Fragrant White Sand Verbena

Plants perennial, sometimes nearly acaulescent. Stems decumbent to erect, elongate, glandular-pubescent, infrequently glabrous. Leaves: petiole 1-4 cm; blade ovate elliptic-oblong to ovate, 1.5-6 × 0.5-3.5 cm, margins entire to sinuate, often undulate, adaxial surface glabrous or puberulent, abaxial surface thinly puberulent to pubescent. Inflorescences: peduncle longer than subtending petiole; bracts ovate to obovate, 5-20 × 3-10 mm, scarious, apex obtuse to acute, glandular-pubescent to villous; flowers 25-75. Perianth: tube rose to greenish, 10-20 mm, limb white, 5-8 mm diam. Fruits broadly turbinate, apex truncate or rounded and slightly beaked, or fruit ± rhombic in profile, 5-12 × 4-8.5 mm, scarious, tapered at both ends; wings (2-)5 (often 2 on periphery of inflorescence and folded together), dilated distally and flattened perpendicular to plane of lamina, dilations longer than wide, thin walled, cavities extending throughout.

Ariz., Colo., Nev., N.Mex., Utah, Wyo.

Flowering spring-fall.

Sandy or gravelly soils, desert grasslands, scrub; 700-2500m.

Abronia fragrans Nuttall ex Hooker var. elliptica (A. Nelson) M. E. Jones; A. nana S. Watson var. harrisii S. L. Welsh

Abronia oumila Rvdb. Bull. Torrev C
1902.
Abronia sparsifolia Standley, Contr. U. S. Nat. Herb. 12: 322. 1909.
Perennial, from a slender woody root; stems few or numerous, ascending or decumbent, 1-4 dm. long, stout, whitish, minutely viscid-puberulent or glabrate, sparsely branched, the ititernodes short or elongate; petioles slender, 1-3 cm. long, puberulent or glabrate; leaf -blades oyate-oblong, ovate-orbicular, deltoid-ovate, or elliptic-oblong, 1.5-3.5 cm. long, 0.7-2.5 cm. wide, subcordate to obtuse at the base, rounded or obtuse at the apex, entire, thick and succulent, often glaucescent beneath, minutely puberulent or glabrate; peduncles slender, 2-5 cm. long, viscid-puberulent; bracts broadly ovate or oval, 5-10 mm. long, 4-5 mm. wide, acute or attenuate, scarious, whitish, viscid-puberulent; flowers numerous, the perianth 13-18 mm. long, viscid-puberulent, the tube slender, the limb about 5 mm. broad; fruit turbinate, about 5 mm. long, subcoriaceous stramineous or olivaceous, finely puberulent, usually deeply 5-lobed, the lobes compressed, truncate and flat above; seed elliptic-oblong, 2.5 mm. long, brown.
Type locality: Emery, Utah, at an altitude of 2100 meters. Distribution: Southern Utah and adjacent Nevada and Arizona.

Abronia elliptica A. Nelson, Bull. Torrey Club 26: 7. 1899
Abronia Bakeri Greene, PL Baker. 3: 32. 1901.
Abronia fragrans elliptica Heimerl; Rydb. Bull. Torrey Club 29: 684. 1902.
Abronia glabra Rydb. Bull. Torrey Club 29: 685. 1902.
Abronia fragrans elliptica M. B. Jones, Contr. West. Bot. 11: 3. 1903.
Abronia ramosa Standley, Contr. U. S. Nat. Herb. 12: 321. 1909.
Abronia glabrifolia Standley, Contr. U. S. Nat. Herb. 12: 321. 1909.
Perennial, often from a woody root; stems usually numerous, erect or decumbent, 1-5 dm. long, stout, usually copiously branched, whitish or tinged with red, glabrous or finely viscid-puberulent, especially above; petioles slender, 1-4 cm. long, puberulent or glabrate; leaf-blades usually oval, oval-oblong, or ovate-oblong, sometimes deltoid or elliptic-ovate, or those of the lowest leaves suborbicular, 1-5.5 cm. long, 0.8-3.5 cm. wide, rounded, truncate, or obtuse at the base, rounded or obtuse at the apex, succulent, entire or subsinuate, usually pale or glaucescent beneath, minutely puberulent or glabrous; peduncles slender, 2-12 cm. long, glabrous or puberulent; bracts broadly oval or obovate-oval, 10-12 mm. long, 5-7 mm. wide, rounded and often apiculate at the apex, scarious, white or greenish-white, glabrous or minutely puberulent; flowers very numerous, the perianth 18-22 mm. long, viscid-puberulent, the tube very slender, greenish-white or pinkish, the limb 5-8 mm. broad, white; fruit turbinate, or the outer ones biturbinate, 5-8 mm. long, subcoriaceous, puberulent or short-villous above, stramineous or olivaceous, usually deeply 5-lobed, the lobes compressed, finely reticulateveined, rounded or truncate above and flattened at the apex; seed elliptic-oblong, 2.5-3 mm. long,
brown, lustrous.
Typb locality: Green River, Wyoming.
Distribution: In dry, clay or sandy soil, Wyoming to northeastern Arizona and northwestern

Abronia salsa Rydb. Bull. Torrey Club 29: 684. 1902
Abronia fallax Heimerl; Rydb. Bull. Torrey Club 29: 684. 1902. Abronia fragrans pterocarpa M. E. Jones, Contr. West. Bot. 11: 3. 1903„
Perennial, from a somewhat woody elongate root; stems few or numerous, 2-5 dm. long, erect or decumbent, stout, whitish or stramineous, short-villous or puberulent below or sometimes glabrate, viscidvillous above with short hairs, the internodes elongate; petioles slender, 1-4 cm. long, viscid-puberulent or short-villous; leaf-blades oblong, oval, elliptic-oblong, or oblong-ovate, rarely deltoid-ovate or suborbicular, sometimes lanceolate, 2-5.5 cm. long, 0.7-3 cm. wide, rounded to acute at the base, broadly rounded to acute at the apex, entire or subsinuate, concolorous or paler beneath, densely viscid-puberulent; peduncles slender or stout, 1.5-8 cm. long, viscidvillous; bracts broadly oval or rounded-ovate, 10-16 mm. long, 5-12 mm. wide, acute, white, scarious, viscidvillous or puberulent; flowers numerous, the perianth 18-25 mm. long, the tube very slender, villous, greenish, the limb 5-8 mm. broad, white; fruit turbinate, 5-7 mm. long, coriaceous, whitish-stramineous, short-villous or puberulent, usually deeply 5-lobed, the lobes compressed and winglike, finely reticulate-veined, truncate at the summit and flattened; seed narrowly obovoid, 2.5-3 mm. long, dark-brown, lustrous.
Type locality: Salt Lake pity, Utah.
Distribution: In sandy soil, Utah.