The parrot family is so ancient that there is no information in the fossil record as to their closest relative. Through electrophoretic analysis of eggshells and egg whites it is currently thought that pigeons are parrots closest relatives (Sick, 1993).
Scarlet macaws communicate with a variety of vocalizations and postures. Mated pairs are engaging in tactile communication when preening.
Scarlet macaws have excellent vision and hearing.
Communication Channels: visual ; tactile ; acoustic
Perception Channels: visual ; tactile ; acoustic ; chemical
The habitat of scarlet macaws is threatened due to forest destruction in the deep rainforest habitats where they live. Also, poachers seek out the parrots and will even cut down the tree where the nest is located to access the young or will shoot the adults for food (Ridgely and Gwynne, 1989). Cutting down trees to access macaws limits the number of places to nest and this practice will eventually limit the numbers of young raised.
Efforts have been made to slow population declines of scarlet macaws. The World Parrot Trust was formed in 1989 to protect parrots in their natural environment. Also, there is a trend towards breeders providing feathers from the birds that they sell so that other macaws will not be poached solely for feathers (Sick, 1993).
Nine of the sixteen species of macaws are listed on Appendix I of CITES, including scarlet macaws. Reproductive rates in the wild are low for a number of reasons, including a natural scarcity of suitable nesting sites. Some conservation organizations have found that macaw species will nest in artificial cavities and have supplemented certain areas with artifical nesting boxes.
US Federal List: no special status
CITES: appendix i
IUCN Red List of Threatened Species: least concern
There are no negative impacts of macaw species on humans.
The illegal, international parrot trade brings in large revenues each year due to the high demand for these colorful birds. An individual scarlet macaw may be sold for more than $1,000. Also, birds may be hunted for meat and the feathers traded for money. Current law dictates that it is illegal to trade in Ara macao individuals due to their CITES Appendix I status (Ridgely and Gwynne, 1989).
Scarlet macaws are more valuable to people as valuable and beautiful members of tropical forests, where their presence has significant ecotourism benefits.
Positive Impacts: pet trade ; body parts are source of valuable material; ecotourism
Scarlet macaws are important seed predators of large tree fruits in the ecosystems in which they live. They may influence the generation of forest tree species.
Scarlet macaws primarily eat fruit and nuts, and will occasionally supplement their diet with nectar and flowers. Ara macao individuals are known to consume fruits before they are ripe. Premature fruits have a tougher skin and pulp that is difficult to access unless the bird has a beak large enough to tear into it. By accessing these fruits before they are available to other animals, they may gain a competitive advantage. Scarlet macaws are also able to break open the toughest nuts. Parrots have more movement in their beaks than do other birds, which allows for a more powerful bill. This ability creates an important food resource for the parrots because not a lot of other animals are able to access such a large variety of nuts (Aditays, 2000). There are structures on the inside of their beaks that allow scarlet macaws to press the hard seed between their tongue and palate and grind the seed so that it can be digested (Sick, 1993).
Scarlet macaws occasionally consume clay found on the banks of rivers. This aids in digestion of the harsh chemicals such as tannins that are ingested when eating premature fruit (Aditays, 2000).
Plant Foods: seeds, grains, and nuts; fruit; nectar; flowers
Primary Diet: herbivore (Frugivore , Granivore )
Ara macao is found in southern Mexico, Central America, and South America. In South America, the species is found as far south as northeastern Argentina. Ara macao is most common throughout the Amazon basin.
Biogeographic Regions: nearctic (Native ); neotropical (Native )
Scarlet macaws are found high in the canopy of rainforest habitats below 1,ooo m (Slud, 1964).
Range elevation: 1000 (high) m.
Habitat Regions: tropical ; terrestrial
Terrestrial Biomes: rainforest
Large macaws may live up to 75 years in captivity. Typical lifespans in the wild and in captivity are closer to 40 to 50 years.
Range lifespan
Status: wild: 33 (high) years.
Typical lifespan
Status: wild: 40 to 50 years.
Average lifespan
Status: captivity: 64.0 years.
Scarlet macaws are brightly colored birds with feathers ranging in color bands from scarlet on their head and shoulders, to yellow on their back and mid wing feathers and blue on the wing tips and tail feathers. The face has short white feathers. This area surrounds the light yellow colored eyes. The long, thick beak is light on the top and dark black on the bottom. The legs and feet are also black (Aditays, 2000).
Body length is approximately 89 cm, with the tail comprising approximately 1/3 - 1/2 of this. Tail feathers of males may be longer than females. Also, bills of males may be slightly larger (Sick, 1993).
Average mass: 1200 g.
Average length: 89 cm.
Other Physical Features: endothermic ; homoiothermic; bilateral symmetry
Sexual Dimorphism: sexes alike
Average mass: 1040 g.
As adults scarlet macaws may escape most predation by virtue of their size and flight. Young may be taken in the nest by arboreal predators such as snakes, monkeys, and other small carnivores. Adults and fledglings may also be taken by large cats, such as jaguars, and by eagles and hawks.
Scarlet macaws form monogamous pair bonds that last for life.
Mating System: monogamous
Breeding in Ara macao occurs about every one to two years. The clutch size is 2 to 4 white, rounded eggs with an incubation period of 24 to 25 days. Females mainly incubate the eggs. After hatching, the young may stay with their parents for one to two years. The male feeds the young by regurgitating and liquefying food (Sick, 1993). The parents will not raise another set of eggs until the previous young have become independent (Aditays, 2000). Scarlet macaws reach sexual maturity at three or four years of age (Sick, 1993).
Breeding interval: Breeding occurs every one to two years.
Breeding season: Breeding may occur year-round.
Range eggs per season: 2 to 4.
Range time to hatching: 24 to 25 days.
Range time to independence: 1 to 2 years.
Average age at sexual or reproductive maturity (female): 3-4 years.
Average age at sexual or reproductive maturity (male): 3-4 years.
Key Reproductive Features: iteroparous ; year-round breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; oviparous
Both male and female scarlet macaws care for their young. Scarlet macaws have an extended period of dependence on their parents, with perhaps some significant learning occuring before they become sexually mature and independent.
Parental Investment: no parental involvement; altricial ; pre-fertilization (Protecting: Male, Female); pre-hatching/birth (Protecting: Female); pre-weaning/fledging (Provisioning: Male, Protecting: Male, Female); post-independence association with parents; extended period of juvenile learning
The scarlet macaw (Ara macao) is a large yellow, red and blue Neotropical parrot native to humid evergreen forests of the Americas. Its range extends from southeastern Mexico to Peru, Ecuador, Colombia, Bolivia, Venezuela and Brazil in lowlands of 500 m (1,600 ft) (at least formerly) up to 1,000 m (3,300 ft), the Caribbean island of Trinidad, as well as the Pacific island of Coiba.[1] Formerly, the northern extent of its range included southern Tamaulipas. In some areas, it has suffered local extinction because of habitat destruction, or capture for the parrot trade, but in other areas, it remains fairly common. It is the national bird of Honduras. Like its relative the blue-and-yellow macaw, the scarlet macaw is a popular bird in aviculture as a result of its striking plumage.
The scarlet macaw was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae under the binomial name Psittacus macao.[3] The scarlet macaw is now placed in the genus Ara (Lacépède, 1799), one of 6 genera of Central and South American macaws.[4]
The two subspecies can be recognized by size and color detail in the feathers on the wings:[4]
In May 2013, it was announced that a team of scientists, led by Dr. Christopher M. Seabury and Dr. Ian Tizard of Texas A&M University had sequenced the complete genome of the scarlet macaw.[5][6] Based on this genome, species-specific microsatellite genetic markers were developed to aid genetic studies throughout the range of the species.[7] These genetic markers were later validated[8] on the trace amount of DNA acquired from feathers, and applied to study red-and-green macaws in a tropical landscape where DNA can degrade very quickly.[9] These markers were proved to be useful to study their population genetics[8] and identification of individuals in the landscape of the Peruvian Amazon.[10]
It is about 81 centimeters (32 in) long, of which more than half is the pointed, graduated tail typical of all macaws, though the scarlet macaw has a larger percentage of tail than the other large macaws. The average weight is about 1 kilogram (2 lb 3 oz). The plumage is mostly scarlet, but the rump and tail-covert feathers are light blue, the greater upper wing coverts are yellow, the upper sides of the flight feathers of the wings are dark blue as are the ends of the tail feathers, and the undersides of the wing and tail flight feathers are dark red with metallic gold iridescence. Some individuals may have green in the wings.
There is bare white skin around the eye and from there to the bill. Tiny white feathers are contained on the face patch. The upper mandible is mostly pale horn in color and the lower is black. Juveniles have dark eyes; adults have light yellow eyes.
It is frequently confused with the slightly larger green-winged macaw, which has more distinct red lines in the face and no yellow in the wing.
Scarlet macaws make very loud, high and sometimes low-pitched, throaty squawks, squeaks and screams designed to carry many kilometers to call for their groups.
The scarlet macaw can live up to 75[11] or even 90[12] years in captivity, although a more typical lifespan is 40 to 50 years.[12][11]
A typical sighting is of a single bird or a pair flying above the forest canopy, though in some areas flocks can be seen. They often gather at clay licks.[13][14] Scarlet macaws communicate primarily through raucous honks; however, vocal communication is highly variable, and captive macaws are known to be adept mimics of human speech.
Wild scarlet macaws feed on fruits, nuts, seeds, flowers and nectar.[15]
They also love to eat insects and larvae. They are seen feeding heavily on bugs, snails and foliage. Snails and bugs are great source of protein, as they need additional protein during breeding seasons.
In Costa Rica's Central Pacific they have learned to feed on introduced Teak trees (Tectona grandis) and Almond Beach Trees. Local non-profit organizations have planted hundreds of those trees along the coastline from the Tárcoles River basin to Esterillos Beach which had helped increase the population drastically. The combined efforts and the correct ecotourism has also an important role in the conservation of such majestic birds. Tour companies along the Tarcoles River and its mangroves have bet on the importance of birdwatching as an asset for the growth on its population.
While comparatively docile at most times of the year, scarlet macaws may be formidably aggressive during periods of breeding. Scarlet macaws are monogamous birds, with individuals remaining with one partner throughout their lives. The hen lays two or three white eggs in a large tree cavity.[16] The female incubates the eggs for about five weeks, and the chicks fledge from the nest about 90 days after hatching[17] and leave their parents about a year later. Juveniles reach sexual maturity at five years of age.
The South American range is extensive and covers the Amazon forest, extending to Peru east of the Andes, to Bolivia.[1] In Bolivia, it is very present in the Aquicuana Reserve, located in the Beni Department, near the city of Riberalta, the Capital of the Bolivian Amazon.
In Central America, the range extends from extreme eastern and southern Mexico and Panama through Guatemala, El Salvador, Honduras and Belize, the island of Coiba and infrequently on the mainland of Panama, and in Costa Rica in isolated regions on the Pacific Coast; the Nicoya Peninsula the Carara National Park and Peninsula de Osa.
The scarlet macaw has escaped or been deliberately released in to Florida, but there is no evidence that the population is breeding and may only persist due to continuing releases or escapes.[18] An introduced population exists in Puerto Rico.
Scarlet macaws inhabit humid lowland subtropical rain forests, open woodlands, river edges, and savannas.
The habitat of scarlet macaws is also considered to have the greatest latitudinal range for any bird in the genus Ara, as the estimated maximum territorial range covers 6,700,000 km2. Nevertheless, the scarlet macaw's habitat is fragmented, and the bird is mostly confined to tiny populations scattered throughout its original range in Middle America.[19] However, as they still occur in large numbers over most of their original range in South America, the species is classified by IUCN as least concern.[1] Its wild population is currently estimated between 20,000 and 50,000 individuals.
Commercial international trade in the species (including parts and derivatives) is prohibited by the bird's listing under CITES Appendix 1 due to predation for the pet and cage bird trade.[20]
The northern subspecies, A. m. cyanopterus, is listed as endangered by the USFWS.[21] The USFWS estimates that only 2,000–3,000 birds of the northern subspecies remain in the wild.[22]
The scarlet macaw is an early example of a parrot breeding in captivity. Captive breeding occurred in Northern Mexico at Paquime (also called Casas Grandes) and very likely Southwest New Mexico Mimbres Valley in the 11th century. Breeding pens, perches, bones, and eggshell fragments have been uncovered. The straightforward nature of scarlet macaw breeding and the value of their plumes in trade created a market for trade wherein the animals were used in religious rites north to the Colorado Plateau region.[23]
Today the scarlet macaw is found worldwide in captivity, but is best represented in captivity in the Americas. Captive techniques developed from the pet trade have positively affected wild populations: in areas with low macaws populations, the "extra" babies that typically die in the nest may be reared by humans hands and released into the wild to bolster the population, as has been done by the Tambopata Macaw Project. Their captive diet, egg incubation, assisted hatching, hand rearing, co-parenting, parent-rearing, fledgling, maturation, and breeding are well understood within the avicultural community (AFA Watchbird magazine).
The scarlet macaw (Ara macao) is a large yellow, red and blue Neotropical parrot native to humid evergreen forests of the Americas. Its range extends from southeastern Mexico to Peru, Ecuador, Colombia, Bolivia, Venezuela and Brazil in lowlands of 500 m (1,600 ft) (at least formerly) up to 1,000 m (3,300 ft), the Caribbean island of Trinidad, as well as the Pacific island of Coiba. Formerly, the northern extent of its range included southern Tamaulipas. In some areas, it has suffered local extinction because of habitat destruction, or capture for the parrot trade, but in other areas, it remains fairly common. It is the national bird of Honduras. Like its relative the blue-and-yellow macaw, the scarlet macaw is a popular bird in aviculture as a result of its striking plumage.