“Tipimegus kangulun, new species
Description of female—Head about 20 percent of total length, greatest width about 77 percent of length, rostrum constricted, narrow, exceeding middle of article 2 on antenna 1. Eyes small, clear of pigment or darkly stained. Article 1 of peduncle of antenna 1 about 0.8 times as long as wide, about 2.8 times as wide as article 2, ventral margin with about 7 setules, weakly produced dorsal apex with 3 setules, article 2 about 0.9 times as long as article 1, with ventral crescent of 13-14 setae; primary flagellum with 9 articles, about 0.55 times as long as peduncle, bearing several long aesthetascs; accessory flagellum almost as long as primary flagellum, with 8-9 articles. Spine formula on article 4 of antenna 2 = (2-3)-2-l 1-10-7 + 2-3 supernumerary spines, dorsal margin setose, ventral margin with 8-9 groups of 2-5 long to short setae, dense ventrodistal brush of setae; article 5 about 0.5 times as long as article 4, facial spine formula = 0, dorsal margin naked, ventral margin with 6 unpaired setae, 6 ventrodistal long to medium spine-setae, this row set facially; flagellum about 0.75 times as long as articles 4-5 of peduncle combined, with 10 articles. Mandibles with weak to medium palpar hump; right incisor with 4 teeth; left incisor with 5 humps in 2 branches; right lacinia mobilis bifid, distal branch shorter than proximal, subbifid proximal branch simple, pointed, with marginal denticles; left lacinia mobilis with 1 teeth; right rakers 10 plus one rudimentary; left rakers II; molars elongate, conical or columnar, each with one long apical spine, one short apical spine, one medium spine on side with vestigial nonarticulate partner; palp article 1 short, article 2 with 4-5 medium inner apical setae, 3 other shorter inner setae and 2-3 outer setae, article 3 about 0.9 times as long as article 2, oblique apex with 14 spine-setae, basodorsal formula = 3. Inner plate of maxilla 1 ordinary, bearing one long subapical pluseta, 2 apicolateral shorter setae; palp article 2 with 6-7 apical spines and 9 submarginal and facial setae. Inner plate of maxilla 2 shorter and much narrower than outer, outer lacking apicolateral setae, inner with 9-10 medial setae. Inner plate of maxilliped with 2 large, thick apical spines, 6 apicofacial setae. 7 medial setae, outer plate with 14 medial and apical spines, no apicolateral setae; palp article 1 lacking apicolateral seta, article 2 with 3 groups of 1-2 apicolateral setae, medial margin of article 2 moderately setose, article 3 with 11 facial setae, 4 lateral setae, article 4 with 3 apical accessory setules, nail absent. Coxa 1 strongly expanded distally, anterior margin almost straight; main ventral setae of coxae 1-4-14-16-15-32, posterior-most seta of coxae 1-3 shortest; anterior and posterior margins of coxa 4 strongly divergent, posterior margin oblique, almost straight, posterodorsal corner sharp, posterodorsal margin ordinary, almost straight, width-length ratio of coxa 4 = 27:29. Long posterior setae on article 2 of gnathopods 1-2 and pereopods 1-2 = 9-7-8-5, long anteriors = 20(+ 6 facial)-! 9-(15-20)-5, short anteriors = 2-3-2-5, no others. Gnathopods with thin hands broad apically, width ratios of articles 5-6 on gnathopods 1-2 = 30:26 and 27:25, length ratios = 63:50 and 63:45; palmar humps very small, palms transverse; article 5 of gnathopods 1-2 elongate, ovate, posterior margins rounded-flat. Pereopod 2 stouter than pereopod 1, especially article 4; facial setae formula on article 4 = 10-11 and 10, on article 5 = 5 and 5, main spine of article 5 extending to M. 75 on article 6, spine formula of article 6 = 6 + 9 or 7 + 9 plus huge middistal spine; acclivity on inner margin of dactyls of pereopods 1-2 sharp, produced as tooth, emergent setule short, midposterior pluseta short. Coxae 5-7 posteroventral seta formula = 32-32-20, composed of marginal and facial groups. Articles 4-5 of pereopods 3-4 broad, facial spine rows dense; facial ridge formula on article 2 of pereopods 3-5 = 0-1-1; width ratios of articles 2, 4, 5, 6 of pereopod 3 = 53:48:42:18, of pereopod 4 = 63:59:44:20, of pereopod 5 = 55:11:10:5; length ratios of pereopod 3 = 72:44:44:34, of pereopod 4 = 75:62:47:45, of pereopod 5 = 80:23:19:22; article 2 of pereopod 5 reaching middle of article 5, ventrally setose; medial apex on article 6 of pereopods 3 and 5 finely combed; dactyls of pereopods 3-4 elongate. Posteroventral corner of epimeron 1 rounded, posterior margin convex, setose, anteroventral face with 15 short to medium setae, midface with 11 medium set^e; posteroventral corner of epimeron 2 rounded, weakly protuberant, posterior margin convex, setose, facial setae = 1 5 + 2 midfacial, pairs and triads set vertically; posteroventral corner of epimeron 3 weakly protuberant, posterior margin straight, weakly serrate, setose, ventral margin with 16 setae mainly in posterior half, face with occasional sub marginal seta. Urosomite 1 with 4 paired sets of setae, each side with sets of 8 and 2, ventrally with sets of 3 and 2, articulation line complete; urosomite 3 protuberant dorsally. Rami of uropods 1-2 with articulate enlarged sharp apical nails, rami of uropod 2 with accessory nails, continuously spinose; outer ramus of uropod 1 with 9 dorsal spines, irregularly gapped apically, inner with 4 dorsal and 3 lateral (actually medial and lateral) spines; outer ramus of uropod 2 with 6-7 dorsal spines, inner with 2-3 dorsal spines; peduncle of uropod 1 with 3 apicolateral spines, medially with 4 spines; peduncle of uropod 2 with 4-5 dorsal spines, medially with one small apical spine; outer rami of uropods 1-2 and apicolateral corner of peduncle on uropod 2 with weak even comb. Peduncle of uropod 3 with 13-14 ventral spines, dorsally with one lateral spine, one medial spine and setule; rami submasculine, inner extending to M. 100+ on article 1 of outer ramus, apex with 2 setae, medial and lateral margins setose, article 2 of outer ramus short, 0.16, bearing 3 medium to long setae, medial margin of article 1 setose, lateral margin with 3-4 acclivities, spine formula = 2-2-2-2-( + 2), or 1-2-2-2-2, setal formula = 0. Telson ordinary, length-width ratio = 25: 29 or 26:27, not fully cleft, each apex of medium width, rounded, undulant, lateral acclivity shallow, weak, with lateral and medial spines separated by setule, occasionally lateral spine position paired, midlateral setules diverse. Cuticle with ordinary bulbar setules, surface punctate or bearing fine striations in form of linear fingerprint pattern, emergent setules branched.
Observations (female)—Unspined subapex of outer ramus of uropod 1 often with spine vestige subapically, juveniles as small as 3.7 mm also lacking spines apically or juvenile "j," 4.37 mm, with subapical spine and second spinule on face of nail, this spinule also present in superfemale; inner ramus of uropod 1 in juvenile with pair of accessory apical spines; protuberant base of spike on peduncle of uropod 1 becoming elongate in terminal adults.
Variations (female)—Female "w," 7.20 mm: Article 5 of pereopod 3 with 2 rows of posterior facial spines, article 6 of pereopod 4 with 4 rows of posterior facial spines; spine formula on outer ramus of uropod 3 = 2-3-2-2-2. Superfemale "z," 11.25 mm: Comb on apex of article 6 on pereopod 3 absent, article 2 of pereopod 5 with only 6 facial setae; inner ramus of uropod 1 so cylindrical that dorsal (medial) spines rotated out of sight medially (see illustration), peduncle of uropod 2 with 6 dorsal spines, outer ramus with 8, inner with 4.
Description of male—Eyes enlarged. Article 1 of antenna 1 with increased numbers of ventral setules, no medial fuzz; primary flagellum with 9 articles, one calceolus each on articles 1-4, aesthetascs well developed; accessory flagellum with 8 articles. Facial spine formula on article 4 of antenna 2 = 2-12-11-7, dorsomedial margin fuzzy, other setae shortened; article 5 with 8 dorsal sets of male setae and 7 calceoli, ventrodistal apex with 2 setules; flagellar formula = 45, 2-4, 6, 8 . . . 40 or 44, 2-5, 7, 9 . . . 39. Distal member of right lacinia mobilis poorly subbifid, inner margin of article 3 on mandibular palp with 2 accessory setae not in female. Inner plate of maxilliped with 2 main apical spines spread slightly more than in female, 4 medial setae; outer plate with 12 spines; palp article 2 with 3 lateral setae, article 3 with 6 lateral setae. Coxae 3-4 slightly to much longer than in female; ventral setal formula of coxae 1-4 = 10-13-13-27. Long posterior setae on article 2 of gnathopods 1-2 and pereopods 1-2 = 6-6-6-3, long anteriors = 6-9-14-7, short anteriors = 1-2-4-3, no others. Setal and spine formulas of pereopods 1-2 on article 4 = 5 + 5 (separated by gap from other distal setae), on article 5 = 5 and 5, on article 6 = 9 + 10 plus middistal spine, setae on posterodistal margin of article 4 shorter than in female. Article 2 of pereopods 3 and 5 slightly narrower than in female. Epimera 1-2 with numerous short posterodorsal setae, epimera 1-3 slightly broadened, posterior margin of epimeron 3 not shortened. Urosome ventrally with 3 paired sets of setae, lateral set = 7, ventral sets = 3 and 4. Spine formula of uropods: uropod 1 peduncle apicolateral = 3; uropod 2 peduncle dorsal = 6; dorsal spines on outer ramus of uropod 1=9, continuous to apex, but with slightly excessive gap between spines 1-2, of uropod 2 = 8; inner ramus of uropod 1 = 6-7 plus 3, of uropod 2 = 3. Ventral spines on peduncle of uropod 3 = 14-15; spine formula on article 1 of outer ramus = 0-1-1-2-2-2 or 0—1—1—1— 2-2-2-2, setal formula = 3-1-1-1-1-1 or 2-1-2-1- 1-1-1-1. Telson weakly elongate, broadened apically, distal spines scarcely shortened, midbase of
each lobe with numerous acute denticles.
Variations—Aberrant male, 8.0 mm, PPBES 960/3: Eyes scarcely enlarged but penial processes well developed; antenna 2 like that of female; uropod 3 almost fully masculine but antenna 1 and epimera like those of female.
Illustrations—Female maxilla 2 as in Tipimegus thalerus; several spines on ventral side of peduncle on uropod 3 omitted.
Holotype—AM, female "n," 7.60 mm.
Type-locality—SBS C4S4, 17 May 1972, off Malabar, New South Wales, Australia, 69 m, sand.
Voucher Material—SBS E2S4: female "w," 7.20 mm (illus.); male "y," 7.10 mm (illus.); male "u," 6.72 mm (illus.); juvenile "j," 4.37 mm (illus). CPBS 24N: female "z," 11.25 mm (illus.).
Relationship—This species differs from Tipimegus thalerus in the elongate dactyls of pereopods 3-4, the small size of the cuticular setules, the broad apices of the hands on the gnathopods, the widely spread setae on coxae 1-3, and the elongation of the lateral seta on article 2 of uropod 3 which in T. thalerus forms a shorter and distinctive spine. Tipimegus kangulun is characterized by the noncontinuous dorsal spination on uropod 1, particularly well developed in females and juveniles but becoming obscured in males. Male uropod 1 usually has some weak degree of asymmetry in the placement of apical spines and lacks any definite apical spine fully contiguous with the nail. In all but superfemales article 6 of pereopod 3 bears a distal comb which is absent in T. thalerus. Generally the spine forming part of the spike on the peduncle of uropod 1 is shorter in T. kangulun than in T. thalerus.
Remarks—A single male with stunted antenna 2 from JLB AUS 11, Western Australia, is provisionally identified with T. kangulun but the setae on coxae 1-3 are few and narrowly spread; this may be a new species and will be reserved until more material from Western Australia can be collected.
Material—CPBS, 3 samples from 2 stations (3); WPBES, 2 samples from 2 stations <5); PPBES, 14 samples from 10 stations (26) [5 doubtful from 2 samples]; AM, one sample (1); SBS, 6 samples from 4 stations (17).
Distribution—Victoria: Western Port and Port Phillip Bay, 2-13 m, sand. New South Wales: off Sydney, 53-150 m, sand.”
(Barnard & Drummond, 1978: 58-64)
