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Hygrophila (gastropod)

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Hygrophila is a taxonomic superorder of air-breathing freshwater snails, aquatic pulmonate gastropod mollusks within the clade Panpulmonata.[2] [1]

The families in this clade are basically air-breathing freshwater snails including freshwater limpets. The three families with the greatest number of species are the Lymnaeidae (pond snails), the Planorbidae (ramshorn snails) and the Physidae (pouch or bubble snails). These are found in ponds, creeks, ditches, and shallow lakes nearly worldwide.

The snails in this clade have their eyes located at the base of their tentacles, rather than at the tips, as in the true land snails Stylommatophora. They have shells that are thin, translucent, and relatively colorless, and they lack an operculum.

Taxonomy

1997 taxonomy

In the older taxonomy of the Gastropoda (Ponder & Lindberg, 1997) these families were placed in the suborder Basommatophora.

2005 taxonomy

Hygrophila has been a clade in the informal group Basommatophora within the Pulmonata.[3]

Clade Hygrophila:

Note: what was previously the family Ancylidae has been demoted in the new taxonomy to the status of tribe Ancylini Rafinesque, 1815 within the subfamily Planorbinae Rafinesque, 1815 of the family Planorbidae Rafinesque, 1815.

2010 taxonomy

Basommatophora (Siphonarioidea and Amphiboloidea and Hygrophila) have been found polyphyletic and so Jörger et al. (2010)[2] have moved Hygrophila to Panpulmonata.

2020 taxonomy

A comprehensive molecular phylogenetic analysis of Hygrophila based on 3112 sites of the large subunit and 5.8S ribosomal RNA genes resulted in a proposal of a new taxonomic revision of the group.[4]

Synonyms
  • Acroloxoidea Thiele, 1931: synonym of Lymnaeoidea Rafinesque, 1815
  • Planorboidea Rafinesque, 1815: synonym of Lymnaeoidea Rafinesque, 1815

References

  1. ^ a b MolluscaBase eds. (2020). MolluscaBase. Hygrophila. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=382243 on 2020-12-14
  2. ^ a b Jörger K. M., Stöger I., Kano Y., Fukuda H., Knebelsberger T. & Schrödl M. (2010). "On the origin of Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia". BMC Evolutionary Biology 10: 323. doi:10.1186/1471-2148-10-323.
  3. ^ Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
  4. ^ Saadi, Ahmed J.; Davison, Angus; Wade, Christopher M. (2020). "Molecular phylogeny of freshwater snails and limpets (Panpulmonata: Hygrophila)". Zoological Journal of the Linnean Society. 190 (2): 518–531. doi:10.1093/zoolinnean/zlz177.
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Hygrophila (gastropod): Brief Summary

provided by wikipedia EN

Hygrophila is a taxonomic superorder of air-breathing freshwater snails, aquatic pulmonate gastropod mollusks within the clade Panpulmonata.

The families in this clade are basically air-breathing freshwater snails including freshwater limpets. The three families with the greatest number of species are the Lymnaeidae (pond snails), the Planorbidae (ramshorn snails) and the Physidae (pouch or bubble snails). These are found in ponds, creeks, ditches, and shallow lakes nearly worldwide.

The snails in this clade have their eyes located at the base of their tentacles, rather than at the tips, as in the true land snails Stylommatophora. They have shells that are thin, translucent, and relatively colorless, and they lack an operculum.

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Hygrophila (Gastropoda) ( Spanish; Castilian )

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Hygrophila es un clado taxonómica de los caracoles de agua dulce que respiran aire, moluscos gasterópodos pulmonados acuáticos dentro del clado Panpulmonata.

Taxonomía

1997

Estas familias fueron colocados en el suborden Basommatophora (Taxonomía de Gastropoda (Ponder & Lindberg, 1997)).

2005

Ha sido un clado en el grupo informal Basommatophora dentro de la Pulmonata.

Clado Hygrophila:

2010

Basommatophora (Siphonarioidea y Amphiboloidea y Hygrophila) se ha encontrado en el grupo polifilético y así Jörger trasladó de Hygrophila a Panpulmonata.

Referencias

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Hygrophila (Gastropoda): Brief Summary ( Spanish; Castilian )

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Hygrophila es un clado taxonómica de los caracoles de agua dulce que respiran aire, moluscos gasterópodos pulmonados acuáticos dentro del clado Panpulmonata.

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Hygrophila (gastéropode) ( French )

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Les Hygrophila sont un super-ordre (ou un ordre selon les classifications) de mollusques gastéropodes de l'infra-classe des Pulmonata.

Description et caractéristiques

Ce taxon correspond à l'ancien groupe des Basommatophora, une fois ôtés les groupes discutés des Amphiboloidea et Siphonarioidea[1].

Taxonomie

Selon World Register of Marine Species (13 mai 2016)[1] :

Références taxinomiques

Notes et références

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Hygrophila (gastéropode): Brief Summary ( French )

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Les Hygrophila sont un super-ordre (ou un ordre selon les classifications) de mollusques gastéropodes de l'infra-classe des Pulmonata.

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Hygrophila (zoologia) ( Italian )

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Hygrophila Férussac, 1822 è un superordine di molluschi gasteropodi d'acqua dolce.[1]

Descrizione

Il raggruppamento comprende tutte le chiocciole d'acqua dolce in grado di respirare l'aria. Gli occhi di questi molluschi sono posti alla base delle loro appendici cefaliche, anziché alla punta di esse, come avviene negli Stylommatophora. Sono dotati di conchiglia sottile, traslucida, priva di opercolo, con un'ampia variabilità di forme (coniche, spiraliformi, discoidali, patelliformi), con avvolgimento destrorso o sinistrorso.[2]

Biologia

Questi molluschi assimilano l'ossigeno atmosferico attraverso la superficie interna del loro mantello, dove trattengono una bolla d'aria, che deve essere rinnovata periodicamente.[3]

Distribuzione e habitat

Il raggruppamento ha una distribuzione cosmopolita essendo diffuso in stagni, ruscelli, fossati e laghi poco profondi di tutti i continenti eccetto l'Antartide.

Tassonomia

Il superordine Hygrophila comprende due superfamiglie e otto famiglie:[1]

Nelle classificazioni precedenti le famiglie di questo taxon erano inquadrate tra i Basommatophora[4][5], raggruppamento rivelatosi polifiletico[6].

Alcune specie

Note

  1. ^ a b (EN) MolluscaBase eds. 2020, Hygrophila, in WoRMS (World Register of Marine Species). URL consultato il 3 novembre 2020.
  2. ^ (EN) Saadi A.J., Davison A., Wade C.M., Molecular phylogeny of freshwater snails and limpets (Panpulmonata: Hygrophila), in Zoological Journal of the Linnean Society, vol. 190, n. 2, 2020, pp. 518-531, DOI:10.1093/zoolinnean/zlz177.
  3. ^ Planorbidae, su acquariofiliaconsapevole.info.
  4. ^ (EN) Ponder W. & Lindberg D.R., Towards a phylogeny of gastropod molluscs; an analysis using morphological characters, in Zoological Journal of the Linnean Society, vol. 119, 1997, pp. 83-265.
  5. ^ (EN) Bouchet P., Rocroi J.-P., Classification and nomenclator of gastropod families, in Malacologia, 47(1-2), 2005, pp. 1-397.
  6. ^ (EN) Jörger K M., Stöger I., Kano Y., Fukuda H., Knebelsberger T. & Schrödl M., On the origin of Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia, in BMC Evolutionary Biology, vol. 10, 2010, p. 323, DOI:10.1186/1471-2148-10-323.

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Hygrophila (zoologia): Brief Summary ( Italian )

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Hygrophila Férussac, 1822 è un superordine di molluschi gasteropodi d'acqua dolce.

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Hygrophila (clade) ( Dutch; Flemish )

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De Hygrophila zijn een clade van zoetwaterslakken (Gastropoda).

Taxonomie

Jörger et al. (2010)[1] hebben de hoofdgroepen binnen de Heterobranchia opnieuw ingedeeld. Ze creëerden de nieuwe clades Euopisthobranchia en Panpulmonata.[1]

Onderstaande cladogram toont de relatie binnen de Heterobranchia zoals voorgesteld door Jörger et al. (2010):[1]

Heterobranchia

Lower Heterobranchia (inclusief Acteonoidea) - Lower Heterobranchia vormt geen clade in de studie door Jörger et al. (2010):[1]


Euthyneura

Nudipleura



Euopisthobranchia

Umbraculoidea





Runcinacea




Anaspidea



Pteropoda





Cephalaspidea s.s.




Panpulmonata

Siphonarioidea



Sacoglossa






Glacidorboidea




Amphiboloidea



Pyramidelloidea






Hygrophila




Acochlidiacea


Eupulmonata

Stylommatophora




Systellommatophora




Ellobioidea




Otinoidea



Trimusculoidea













In detail is de taxonomische indeling van de clade als volgt:[2]

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Hygrophila (clade): Brief Summary ( Dutch; Flemish )

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De Hygrophila zijn een clade van zoetwaterslakken (Gastropoda).

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Hygrophila (orde) ( Dutch; Flemish )

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Hygrophila is de naam van een orde van slakken (Gastropoda). Deze indeling wordt niet meer door alle moderne auteurs geaccepteerd; veel auteurs maken tegenwoordig gebruik van een indeling in clades.[1][2]

Superfamilies

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Hygrophila (orde): Brief Summary ( Dutch; Flemish )

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Hygrophila is de naam van een orde van slakken (Gastropoda). Deze indeling wordt niet meer door alle moderne auteurs geaccepteerd; veel auteurs maken tegenwoordig gebruik van een indeling in clades.

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喜濕螺總目 ( Chinese )

provided by wikipedia 中文维基百科

喜濕螺類(學名:Hygrophila)是生物分类学的一個支序,屬於異鰓類直神經類泛有肺類支序,包括有所有會呼吸的淡水螺及所有水生的真有肺類軟體動物,由Jörger et al.於2010年10月建立[2]。2017年12月,本支序成為了總目分類單元,即喜濕螺總目[3]

本支序之下的所有科皆為會呼吸的淡水螺,包括淡水帽貝。 本分類最多不同物種的三個科分別為:

這些物種可在池塘、小溪、溝渠及淺湖中發現,遍及全世界。

本支序物種的眼睛均長在觸角的底部,而不是頂部。這一點跟陸生的柄眼類物種(Stylommatophora)相當。牠們的殼薄而透明,看起來似乎無色,而且沒有口蓋覆蓋。

分類

1997年分類

根據腹足纲分类表 (1997年),現時本支序的科原來都放在基眼亚目(Basommatophora Keferstein in Bronn, 1864)之下。

2005年分類

根據2005年的《布歇特和洛克羅伊的腹足類分類》,原來的基眼亚目在2005年的分類裡成為了有肺目之下的一個非正式群組,而本支序仍然在基眼亞目之下[4]

喜濕螺類支序:

2010年分類

基眼類(即:SiphonarioideaAmphiboloidea及喜濕螺類)確認為並系群,為保持分類的單系性,Jörger et al. (2010)[2]把本支序連同其他相關的分類合併成為泛有肺類(Panpulmonata)支序。

2017年分類

根據2017年的《布歇特等人的腹足類分類》,本分類支序成為了Tectipleura Subterclass之下的一個總目分類單元[3][1]。過往的基眼目被認為是喜濕螺總目的異名,儘管昔日的基眼目所包含的分類單元更多。

原來的四個總科現時綜合成為下列兩個總科[3]

參考文獻

  1. ^ 1.0 1.1 WoRMS. Hygrophila. 世界海洋物种目录(庫存版本). [2018-04-02]. (原始内容存档于2018-02-02).
  2. ^ 2.0 2.1 Jörger, K. M.; Stöger, I.; Kano, Y.; Fukuda, H.; Knebelsberger, T.; Schrödl, M. On the origin of Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia. BMC Evolutionary Biology. 2010, 10: 323. doi:10.1186/1471-2148-10-323 (英语).
  3. ^ 3.0 3.1 3.2 Bouchet, P.; Rocroi, J.P.; Hausdorf, B.; Kaim , A.; Kano , Y.; Nützel , A.; Parkhaev, P.; Schrödl, M.; Strong , E.E. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 2017, 61 (1-2): 1–526. doi:10.4002/040.061.0201 (英语).
  4. ^ Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á.; Warén, A. Bouchet P.; Rocroi J.-P., 编. Classification and nomenclator of gastropod families. Malacologia: International Journal of Malacology (Hackenheim, Germany / Ann Arbor, USA: ConchBooks). 2005, 47 (1-2): 397 pp. ISBN 3-925919-72-4. ISSN 0076-2997.
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喜濕螺總目: Brief Summary ( Chinese )

provided by wikipedia 中文维基百科

喜濕螺類(學名:Hygrophila)是生物分类学的一個支序,屬於異鰓類直神經類泛有肺類支序,包括有所有會呼吸的淡水螺及所有水生的真有肺類軟體動物,由Jörger et al.於2010年10月建立。2017年12月,本支序成為了總目分類單元,即喜濕螺總目。

本支序之下的所有科皆為會呼吸的淡水螺,包括淡水帽貝。 本分類最多不同物種的三個科分別為:

椎实螺科 扁卷螺科 膀胱螺科

這些物種可在池塘、小溪、溝渠及淺湖中發現,遍及全世界。

本支序物種的眼睛均長在觸角的底部,而不是頂部。這一點跟陸生的柄眼類物種(Stylommatophora)相當。牠們的殼薄而透明,看起來似乎無色,而且沒有口蓋覆蓋。

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히그로필라류 ( Korean )

provided by wikipedia 한국어 위키백과

히그로필라류(Hygrophila)는 공기 호흡하는 민물 달팽이 분류군 중의 하나로, 범유폐류에 속하는 수생 유폐류복족류 연체동물이다.[1]

이 분류군에 있는 과들은 민물 삿갓조개류를 포함하여 기본적으로 공기 호흡하는 민물 달팽이이다. 종의 수가 가장 많은 3개 과는 물달팽이과, 또아리달팽이과 그리고 왼돌이물달팽이과이다. 이 종들은 못, 샛강, 개천, 얕은 호수 등 거의 전 세계에서 발견된다.

이 분류군에 있는 달팽이들은 눈이 육상의 참달팽이 병안류처럼 촉각 끝단보다는 촉각 하부에 위치한다. 얇고, 반투명의 비교적 무채색의 조가비 껍질을 지니고 있으며, 덮개딱지가 없다.

1997년 분류

이전 분류 체계인 1997년 복족류 분류(폰더 & 린더버그)는 이들 과를 기안아목으로 분류했다.

2005년 분류

히그로필라류를 유폐류 내의 비공식군 기안류에 속하는 분류군의 하나로 분류했다.[2]

참고: 이전 분류 체계의 민물삿갓조개과 (Ancylidae)는 새 분류 체계에서 또아리물달팽이과(Planorbidae)의 또아리물달팽이아과(Planorbinae)에 속하는 민물삿갓조개족(Ancylini)으로 강등되었다.

2010년 분류

기안류(고랑따개비상과, 암피볼라상과, 히그로필라류)가 다계통군임이 밝혀짐에 따라, 2010년 요르거(Jörger)와 그의 공저자들은[1] 히그로필라류를 범유폐류로 옮겼다.

2010년 요르거(Jörger) 등이 제안한 분기도는 이새류의 계통분류학적 관계를 아래와 같이 보여준다.[1]

이새류

하이새류 (비자고둥상과 포함) - 하이새류는 요르거(Jörger) 등의 연구(2010년)에서는 별도의 분기군을 형성하지 않는다.[1]

직신경류

나측류

    진후새류

산각류

       

룬키나류

     

무순류

   

익족류 (무각익족류, 유각익족류)

       

두순류 s.s.

      범유폐류    

고랑딱개비류

   

낭설류

         

글라키도르비스류

     

암피볼라류

   

회오리고둥류

         

히그로필라류

     

아코클리디움류

진유폐류

병안류

     

수안류

     

대추귀고둥류

     

오티나류

   

트리무스쿨루스류

                       

각주

  1. Jörger K. M., Stöger I., Kano Y., Fukuda H., Knebelsberger T. & Schrödl M. (2010). "On the origin of Acochlidia and other enigmatic euthyneuran gastropods, with implications for the systematics of Heterobranchia". BMC Evolutionary Biology 10: 323. doi:10.1186/1471-2148-10-323.
  2. Bouchet P.; Rocroi J.; Frýda J.; Hausdorf B.; Ponder W.; Valdés Á.; Warén A. (2005). “복족류 분류 (부쉐 & 로크루아, 2005년)”. 《Malacologia: International Journal of Malacology》 (Hackenheim, Germany: ConchBooks) 47 (1-2): 1–397. ISBN 3925919724. ISSN 0076-2997.
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Classification

provided by World Register of Marine Species
This is what is left of the basommatophores when removing the controversial groups Amphiboloidea and Siphonarioidea. Original spelling "Hygrophiles" (vernacular), latinized by Herrmannsen (1846 [in 1846-1852]: 547). Established as a suborder containing the family Lymnaeidae. Ranked by Starobogatov (1970b: 46) as an order containing the superfamilies Chilinoidea, Latioidea, and Lymnaeoidea.
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