↑Herron, Matthew D.; Castoe, Todd A.; Parkinson, Christopher L. (2004). “Sciurid phylogeny and the paraphyly of Holarctic ground squirrels (Spermophilus)”. 《Molecular Phylogenetics and Evolution》 31 (3): 1015–30. doi:10.1016/j.ympev.2003.09.015. PMID15120398.