World Register of Marine Species (6 mars 2014)[3] :

Liste des genres

Selon ITIS (6 mars 2014)[2] :

Selon World Register of Marine Species (6 mars 2014)[3] :

Références taxinomiques

Notes et références

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Stomiidae: Brief Summary ( French )

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Dragons à écailles

Les Stomiidae (Stomiidés en français), communément appelés Dragons à écailles, sont une famille de poissons téléostéens vivant dans les abysses.

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Stomiidae ( Italian )

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Gli Stomidi (Stomiidae) sono una famiglia di pesci abissali, appartenenti all'ordine Stomiiformes.

Descrizione

Benché innocui all'uomo e lunghi in media 20 cm, questi pesci hanno sviluppato incredibili strategie di caccia, che comprendono dalla mascella estroflessibile del pesce mandibola (Malacosteus niger) al barbiglio penzolante dei draghi di mare (gen.Idiacanthus). Altra strana caratteristica di alcuni di questi pesci è la totale mancanza di scaglie e hanno nella maggior parte dei casi un colore nero o marrone, per mimetizzarsi nel buio totale degli abissi marini. Come tutti i predatori di profondità, dispongono ovviamente di fotofori allineati su tutto il corpo, che servono per attrarre la preda verso il pesce.

Alimentazione

Benché si nutrano di piccoli pesci e crostacei, non capita raramente che riescano a inghiottire prede grandi anche la metà di essi, dato il fatto che hanno stomaco e bocca espandibili. Possono però finire anch'essi prede di delfini, capodogli e altri grandi animali capaci di arrivare a notevoli profondità.

Generi

Attualmente si contano 277 specie di Stomidi suddivisi in 27 generi, ma visti gli ambienti difficilmente raggiungibili dall'uomo, si presuppone che alcune specie non siano ancora state scoperte.

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Stomiidae: Brief Summary ( Italian )

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Gli Stomidi (Stomiidae) sono una famiglia di pesci abissali, appartenenti all'ordine Stomiiformes.

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Didžiažiotės švietenės ( Lithuanian )

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Didžiažiotės švietenės (lot. Stomiidae, angl. Barbeled dragonfishes, vok. Barten-Drachenfische) – švietenžuvių (Stomiiformes) šeima. Paplitusios Atlanto, Indijos ir Ramiajame vandenynuose.

Sistematika

Šeima skirstoma į 6 pošeimius, kuriuose yra 27 gentys, 279 rūšys.

Galerija

Nuorodos


Vikiteka

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Didžiažiotės švietenės: Brief Summary ( Lithuanian )

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Didžiažiotės švietenės (lot. Stomiidae, angl. Barbeled dragonfishes, vok. Barten-Drachenfische) – švietenžuvių (Stomiiformes) šeima. Paplitusios Atlanto, Indijos ir Ramiajame vandenynuose.

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Stomiidae ( Dutch; Flemish )

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Vissen

Stomiidae zijn een familie van straalvinnige vissen uit de orde van Draakvisachtigen (Stomiiformes).[1]

Onderfamilies

Bronnen, noten en/of referenties
  1. (en) Stomiidae. FishBase. Ed. Ranier Froese and Daniel Pauly. 10 2011 version. N.p.: FishBase, 2011.
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Stomiidae: Brief Summary ( Dutch; Flemish )

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Stomiidae zijn een familie van straalvinnige vissen uit de orde van Draakvisachtigen (Stomiiformes).

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Wężorowate ( Polish )

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Commons Multimedia w Wikimedia Commons
 src=
Kladogram rodziny Stomiidae[2]
 src=
Przedstawiciele wężorowatych na rycinie z 1912 roku

Wężorowate[3] (Stomiidae) – rodzina drapieżnych, morskich ryb promieniopłetwych z rzędu wężorokształtnych (Stomiiformes).

Zasięg występowania

Występują w wodach mezopelagialnych i batypelagialnych wszystkich oceanów świata z wyjątkiem mórz polarnych[4].

Charakterystyka

Ciało silnie wydłużone, bocznie spłaszczone. Głowa zaokrąglona, z dużym i głębokim otworem gębowym wyposażonym w długie i ostre zęby, przystosowanym do chwytania i połykania dużych ofiar. Narządy świetlne ułożone wzdłuż boków. Niektóre gatunki mają pierwszy promień płetwy grzbietowej przekształcony w długą wić pełniącą funkcję wabika. U innych taka wić wyrasta na podbródku. Ciało nagie lub pokryte cienkimi łuskami[3].

Do najbardziej znanych przedstawicieli wężorowatych należą idiakanty (Idiacanthus).

Klasyfikacja

Rodzaje zaliczane do tej rodziny są zgrupowane w podrodzinach[5]:

Chauliodontinae: Chauliodus

Stomiinae: Stomias

Astronesthinae: AstronesthesBorostomiasEupogonesthesHeterophotusNeonesthesRhadinesthes

Melanostomiinae: BathophilusChirostomiasDiplostomiasEchiostomaEustomiasFlagellostomiasGrammatostomiasLeptostomiasMelanostomiasOdontostomiasOpostomiasPachystomiasPhotonectesTactostomaThysanactisTrigonolampa

Malacosteinae: AristostomiasMalacosteusPhotostomias

Idiacanthinae: Idiacanthus

Rodzajem typowym rodziny jest Stomias.

Zobacz też

Przypisy

  1. Stomiidae, w: Integrated Taxonomic Information System (ang.).
  2. WL Fink. Phylogenetic interrelationships of the stomiid fishes (Teleostei: Stomiiformes). „Miscellaneous Publications”. 171, s. 1–127, 1985. Museum of Zoology, University of Michigan (ang.).
  3. a b Stanisław Rutkowicz: Encyklopedia ryb morskich. Gdańsk: Wydawnictwo Morskie, 1982, s. 54. ISBN 83-215-2103-7.
  4. Nelson 2006 ↓, s. 210.
  5. Eschmeyer, W. N. & Fricke, R.: Catalog of Fishes electronic version (7 August 2012) (ang.). California Academy of Sciences. [dostęp 17 sierpnia 2012].

Bibliografia

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Wężorowate: Brief Summary ( Polish )

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 src= Kladogram rodziny Stomiidae  src= Przedstawiciele wężorowatych na rycinie z 1912 roku

Wężorowate (Stomiidae) – rodzina drapieżnych, morskich ryb promieniopłetwych z rzędu wężorokształtnych (Stomiiformes).

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Stomiidae ( Portuguese )

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Stomiidae é uma família de peixes da ordem Stomiiformes. A família Stomiidae representa uma das famílias de peixes dominantes nos ecossistemas mesopelágicos, exibindo uma matriz de especializações para uma existência predatória neste ambiente; Boca enorme gapes com dentes proeminentes, estômagos distensíveis, alongados corpos escuros com fotóforos, barbatanas e queixo com tecido bioluminescente.

Anatomia

Em contraste com a sua pele pigmentada escura, o peixe-dragão está equipado com dentes transparentes.[1]

Classificação

Família Stomiidae

Referências

  • Nelson, Joseph, S. (2006). Fishes of the World. John Wiley & Sons, Inc.. ISBN 0-471-25031-7.
  1. Malewar, Amit (7 de junho de 2019). «What makes the teeth of deep-sea dragonfish transparent?». Tech Explorist (em inglês). Consultado em 12 de julho de 2019
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Stomiidae: Brief Summary ( Portuguese )

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Stomiidae é uma família de peixes da ordem Stomiiformes. A família Stomiidae representa uma das famílias de peixes dominantes nos ecossistemas mesopelágicos, exibindo uma matriz de especializações para uma existência predatória neste ambiente; Boca enorme gapes com dentes proeminentes, estômagos distensíveis, alongados corpos escuros com fotóforos, barbatanas e queixo com tecido bioluminescente.

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Drakfiskar ( Swedish )

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Drakfiskar[1] (Stomiidae[2]) är en familj av fiskar som ingår i ordningen drakfiskartade fiskar (Stomiiformes).[2] Enligt Catalogue of Life omfattar familjen Stomiidae 280 arter[2]. Fishbase listar 287 arter.[3]

Familjens medlemmar förekommer i Atlanten, Indiska oceanen och Stilla havet. De har oftast en mörk kroppsfärg. Det vetenskapliga namnet är bildat av det grekiska ordet stoma (mun).[3]

Taxonomi

Släkten enligt Catalogue of Life[2]:

Fishbase listar ytterligare ett släkte.[3]

Källor

  1. ^ [a b] Drakfiskar, Nationalencyklopedin, läst 2017-04-03.
  2. ^ [a b c d] Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (red.) (28 april 2011). ”Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist.”. Species 2000: Reading, UK. http://www.catalogueoflife.org/annual-checklist/2011/search/all/key/stomiidae/match/1. Läst 24 september 2012.
  3. ^ [a b c] Stomiidae, Fishbase, läst 2017-04-03.

Externa länkar

Mört, Nordisk familjebok.png Denna fiskrelaterade artikel saknar väsentlig information. Du kan hjälpa till genom att tillföra sådan.
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Drakfiskar: Brief Summary ( Swedish )

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Drakfiskar (Stomiidae) är en familj av fiskar som ingår i ordningen drakfiskartade fiskar (Stomiiformes). Enligt Catalogue of Life omfattar familjen Stomiidae 280 arter. Fishbase listar 287 arter.

Familjens medlemmar förekommer i Atlanten, Indiska oceanen och Stilla havet. De har oftast en mörk kroppsfärg. Det vetenskapliga namnet är bildat av det grekiska ordet stoma (mun).

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Стомієві ( Ukrainian )

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Поширення

Родина розповсюджена у Атлантичному, Індійському, Тихому океанах.

Опис

Більшість видів темнозабарвлені. Орган світіння у фотофорах відсутні. На підборідді є вусик, пов'язаний з гіпоїдним апаратом. Справжні зяброві тичинки відсутні. Це риби середнього розміру, що сягають 15-40 см завдовжки.

Спосіб життя

Всі стомієві є глибоководними істотами (мешкають на глибині 300–500 м, інколи до 2000 м). Живляться дрібною рибою[1].

Класифікація

Родина містить 28 родів і 285 видів:

Примітки

Джерела


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Stomiidae ( Vietnamese )

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Bài viết này cần thêm chú thích nguồn gốc để kiểm chứng thông tin. Mời bạn giúp hoàn thiện bài viết này bằng cách bổ sung chú thích tới các nguồn đáng tin cậy. Các nội dung không có nguồn có thể bị nghi ngờ và xóa bỏ.

Stomiidae là một họ cá vây tia biển sâu. Chúng khá nhỏ, thường khoảng 15 cm, tới 26 cm.

Tham khảo


Hình tượng sơ khai Bài viết Lớp Cá vây tia này vẫn còn sơ khai. Bạn có thể giúp Wikipedia bằng cách mở rộng nội dung để bài được hoàn chỉnh hơn.
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Stomiidae: Brief Summary ( Vietnamese )

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Stomiidae là một họ cá vây tia biển sâu. Chúng khá nhỏ, thường khoảng 15 cm, tới 26 cm.

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Стомиевые ( Russian )

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Царство: Животные
Подцарство: Эуметазои
Без ранга: Вторичноротые
Подтип: Позвоночные
Инфратип: Челюстноротые
Группа: Рыбы
Группа: Костные рыбы
Подкласс: Новопёрые рыбы
Инфракласс: Костистые рыбы
Надотряд: Osmeromorpha
Семейство: Стомиевые
Международное научное название

Stomiidae

Синонимы
  • Stomiatidae
Подсемейства Wikispecies-logo.svg
Систематика
на Викивидах
Commons-logo.svg
Изображения
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ITIS 162282NCBI 123351EOL 5074FW 265829

Сто́миевые (лат. Stomiidae) — семейство глубоководных рыб из отряда стомиеобразных, или иглоротов (Stomiiformes). Атлантический, Индийский и Тихий океаны. Содержит 28 родов и около 285 видов. Большинство видов тёмноокрашенные. Орган свечения в фотофорах отсутствует. На подбородке имеется усик, связанный с гипоидным аппаратом. Истинные жаберные тычинки отсутствуют[1]. Хищники, поедающие мелких рыб и планктонных рачков. Обитают, как правило, на глубинах от 300 до 500 м (но могут быть встречены на глубине до 2000 м)[2].

Систематика

Включаемым в Stomiidae подсемействам иногда придают статус самостоятельных семейств[1].

Примечания

  1. 1 2 Нельсон Дж. С. Рыбы мировой фауны / Пер. 4-го перераб. англ. изд. Богуцкой Н. Г., науч. ред-ры Насека А. М., Герд А. С. — М.: Книжный дом «ЛИБРОКОМ», 2009. — С. 393—395. — 880 с. — ISBN 978-5-397-00675-0
  2. Жизнь животных: в 6-ти томах. — М.: Просвещение. Под редакцией профессоров Н. А. Гладкова, А. В. Михеева. 1970.
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Авторы и редакторы Википедии

Стомиевые: Brief Summary ( Russian )

provided by wikipedia русскую Википедию

Сто́миевые (лат. Stomiidae) — семейство глубоководных рыб из отряда стомиеобразных, или иглоротов (Stomiiformes). Атлантический, Индийский и Тихий океаны. Содержит 28 родов и около 285 видов. Большинство видов тёмноокрашенные. Орган свечения в фотофорах отсутствует. На подбородке имеется усик, связанный с гипоидным аппаратом. Истинные жаберные тычинки отсутствуют. Хищники, поедающие мелких рыб и планктонных рачков. Обитают, как правило, на глубинах от 300 до 500 м (но могут быть встречены на глубине до 2000 м).

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巨口魚科 ( Chinese )

provided by wikipedia 中文维基百科
  • 見內文

巨口魚科学名Stomiidae),是輻鰭魚綱巨口魚目的其中一科。

目录

分類

巨口魚科下分27個屬,如下:

星衫魚亞科(Astronesthinae)

星衫魚屬(Astronesthes

主条目:星衫魚屬

掠食巨口魚屬(Borostomias

真芒巨口魚屬(Eupogonesthes

異星杉魚屬(Heterophotus

新衫魚屬(Neonesthes

細杉魚屬(Rhadinesthes

蝰魚亞科(Chauliodontinae)

蝰魚屬(Chauliodus

奇棘魚亞科(Idiacanthinae)

奇棘魚屬(Idiacanthus

柔骨魚亞科(Malacosteinae)

奇巨口魚屬(Aristostomias

柔骨魚屬(Malacosteus

  • Malacosteus australis
  • 黑柔骨魚Malacosteus niger

光巨口魚屬(Photostomias

  • Photostomias atrox
  • Photostomias goodyeari
  • 格氏光巨口鱼Photostomias guernei
  • Photostomias liemi
  • Photostomias lucingens
  • Photostomias tantillux

黑巨口魚亞科(Melanostomiinae)

深巨口魚屬(Bathophilus

肢巨口魚屬(Chirostomias

刺巨口魚屬(Echiostoma

真巨口魚屬(Eustomias

鞭鬚巨口魚屬(Flagellostomias

裸巨口魚屬(Grammatostomias

纖巨口魚屬(Leptostomias

黑巨口魚屬(Melanostomias

強牙巨口魚屬(Odontostomias

脂巨口魚屬(Opostomias

厚巨口魚屬(Pachystomias

袋巨口魚屬(Photonectes

箭口魚屬(Tactostoma

纓光魚屬(Thysanactis

三線巨口魚屬(Trigonolampa

巨口魚亞科(Stomiinae)

巨口魚屬(Stomias

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巨口魚科: Brief Summary ( Chinese )

provided by wikipedia 中文维基百科

巨口魚科(学名:Stomiidae),是輻鰭魚綱巨口魚目的其中一科。

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维基百科作者和编辑

스토미아스과 ( Korean )

provided by wikipedia 한국어 위키백과

스토미아스과(Stomiidae)는 앨퉁이목에 속하는 조기어류 과의 하나이다.[1] 아주 작은 물고기로 보통 15cm정도이고, 최대 26cm에 불과하다. 바이퍼피시 등을 포함하고 있다.

하위 분류

스토미아스과는 다음과 같이 분류한다.[1]

각주

  1. (영어) "Stomiidae". FishBase. Ed. Rainer Froese and Daniel Pauly. 2015년 1월 version. N.p.: FishBase, 2015년.
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Distribution

provided by World Register of Marine Species
Distribution: Atlantic, Indian, and Pacific. Elongate body. Dorsal fin origin far behind pelvics, above anal fin. Chin barbel long. Adipose fin absent.

Reference

MASDEA (1997).

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contributor
Edward Vanden Berghe [email]