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Lifespan, longevity, and ageing

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Maximum longevity: 5 years (wild) Observations: The salmon ages and dies shortly after spawning. It is not considered ageing but rather sudden death.
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The scientific name Oncorhynchus kisutch was derived from the Greek roots onkos, (hook), rynchos (nose), and kisutch, the ordinary name in Siberia and Alaska. Some other common names for Oncorhynchus kisutch are coho, silver, blueback, and hook nose salmon. (StreamNet, 1996)

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Kyle Smith, University of Michigan-Ann Arbor
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Behavior

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Perception Channels: tactile ; chemical

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Kyle Smith, University of Michigan-Ann Arbor
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Conservation Status

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There has been a serious decline of wild coho stocks in the Strait of Georgia, due to loss of habitat (from dams) and overfishing. A plan of harvest management has been developed and implemented by the Dept. of Fisheries and Oceans. (Pacific Salmon Foundation, 1995)

US Federal List: no special status

CITES: no special status

State of Michigan List: no special status

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Kyle Smith, University of Michigan-Ann Arbor
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Benefits

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Oncorhynchus kisutch make a good meal. Often, coho are either sold frozen or canned by commercial fisherman. Twice as many coho salmon are harvested commercially in North America than in Asia. During the late 1950's about 10 million coho salmon per year were caught. These numbers declined to 4.5 million coho per year in the 60's and have now risen to about 10.5 million coho per year. Fishing for coho is also an important sport for trollers and flyfisherman. (McNeil et. al, 1980)

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Kyle Smith, University of Michigan-Ann Arbor
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Trophic Strategy

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In fresh water streams and rivers, juvenile coho salmon defend territories and compete for limited feeding sites with other fish. In order to attain these feeding sites, they may charge or chase other fish away, as growth and fitness depend on it. Coho salmon attack prey by sprinting and striking. They are able to maneuver well in order to capture prey. In fresh water, coho eat insects and smaller fish. When they move to the ocean they begin to grow rapidly due to the abundance of food. Once these fish reach the ocean their diet changes to other fish, such as herring and squid (Groot et. al., 1995).

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Kyle Smith, University of Michigan-Ann Arbor
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Distribution

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Oncorhynchus kisutch, commonly known as coho salmon, are primarily found in coastal waters. Coho salmon do not range widely in the open ocean like that of many other salmon species. Oncorhynchus kisutch are also found in fresh water, during their first year and later while spawning. They are found in fresh streams of the North Pacific, ranging from Baja, California to Alaska (Wheeley, 1985).

Biogeographic Regions: nearctic (Native ); arctic ocean (Native ); atlantic ocean (Native ); pacific ocean (Native )

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Kyle Smith, University of Michigan-Ann Arbor
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Tanya Dewey, Animal Diversity Web
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Habitat

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Coho Salmon live in fresh and ocean water. In fresh water, coho like relatively slow moving water with fine gravel. In the ocean, coho tend to live closer to shores than in the open ocean. (StreamNet, 1996)

Aquatic Biomes: rivers and streams; coastal

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Life Expectancy

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Average lifespan
Status: wild:
4.1 years.

Average lifespan
Status: wild:
5.0 years.

Average lifespan
Status: wild:
4.0 years.

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Kyle Smith, University of Michigan-Ann Arbor
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Morphology

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Oncorhynchus kisutch typically spend the first year of their life in fresh water. Upon moving to the ocean, growth increases dramatically. At about the third year, coho salmon reach maturity. They weigh from 6 to 12 pounds and can be up to 38 inches in length.

Oncorhynchus kisutch are deep-bodied salmon with unique color characteristics. The dorsal surface is a metallic blue while the sides are a silver color. Black spots appear on the back and upper lobe of the caudal fin. A lateral line is also present. The line is curved toward the front of the fish and straightens out as it approaches the back of the fish. While spawning, the fish's back and belly turn dark. Spawning males' sides develop a bright red line and their jaws become hooked (Clemens and Wilby, 1961).

Range mass: 3 to 6 kg.

Other Physical Features: ectothermic ; heterothermic ; bilateral symmetry

Sexual Dimorphism: sexes colored or patterned differently

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Kyle Smith, University of Michigan-Ann Arbor
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Reproduction

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At three to four years of age, coho salmon reach sexual maturity. From September to October, coho swim to the fresh water streams where they were born (up to 400 miles). Once the fish reach their natal site females dig a nest in a gravel-type area. After the nest is made a female and one male (occasionally 2) breed. In this breeding, the female lays her eggs and the male's sperm is spread over them, thus fertilizing them. After fertilization has occurred, the eggs are buried by other female coho that are digging their nests. Following reproduction, males and females die, giving the waters more nutrients. Larvae hatch 6-8 weeks after fertilization. The larvae remain in the gravel for 2-3 weeks. Coho live in streams or rivers for about a year until they move to the ocean. (Wheeler, 1985)

Average age at sexual or reproductive maturity (male)
Sex: male:
1095 days.

Average age at sexual or reproductive maturity (female)
Sex: female:
1095 days.

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Smith, K. 2000. "Oncorhynchus kisutch" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Oncorhynchus_kisutch.html
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Comprehensive Description

provided by EOL staff

The Coho or Silver Salmon (Oncorhynchus kisutch) has black spots on the back and upper lobe of the caudal fin. The gums are light at the base of the teeth. At sea, these salmon are metallic blue above and silver below. Breeding males have dusky green on the back and head, with bright red sides and a black belly; females have bonze to pink-red sides. Spawning males develop a strongly hooked snout and large teeth. Maximum length is around 100 cm. Chinook Salmon (O. tshawytscha) are very similar in appearance to Coho Salmon while at sea, but Chinook Salmon are larger, have black spots on both the upper and lower caudal fin lobes, and have gums that are dark at the base of the teeth. (Page and Burr 1991)

Coho Salmon occur in northeastern Asia and, in North America, in Arctic and Pacific drainages from Point Hope, Alaska, to Monterey Bay, California (occasionally as far south as Baja California). These fish are anadromous, returning from the ocean to the coastal streams or rivers where they were born to spawn. They reproduce just once, then die. (Page and Burr 1991)

For detailed information on the biology and status of this species, including conservation issues, see this resource from the NOAA Fisheries Office of Protected Resources.

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Benefits

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Caught by gillnets and trawl nets. The total catch reported for this species to FAO for 1999 was 15 466 t. The countries with the largest catches were USA (13 259 t) and Russian Federation (1 668 t). Marketed fresh, dried/salted, smoked, canned, cured, and frozen. Eaten steamed, fried, broiled, boiled, microwaved and baked.

Brief Summary

provided by FAO species catalogs
Anadromous, adults enter rivers from autumn to winter and spawn about 2,500 to 7,000 eggs in headwaters. Fry emerge from bottom gravel in spring, spending about one year in the river. Instead of territorial behavior in the parr, the smolt make schools and are carried out to sea. Young fish remain close to shore. Immature migrate more than 1600 km offshore.Main food items are insects in the river, small fishes such as sandlance and herring and many crustaceans near shore, and squids and krill in the open ocean.After 1 or 2 years of at sea, adults return to the river.

Size

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To about 88 cm and 14 kg; usually to around 3.6 kg at which weight it measures around 50-76 cm.

Distribution

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A Pacific fish (in rivers from Kamchatka, through Alaska to California. Incidental migrations to rivers of Shiribetsu and Yurapu of Hokkaido have been reported), introduced into northern rivers of France; in 1973 and 1974, 50,000 yearlings escaped into the Varenne (Normandy) from a fish farm, and 10,000 in 1975, with lesser numbers in southern France and southern Brittany in 1976-79; in 1975-77 up to 25 adults were caught in the Varenne, in rivers to the north, and around the Channel Is. Also introduced to the Great Lakes of America, Argentina, and Chile.

Diagnostic Description

provided by FAO species catalogs
Body elongate, becoming deeper with age, spawning males a little hump-backed. Tip of upper jaw reaching well behind eye; snout and lower jaw becoming hooked and teeth enlarged in spawning males. Gillrakers 18-25. Piloric coeca 45-81. Dorsal finrays iii-iv + 8-10, and an adipose fin behind it; base below about middle of dorsal fin. Anal finrays iv-v + 12-14. Caudal fin emarginate. Scales moderate, 120-140 in lateral line. Vertebrae 61-69. Colour silvery, with black spots on upper flanks, base of dorsal fin and upper lobe of caudal fin in adults.

Life Cycle

provided by Fishbase
Adults migrate from the sea or lake, school at mouths of rivers, and move upstream when rains increase river flow (Ref. 1998). As a rule, the winter entries (December and January) occur in the southern part of the range, with appearance in fresh water becoming progressively earlier to the north (Ref. 30381, 30382, 30383). At the spawning area, the female finds a spot and digs a pitt. At this point she is aggressive toward other females. While digging, an attendant male courts her or is busy driving away other males. As soon as the pitt is completed, the female drops into it and is immediately followed by the male. The pair are side by side, they open their mouth, quiver and release egg and sperm (Ref. 27547). At this point, other males move in and release sperm into the nest (Ref. 1998). The female quickly moves to the upstream edge of the nest and starts digging a new pitt, covering the eggs. The whole process is repeated for several days until the female deposits all her eggs. The male then leaves and may seek another female. The spent female usually continues to dig, until she dies (Ref. 27547).Reproductive strategy: synchronous ovarian organization, determinate fecundity (Ref. 51846).
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Trophic Strategy

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The fish occur in the ocean or in lakes; adults return to the rivers where they were born (Ref. 27547). The young fish emerge in springtime and they usually live in fresh water for 1-2 years (sometimes up to 4 years, Ref. 27547); later they migrate at night to freshwater lakes or to the sea (Ref. 1998). Epipelagic (Ref. 58426). The fish that stay more than two years in fresh water and become sexually ripe without ever going to sea, are called residuals; they never spawn (Ref. 27547). Young fish in lakes and rivers eat mainly insects; they stay almost entirely in deep parts of the river and soon become strongly territorial (Ref. 27547). Upon reaching the sea, the smolts remain close to the coast for a certain time, eating planktonic crustaceans (Ref. 27547). As they grow, they migrate farther out into the sea and hunt larger organisms (Ref. 27547) such as jellyfish, squids and fishes (Ref. 58426). They are hunted by various fishes, birds (mergansers, loons and kingfishers), mammals and lampreys (Ref. 1998).
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Morphology

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Dorsal spines (total): 0; Dorsal soft rays (total): 9 - 13; Analspines: 0; Analsoft rays: 12 - 17; Vertebrae: 61 - 69
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Migration

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Anadromous. Fish that ascend rivers to spawn, as salmon and hilsa do. Sub-division of diadromous. Migrations should be cyclical and predictable and cover more than 100 km.
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Susan M. Luna
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Diseases and Parasites

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Epitheliocystis. Bacterial diseases
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Diseases and Parasites

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Enteric Redmouth Disease. Bacterial diseases
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Diagnostic Description

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Characterized by the presence of small black spots on the back and on the upper lobe of the caudal fin, and by the lack of dark pigment along the gum line of the lower jaw (Ref. 27547). The gill rakers are rough and widely spaced; the lateral line is nearly straight (Ref. 27547). The adipose fin is slender; the pelvic fins have an axillary extension (Ref. 27547). Fish in the sea are dark metallic blue or greenish on the back and upper sides, a brilliant silver color on middle and lower sides, and white below; small black spots are present on the back and upper sides and on the upper lobe of the caudal fin (Ref. 27547). During the spawning season fish turn dark to bright green on head and back, bright red on the sides, and often dark on the belly (Ref. 27547). Females are less brightly colored than males (Ref. 27547).
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Biology

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The fish occur in the ocean or in lakes; adults return to the rivers where they were born (Ref. 27547). The young fish emerge in springtime and they usually live in fresh water for 1-2 years (sometimes up to 4 years, Ref. 27547); later they migrate at night to freshwater lakes or to the sea (Ref. 1998). Epipelagic (Ref. 58426). The fish that stay more than two years in fresh water and become sexually ripe without ever going to sea, are called residuals; they never spawn (Ref. 27547). Young fish in lakes and rivers eat mainly insects; they stay almost entirely in deep parts of the river and soon become strongly territorial (Ref. 27547). Upon reaching the sea, the smolts remain close to the coast for a certain time, eating planktonic crustaceans (Ref. 27547). As they grow, they migrate farther out into the sea and hunt larger organisms (Ref. 27547) such as jellyfish, squids and fishes (Ref. 58426). They are hunted by various fishes, birds (mergansers, loons and kingfishers), mammals and lampreys (Ref. 1998). This kind is traded as fresh fish, dried or salted, smoked, canned, preserved and frozen (Ref. 9988). They are steamed, grilled, broiled, cooked in the microwave and baked (Ref. 9988).
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Importance

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fisheries: highly commercial; aquaculture: commercial; gamefish: yes; price category: medium; price reliability: reliable: based on ex-vessel price for this species
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Coho salmon

provided by wikipedia EN

The coho salmon (Oncorhynchus kisutch; Karuk: achvuun[1]) is a species of anadromous fish in the salmon family and one of the five Pacific salmon species. Coho salmon are also known as silver salmon or "silvers". The scientific species name is based on the Russian common name kizhuch (кижуч).

Description

Coho salmon from Oregon

During their ocean phase, coho salmon have silver sides and dark-blue backs. During their spawning phase, their jaws and teeth become hooked. After entering fresh water, they develop bright-red sides, bluish-green heads and backs, dark bellies and dark spots on their backs. Sexually maturing fish develop a light-pink or rose shading along the belly, and the males may show a slight arching of the back. Mature adults have a pronounced red skin color with darker backs and average 28 inches (71 cm) and 7 to 11 pounds (3.2 to 5.0 kg), occasionally reaching up to 36 pounds (16 kg). They also develop a large kype (hooked beak) during spawning.[2] Mature females may be darker than males, with both showing a pronounced hook on the nose.[3]

Reproduction

The eggs hatch in the late winter or early spring after six to seven weeks in the redd. Once hatched, they remain mostly immobile in the redd during the alevin life stage, which lasts for 6–7 weeks. Alevin no longer have the protective egg shell, or chorion, and rely on their yolk sacs for nourishment during growth. The alevin life stage is very sensitive to aquatic and sedimental contaminants. When the yolk sac is completely resorbed, the alevin leaves the redd. Young coho spend one to two years in their freshwater natal streams, often spending the first winter in off-channel sloughs, before transforming to the smolt stage. Smolts are generally 100–150 mm (3.9–5.9 in) and as their parr marks fade and the adult's characteristic silver scales start to dominate. Smolts migrate to the ocean from late March through July. Some fish leave fresh water in the spring, spend summer in brackish estuarine ponds, and then return to fresh water in the fall. Coho salmon live in salt water for one to three years before returning to spawn. Some precocious males, known as "jacks", return as two-year-old spawners. Spawning males develop kypes, which are strongly hooked snouts and large teeth.

Range

The traditional range of the coho salmon runs along both sides of the North Pacific Ocean, from Hokkaidō, Japan and eastern Russia, around the Bering Sea to mainland Alaska, and south to Monterey Bay, California.[4] Coho salmon have also been introduced in all the Great Lakes, as well as many landlocked reservoirs throughout the United States.[5] A number of specimens, (more than 20), were caught in waters surrounding Denmark and Norway in 2017. Their source is currently unknown, but the salmon species is farmed at several locations in Europe, making it probable that the animal has slipped the net at such a farm.[6]

Human uses

Freshly caught coho

Fisheries

The total North Pacific harvest of coho salmon in 2010 exceeded 6.3 million fish, of which 4.5 million were taken in the United States and 1.7 million in Russia. This corresponds to some 21,000 tonnes in all.[7] Coho salmon are the backbone of the Alaskan troll fishery; however, the majority are caught by the net fishery (gillnet and seine). Coho salmon average 3.5% by fish and 5.9% by weight of the annual Alaska salmon harvest.[8] The total North Pacific yields of the pink salmon, chum salmon and sockeye salmon are some 10–20 fold larger by weight.[7]

Game fish

In North America, coho salmon is a game fish in fresh and salt water from July to December, especially with light fishing tackle. It is one of the most popular sport fish in the Pacific Northwest of the United States and Canada. Its popularity is due in part to the reckless abandon which it frequently displays chasing bait and lure while in salt water, and the large number of coastal streams it ascends during its spawning runs. Its habit of schooling in relatively shallow water, and often near beaches, makes it accessible to anglers on the banks, as well as in boats.[9]

It is also pursued by fly fisherman in salt water.[10]

Nutritional value

Ocean-caught coho is regarded as excellent table fare. It has a moderate to high amount of fat, which is considered to be essential when judging taste. Only spring chinook and sockeye salmon have higher levels of fat in their meat. When smoking coho it is best to use a cold-smoking rather than hot-smoking process, due to their lower fat content compared to sockeye and chinook.

Cultural tradition

Historically coho, along with other species, has been a staple in the diet of several indigenous peoples, who would also use it to trade with other tribes farther inland. The coho salmon is also a symbol of several tribes, representing life and sustenance.

Ecology

In their freshwater stages, coho feed on plankton and aquatic invertebrates in the benthos and water column, such as Chironomids, midge larvae, and terrestrial insects that fall into the water.[11] Upon entering the marine environment, they switch to a diet of plankton and fish, with fish making up most of their diets after a certain size.[12] Adult coho feed on a vast variety of prey items that depend on the region they reside in during their second year at sea. Spawning habitats are small streams with stable gravel substrates.

Salmonid species on the west coast of the United States have experienced dramatic declines in abundance during the past several decades as a result of human-induced and natural factors.[13]

Conservation

Coho salmon, Tillamook State Forest, Oregon

The U.S. National Marine Fisheries Service (NMFS) has identified seven populations, called Evolutionary Significant Units (ESUs), of coho salmon in Washington, Oregon and California.[14][15] Four of these ESUs are listed under the U.S. Endangered Species Act (ESA).[16] These are the Lower Columbia River (threatened), Oregon Coast (threatened), Southern Oregon and Northern California Coasts (threatened), and Central California Coast (endangered). The long-term trend for the listed populations is still downward, though there was one recent good year with an increasing trend in 2001.[17]

The Puget Sound/Strait of Georgia ESU in Washington is an NMFS "Species of Concern".[18] Species of Concern are those species for which insufficient information prevents resolving the U.S. National Oceanic and Atmospheric Administration's concerns regarding status and threats and whether to list the species under the ESA.

On May 6, 1997, NMFS, on behalf of the Secretary of Commerce, listed as threatened the Southern Oregon/Northern California Coast coho salmon ESU.[19] The coho salmon population in the Southern Oregon/Northern California region has declined from an estimated 150,000–400,000 naturally spawning fish in the 1940s to fewer than 10,000 naturally producing adults today. These reductions are due to natural and man-made changes, including short-term atmospheric trends (such as El Niño, which causes extremes in annual rainfall on the northern California coast), predation by the California sea lion and Pacific harbor seal, and commercial timber harvesting.

More than 680,000 coho salmon returned to Oregon in 2009, double that of 2007. The Oregon Department of Fish and Wildlife required volunteers to herd fish into hatchery pens. Some creeks were reported to have so many fish, "you could literally walk across on the backs of coho," claimed a Portland television station. Lower temperatures in 2008 North Pacific waters brought in fatter plankton, which, along with greater outflows of Columbia River water, fed the resurgent populations. The 2009 run was so large, food banks were able to freeze 40 tonnes (39 long tons; 44 short tons) for later use.[20]

See also

Footnotes

  1. ^ Gehr, Susan; Bright. Karuk Dictionary.
  2. ^ "Coho Salmon" (PDF). Adfg.alaska.gov. Retrieved 18 March 2022.
  3. ^ "Coho Salmon (Oncorhynchus kisutch) - Office of Protected Resources - NOAA Fisheries". Nmfs.noaa.gov. Retrieved 2013-05-13.
  4. ^ Peter B. Adams; et al. (September 2007). "Coho Salmon Are Native South of San Francisco Bay: A Reexamination of North American Coho Salmon's Southern Range Limit". Fisheries. 32 (9): 441–451. doi:10.1577/1548-8446(2007)32[441:CSANSO]2.0.CO;2. ISSN 1548-8446.
  5. ^ "Coho Salmon, Oncorhynchus kisutch". Michigan.gov. Retrieved 18 March 2022.
  6. ^ "Danish fishermen catch salmon not from Denmark". The Local Denmark. The Local. 27 September 2017. Retrieved 26 October 2017.
  7. ^ a b Annual Statistics 2010: Commercial salmon catch by species and country North Pacific Anadromous Fish Commission Statistical Yearbook. Retrieved 2015 March 16. These numbers do not include fish taken in Russian waters by foreign fleet.
  8. ^ (Alaska Seafood Marketing Institute, 2003, p. 2)
  9. ^ Sisnyak, Nancy; Ragan, Ryan. "Fishing for Coho Salmon". Adfg.alaska.gov. Alaska Department of Fish and Game. Retrieved 8 December 2020.
  10. ^ Caputi, Gary (28 August 2020). "Fly Fishing for Silver Salmon". Saltwatersportsman.com. Retrieved 8 December 2020.
  11. ^ Johnson, James; Ringler, Neil (1980). "Diets of juvenile coho salmon (Oncorhynchus kisutch) and steelhead trout (Salmo gairdneri) relative to prey availability". Canadian Journal of Zoology. 58 (4): 553–558. doi:10.1139/z80-077.
  12. ^ Daly, Elizabeth; Brodeur, Richard; Weitkamp, Laurie (2009). "Ontogenetic Shifts in Diets of Juvenile and Subadult Coho and Chinook Salmon in Coastal Marine Waters: Important for Marine Survival?". Transactions of the American Fisheries Society. 138 (6): 1420–1438. doi:10.1577/T08-226.1.
  13. ^ "Pacific salmonids threats". U.S. National Marine Fisheries Service. 9 July 2021.
  14. ^ "Evolutionary Significant Units". U.S. National Marine Fisheries Service. 2005.
  15. ^ "Coho salmon ESUs". Archived from the original on 2013-02-02. Retrieved 2008-01-28.
  16. ^ "Endangered Species Act". Nmfs.noaa.gov. 9 July 2021.
  17. ^ "2005 status review report" (PDF). Archived from the original (PDF) on 2008-10-02. Retrieved 2008-01-28.
  18. ^ "Species of Concern". Nmfs.noaa.gov. 9 July 2021.
  19. ^ 62 Fed.Reg. 24588
  20. ^ Millman, Joel (January 21, 2010). "Fish Boom Makes Splash in Oregon". Wall Street Journal. Retrieved January 21, 2010.

References

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Coho salmon: Brief Summary

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The coho salmon (Oncorhynchus kisutch; Karuk: achvuun) is a species of anadromous fish in the salmon family and one of the five Pacific salmon species. Coho salmon are also known as silver salmon or "silvers". The scientific species name is based on the Russian common name kizhuch (кижуч).

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Diet

provided by World Register of Marine Species
Feeds on insects and crustaceans

Reference

North-West Atlantic Ocean species (NWARMS)

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Kennedy, Mary [email]

Distribution

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New Brunswick to Chesapeake Bay

Reference

North-West Atlantic Ocean species (NWARMS)

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Kennedy, Mary [email]

Habitat

provided by World Register of Marine Species
anadromous species; born in freshwater, spends most of life at sea, returns upriver to spawn

Reference

North-West Atlantic Ocean species (NWARMS)

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WoRMS Editorial Board
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Kennedy, Mary [email]

Habitat

provided by World Register of Marine Species
benthic

Reference

North-West Atlantic Ocean species (NWARMS)

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WoRMS Editorial Board
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Kennedy, Mary [email]

Introduction

provided by World Register of Marine Species
This species has been introduced or released in Dutch waters.

Reference

2. Fish, J. D. & Fish, S. (1996) A student's guide to the seashore. Second Edition. Cambridge University Press, Cambridge.

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