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Onthophagus
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Onthophagus (Palaeonthophagus) marginalis (Gebler 1817)
collect
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maps
names
Scientific Names
Preferred Names
Onthophagus (Palaeonthophagus) marginalis (Gebler 1817)
Species recognized by
EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Onthophagus marginalis
Species recognized by
wikipedia VI
,
Portugal Species List
,
Spain Species List
,
wikipedia ID
,
North Atlantic Species List
,
wikipedia AST
,
NCBI
, and
wikipedia MIN
Onthophagus marginalis
(Gebler 1817)
Species recognized by
GBIF classification
Onthophagus
(
Palaeonthophagus
)
marginalis
(Gebler 1817)
Species recognized by
EOL Dynamic Hierarchy 1.1
and
EOL Dynamic Hierarchy 1.1
Onthophagus (Palaeonthophagus) marginalis (Gebler 1817)
Species recognized by
Catalogue of Life 26 March 2024
and
Dynamic Hierarchy Version 2.2.3
Onthophagus marginalis
Species recognized by
wikidata_hierarchy tar gz
,
wikipedia ES
,
wikipedia PT
,
Global Biotic Interactions
, and
iNaturalist data coverage
Onthophagus marginalis
Recognized by
BHL data coverage
Onthophagus marginalis (Gebler 1817)
Species recognized by
GBIF data coverage
Alternative Names
Onthophagus circumscriptus Faldermann 1835
Not accepted according to
EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Onthophagus marginalis nigrimargo Goidanich 1926
Not accepted according to
EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Onthophagus marmoratus Ménétriès 1832
Not accepted according to
EOL Dynamic Hierarchy April 2022
Reference taxon from World Scarabaeidae Database in Species 2000 & ITIS Catalogue of Life
Onthophagus circumscriptus
Faldermann 1835
Synonym according to
EOL Dynamic Hierarchy 1.1
and
EOL Dynamic Hierarchy 1.1
Onthophagus marginalis nigrimargo
Goidanich 1926
Synonym according to
EOL Dynamic Hierarchy 1.1
and
EOL Dynamic Hierarchy 1.1
Onthophagus marmoratus
Ménétriès 1832
Synonym according to
EOL Dynamic Hierarchy 1.1
and
EOL Dynamic Hierarchy 1.1
Onthophagus obtusicornis
Motschulsky 1859
Heterotypic synonym according to
GBIF classification
Copris marginalis Gebler 1817
Synonym according to
Catalogue of Life 26 March 2024
Onthophagus circumscriptus Faldermann 1835
Not accepted according to
Dynamic Hierarchy Version 2.2.3
Onthophagus marginalis nigrimargo Goidanich 1926
Not accepted according to
Dynamic Hierarchy Version 2.2.3
Onthophagus marmoratus Ménétriès 1832
Not accepted according to
Dynamic Hierarchy Version 2.2.3
Common Names
There are no common names associated with this taxon.
Curated hierarchies for Onthophagus (Palaeonthophagus) marginalis (Gebler 1817)
EOL Dynamic Hierarchy April 2022
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Onthophagus
Onthophagus (Palaeonthophagus) marginalis (Gebler 1817)
(this page)
Onthophagus (Palaeonthophagus) marginalis przewalski Kabakov 2006
Onthophagus (Amphionthophagus) melitaeus (Fabricius 1798)
Onthophagus (Colobonthophagus) piffli Petrovitz 1961
Onthophagus (Colobonthophagus) thai Kabakov 1994
Onthophagus (Colobonthophagus) triceratops Arrow 1913
Onthophagus (Furconthophagus) karenensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus (Hikidaeus) leknamnaous Masumoto & Ochi 2015
Onthophagus (Hikidaeus) sauteri Gillet 1924
Onthophagus (Indachorius) baenzigeri Masumoto, Ochi & Hanboonsong 2007
Onthophagus (Indachorius) heterorrhinus Lansberge 1885
Onthophagus (Indachorius) subsapaensis Kabakov 1994
Onthophagus (Matashia) fossor Arrow 1931
Onthophagus (Micronthophagus) collinsi Krikken & Huijbregts 1987
Onthophagus (Micronthophagus) liui Masumoto, Ochi & Lee 2014
Onthophagus (Micronthophagus) vanofwegeni Krikken & Huijbregts 2008
Onthophagus (Onthophagiellus) thanwaakhomus Masumoto 1992
Onthophagus (Palaeonthophagus) clitellifer Reitter 1894
Onthophagus (Palaeonthophagus) kolenatii Reitter 1892
Onthophagus (Palaeonthophagus) nikolajevi Kabakov 2006
Onthophagus (Palaeonthophagus) parmatus Reitter 1892
Onthophagus (Palaeonthophagus) pygargus Motschulsky 1845
Onthophagus (Palaeonthophagus) rachelis Martin-Piera 1985
Onthophagus (Palaeonthophagus) vacca (Linnaeus 1767)
Onthophagus (Paraphanaeomorphus) jeannelianus Paulian 1945
Onthophagus (Parentius) nigellus (Illiger 1803)
Onthophagus (Phanaeomorphus) gagatoides Kabakov 2006
Onthophagus (Sinonthophagus) rugulosus Harold 1886
Onthophagus (Trichonthophagus) mediofuscatus D' Orbigny 1908
Onthophagus (Trichonthophagus) triacanthus Castelnau 1840
Onthophagus abyssinicus Gillet 1925
Onthophagus annulopunctatus Krikken & Huijbregts 2017
Onthophagus atridorsis D' Orbigny 1902
Onthophagus atriglabrus Howden & Gill 1987
Onthophagus axillaris Boheman 1860
Onthophagus baiyericus Krikken & Huijbregts 2013
Onthophagus batesi Howden & Cartwright 1963
Onthophagus birugifer D' Orbigny 1908
Onthophagus bituberifrons Josso 2018
Onthophagus bocandei D' Orbigny 1904
Onthophagus carpophilus Pereira & Halffter 1961
Onthophagus centurio Lansberge 1885
Onthophagus ceylonicus Harold 1859
Onthophagus chryses Bates 1887
Onthophagus civettae Cambefort 1984
Onthophagus convexicollis Boheman 1858
Onthophagus cribratus Lansberge 1883
Onthophagus crinitus Harold 1869
Onthophagus cruciger MacLeay 1888
Onthophagus cuniculus MacLeay 1864
Onthophagus diversus Reiche 1847
Onthophagus ebenicolor D' Orbigny 1902
Onthophagus egenus Harold 1877
Onthophagus evae Balthasar 1944
Onthophagus excisiceps D' Orbigny 1902
Onthophagus flavimargo D' Orbigny 1902
Onthophagus foulliouxi Cambefort 1971
Onthophagus gabonensis Walter 1989
Onthophagus gibbus D' Orbigny 1913
Onthophagus grandivigilans Masumoto 1995
Onthophagus gurburra Storey & Weir 1990
Onthophagus hemipygus Frey 1964
Onthophagus hircus Billberg 1815
Onthophagus hulstaerti Boucomont 1932
Onthophagus imbutus Sharp 1875
Onthophagus impunctatus D' Orbigny 1904
Onthophagus impurus Harold 1868
Onthophagus inermicollis D' Orbigny 1908
Onthophagus juncticornis D' Orbigny 1908
Onthophagus koebelei Blackburn 1903
Onthophagus mexicanus Bates 1887
Onthophagus miles D' Orbigny 1905
Onthophagus mimikanus Balthasar 1969
Onthophagus mongana Storey & Weir 1990
Onthophagus naevuliger D' Orbigny 1908
Onthophagus nigriceps Raffray 1877
Onthophagus novaeirlandiae Balthasar 1969
Onthophagus orientalis Harold 1868
Onthophagus palamoui Biswas & Chatterjee 1986
Onthophagus paravinctus Tagliaferri & Moretto 2012
Onthophagus pseudoconvexicollis Bai & Yang 2016
Onthophagus puberulus D' Orbigny 1902
Onthophagus pugnax Harold 1868
Onthophagus pullus Roth 1851
Onthophagus pupillatus Kolbe 1886
Onthophagus pygidialis Lansberge 1883
Onthophagus sansibaricus Harold 1878
Onthophagus savanicola Cambefort 1984
Onthophagus sculptilis Gerstaecker 1871
Onthophagus solomonensis Krikken & Huijbregts 2012
Onthophagus spiculatus Boucomont 1914
Onthophagus tamworthi Blackburn 1903
Onthophagus tatsienluensis Balthasar 1942
Onthophagus tessulatus Harold 1871
Onthophagus thainuaensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus tirapensis Biswas & Chatterjee 1985
Onthophagus toraut Krikken & Huijbregts 2011
Onthophagus tricariniger D' Orbigny 1902
Onthophagus tripolitanus Heyden 1890
Onthophagus tshuapae Balthasar 1964
Onthophagus undaticeps D' Orbigny 1902
Onthophagus viridivinosus Kohlmann & Solis 2001
2069 additional siblings truncated for brevity.
See the
resource file
for a full list.
Catalogue of Life 26 March 2024
Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Scarabaeinae Latreille 1802
Onthophagini Streubel 1846
Onthophagus Latreille 1802
Onthophagus subgen. Palaeonthophagus Zunino 1979
Onthophagus (Palaeonthophagus) marginalis (Gebler 1817)
(this page)
Onthophagus (Palaeonthophagus) marginalis marginalis (Gebler 1817)
Onthophagus (Palaeonthophagus) marginalis przewalski Kabakov 2006
Onthophagus (Palaeonthophagus) aerarius Reitter 1892
Onthophagus (Palaeonthophagus) akinini Koenig 1889
Onthophagus (Palaeonthophagus) albarracinus Baraud 1979
Onthophagus (Palaeonthophagus) aleppensis Redtenbacher 1843
Onthophagus (Palaeonthophagus) amirus Kabakov 1982
Onthophagus (Palaeonthophagus) anatolicus Petrovitz 1962
Onthophagus (Palaeonthophagus) andalusicus Waltl 1835
Onthophagus (Palaeonthophagus) angorensis Petrovitz 1963
Onthophagus (Palaeonthophagus) arnoldii Kabakov 1982
Onthophagus (Palaeonthophagus) baraudi Nicolas 1964
Onthophagus (Palaeonthophagus) basipustulatus Heyden 1889
Onthophagus (Palaeonthophagus) bonsae Zunino 1976
Onthophagus (Palaeonthophagus) bytinskii Balthasar 1960
Onthophagus (Palaeonthophagus) carpanetoi Pittino 1982
Onthophagus (Palaeonthophagus) coenobita (Herbst 1783)
Onthophagus (Palaeonthophagus) conspersus Reitter 1892
Onthophagus (Palaeonthophagus) cruciatus Ménétriés 1832
Onthophagus (Palaeonthophagus) dacatrai Pittino 2004
Onthophagus (Palaeonthophagus) dellacasai Pittino & Mariani 1981
Onthophagus (Palaeonthophagus) dorsosignatus Orbigny 1898
Onthophagus (Palaeonthophagus) eulaminicornis Pittino 2006
Onthophagus (Palaeonthophagus) excisus Reiche & Saulcy 1856
Onthophagus (Palaeonthophagus) excubitor Ziani & Gudenzi 2006
Onthophagus (Palaeonthophagus) finschi Harold 1877
Onthophagus (Palaeonthophagus) fissicornis (Steven 1809)
Onthophagus (Palaeonthophagus) fissinasus Fairmaire 1895
Onthophagus (Palaeonthophagus) flagrans Reitter 1892
Onthophagus (Palaeonthophagus) formaneki Reitter 1897
Onthophagus (Palaeonthophagus) fortigibber Reitter 1909
Onthophagus (Palaeonthophagus) fracticornis (Preyssler 1790)
Onthophagus (Palaeonthophagus) furciceps Marseul 1869
Onthophagus (Palaeonthophagus) glasunowi Koshantschikov 1894
Onthophagus (Palaeonthophagus) gorodinskii Kabakov 2008
Onthophagus (Palaeonthophagus) grossepunctatus Reitter 1905
Onthophagus (Palaeonthophagus) hermonensis Baraud 1982
Onthophagus (Palaeonthophagus) hissariensis Kabakov 2006
Onthophagus (Palaeonthophagus) isikdagensis Pittino 2004
Onthophagus (Palaeonthophagus) jiupengensis Masumoto, Lan & Kiuchi 2017
Onthophagus (Palaeonthophagus) joannae Goljan 1953
Onthophagus (Palaeonthophagus) kindermanni Harold 1877
Onthophagus (Palaeonthophagus) kolenatii Reitter 1892
Onthophagus (Palaeonthophagus) koryoensis Kim 1985
Onthophagus (Palaeonthophagus) laticornis Gebler 1823
Onthophagus (Palaeonthophagus) lemur (Fabricius 1781)
Onthophagus (Palaeonthophagus) lemuroides Orbigny 1898
Onthophagus (Palaeonthophagus) leucostigma (Steven 1812)
Onthophagus (Palaeonthophagus) lucidus (Illiger 1800)
Onthophagus (Palaeonthophagus) massai Baraud 1975
Onthophagus (Palaeonthophagus) medius (Kugelann 1792)
Onthophagus (Palaeonthophagus) medvedevi Kabakov 1982
Onthophagus (Palaeonthophagus) merdarius Chevrolat 1865
Onthophagus (Palaeonthophagus) muelleri Novak 1921
Onthophagus (Palaeonthophagus) nebulosus Reiche 1864
Onthophagus (Palaeonthophagus) necessarius Reitter 1892
Onthophagus (Palaeonthophagus) nikolajevi Kabakov 2006
Onthophagus (Palaeonthophagus) nurestanicus Kabakov 1982
Onthophagus (Palaeonthophagus) ocellatopunctatus Waterhouse 1875
Onthophagus (Palaeonthophagus) olsoufieffi Boucomont 1924
Onthophagus (Palaeonthophagus) opacicollis Reitter 1892
Onthophagus (Palaeonthophagus) osellai Pittino 1982
Onthophagus (Palaeonthophagus) ovatus (Linnaeus 1767)
Onthophagus (Palaeonthophagus) panici Petrovitz 1964
Onthophagus (Palaeonthophagus) parmatus Reitter 1892
Onthophagus (Palaeonthophagus) petrovitzianus Pittino 2006
Onthophagus (Palaeonthophagus) pljushtchi Ivanova 2012
Onthophagus (Palaeonthophagus) ponticus Harold 1883
Onthophagus (Palaeonthophagus) pseudocaccobius Reitter 1889
Onthophagus (Palaeonthophagus) psychopompus Ziani & Garakhloo 2010
Onthophagus (Palaeonthophagus) pygargus Motschulsky 1845
Onthophagus (Palaeonthophagus) quadrinodus Reitter 1896
Onthophagus (Palaeonthophagus) rachelis Martín-Piera 1985
Onthophagus (Palaeonthophagus) rodentium Pittino 2004
Onthophagus (Palaeonthophagus) roessneri Ziani 2016
Onthophagus (Palaeonthophagus) ruficapillus Brullé 1832
Onthophagus (Palaeonthophagus) rufimanus Kabakov 1982
Onthophagus (Palaeonthophagus) sacharovskii Olsoufieff 1918
Onthophagus (Palaeonthophagus) samai Ziani 2011
Onthophagus (Palaeonthophagus) scabriusculus Harold 1873
Onthophagus (Palaeonthophagus) semicornis (Panzer 1798)
Onthophagus (Palaeonthophagus) sericatus Reitter 1892
Onthophagus (Palaeonthophagus) shapovalovi Gusakov 2012
Onthophagus (Palaeonthophagus) silus Balthasar 1960
Onthophagus (Palaeonthophagus) similis (Scriba 1790)
Onthophagus (Palaeonthophagus) speculifer Solsky 1876
Onthophagus (Palaeonthophagus) strabo Reitter 1892
Onthophagus (Palaeonthophagus) stylocerus Graëlls 1851
Onthophagus (Palaeonthophagus) suermelii Petrovitz 1963
Onthophagus (Palaeonthophagus) sutleinensis Splichal 1910
Onthophagus (Palaeonthophagus) suturellus Brullé 1832
Onthophagus (Palaeonthophagus) tesquorum Semenov & Medvedev 1927
Onthophagus (Palaeonthophagus) trigibber Reitter 1892
Onthophagus (Palaeonthophagus) trispinus Reitter 1892
Onthophagus (Palaeonthophagus) truchmenus Kolenati 1846
Onthophagus (Palaeonthophagus) vacca (Linnaeus 1767)
Onthophagus (Palaeonthophagus) verticicornis (Laicharting 1781)
Onthophagus (Palaeonthophagus) viridis Ménétriés 1832
Onthophagus (Palaeonthophagus) vitulus (Fabricius 1776)
Onthophagus (Palaeonthophagus) vlasovi Medvedev 1958
Onthophagus (Palaeonthophagus) zuninoi Martín-Piera 1985
Onthophagus (Palaeonthophagus) zuvandi Qaryagdy 1939
6 additional siblings truncated for brevity.
See the
resource file
for a full list.
Dynamic Hierarchy Version 2.2.3
Onthophagus (Palaeonthophagus) marginalis (Gebler 1817)
(this page)
Onthophagus (Palaeonthophagus) marginalis przewalski Kabakov 2006
Onthophagus (Amphionthophagus) melitaeus (Fabricius 1798)
Onthophagus (Colobonthophagus) piffli Petrovitz 1961
Onthophagus (Colobonthophagus) thai Kabakov 1994
Onthophagus (Colobonthophagus) triceratops Arrow 1913
Onthophagus (Furconthophagus) karenensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus (Hikidaeus) leknamnaous Masumoto & Ochi 2015
Onthophagus (Hikidaeus) sauteri Gillet 1924
Onthophagus (Indachorius) baenzigeri Masumoto, Ochi & Hanboonsong 2007
Onthophagus (Indachorius) heterorrhinus Lansberge 1885
Onthophagus (Indachorius) subsapaensis Kabakov 1994
Onthophagus (Matashia) fossor Arrow 1931
Onthophagus (Micronthophagus) collinsi Krikken & Huijbregts 1987
Onthophagus (Micronthophagus) liui Masumoto, Ochi & Lee 2014
Onthophagus (Micronthophagus) vanofwegeni Krikken & Huijbregts 2008
Onthophagus (Onthophagiellus) thanwaakhomus Masumoto 1992
Onthophagus (Palaeonthophagus) clitellifer Reitter 1894
Onthophagus (Palaeonthophagus) kolenatii Reitter 1892
Onthophagus (Palaeonthophagus) nikolajevi Kabakov 2006
Onthophagus (Palaeonthophagus) parmatus Reitter 1892
Onthophagus (Palaeonthophagus) pygargus Motschulsky 1845
Onthophagus (Palaeonthophagus) rachelis Martin-Piera 1985
Onthophagus (Palaeonthophagus) vacca (Linnaeus 1767)
Onthophagus (Paraphanaeomorphus) jeannelianus Paulian 1945
Onthophagus (Parentius) nigellus (Illiger 1803)
Onthophagus (Phanaeomorphus) gagatoides Kabakov 2006
Onthophagus (Sinonthophagus) rugulosus Harold 1886
Onthophagus (Trichonthophagus) mediofuscatus D'Orbigny 1908
Onthophagus (Trichonthophagus) triacanthus Castelnau 1840
Onthophagus abyssinicus Gillet 1925
Onthophagus annulopunctatus Krikken & Huijbregts 2017
Onthophagus atridorsis D'Orbigny 1902
Onthophagus atriglabrus Howden & Gill 1987
Onthophagus axillaris Boheman 1860
Onthophagus baiyericus Krikken & Huijbregts 2013
Onthophagus batesi Howden & Cartwright 1963
Onthophagus birugifer D'Orbigny 1908
Onthophagus bituberifrons Josso 2018
Onthophagus bocandei D'Orbigny 1904
Onthophagus carpophilus Pereira & Halffter 1961
Onthophagus centurio Lansberge 1885
Onthophagus ceylonicus Harold 1859
Onthophagus chryses Bates 1887
Onthophagus civettae Cambefort 1984
Onthophagus convexicollis Boheman 1858
Onthophagus cribratus Lansberge 1883
Onthophagus crinitus Harold 1869
Onthophagus cruciger MacLeay 1888
Onthophagus cuniculus MacLeay 1864
Onthophagus diversus Reiche 1847
Onthophagus ebenicolor D'Orbigny 1902
Onthophagus egenus Harold 1877
Onthophagus evae Balthasar 1944
Onthophagus excisiceps D'Orbigny 1902
Onthophagus foulliouxi Cambefort 1971
Onthophagus gabonensis Walter 1989
Onthophagus gibbus D'Orbigny 1913
Onthophagus grandivigilans Masumoto 1995
Onthophagus gurburra Storey & Weir 1990
Onthophagus hemipygus Frey 1964
Onthophagus hircus Billberg 1815
Onthophagus hulstaerti Boucomont 1932
Onthophagus imbutus Sharp 1875
Onthophagus impunctatus D'Orbigny 1904
Onthophagus impurus Harold 1868
Onthophagus inermicollis D'Orbigny 1908
Onthophagus juncticornis D'Orbigny 1908
Onthophagus koebelei Blackburn 1903
Onthophagus mexicanus Bates 1887
Onthophagus miles D'Orbigny 1905
Onthophagus mimikanus Balthasar 1969
Onthophagus mongana Storey & Weir 1990
Onthophagus naevuliger D'Orbigny 1908
Onthophagus nigriceps Raffray 1877
Onthophagus novaeirlandiae Balthasar 1969
Onthophagus orientalis Harold 1868
Onthophagus palamoui Biswas & Chatterjee 1986
Onthophagus paravinctus Tagliaferri & Moretto 2012
Onthophagus piceiceps D'Orbigny 1904
Onthophagus pseudoconvexicollis Bai & Yang 2016
Onthophagus puberulus D'Orbigny 1902
Onthophagus pugnax Harold 1868
Onthophagus pullus Roth 1851
Onthophagus pupillatus Kolbe 1886
Onthophagus pygidialis Lansberge 1883
Onthophagus sansibaricus Harold 1878
Onthophagus savanicola Cambefort 1984
Onthophagus sculptilis Gerstaecker 1871
Onthophagus solomonensis Krikken & Huijbregts 2012
Onthophagus spiculatus Boucomont 1914
Onthophagus sylvipapuanus Krikken & Huijbregts 2012
Onthophagus tamworthi Blackburn 1903
Onthophagus tatsienluensis Balthasar 1942
Onthophagus tessulatus Harold 1871
Onthophagus thainuaensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus tirapensis Biswas & Chatterjee 1985
Onthophagus toraut Krikken & Huijbregts 2011
Onthophagus tricariniger D'Orbigny 1902
Onthophagus tripolitanus Heyden 1890
Onthophagus tshuapae Balthasar 1964
Onthophagus viridivinosus Kohlmann & Solis 2001
2034 additional siblings truncated for brevity.
See the
resource file
for a full list.
EOL Dynamic Hierarchy 1.1
Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Onthophagus
Onthophagus (Palaeonthophagus) marginalis (Gebler 1817)
(this page)
Onthophagus (Palaeonthophagus) marginalis przewalski Kabakov 2006
Onthophagus (Amphionthophagus) melitaeus (Fabricius 1798)
Onthophagus (Colobonthophagus) piffli Petrovitz 1961
Onthophagus (Colobonthophagus) thai Kabakov 1994
Onthophagus (Colobonthophagus) triceratops Arrow 1913
Onthophagus (Furconthophagus) karenensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus (Hikidaeus) leknamnaous Masumoto & Ochi 2015
Onthophagus (Hikidaeus) sauteri Gillet 1924
Onthophagus (Indachorius) baenzigeri Masumoto, Ochi & Hanboonsong 2007
Onthophagus (Indachorius) heterorrhinus Lansberge 1885
Onthophagus (Indachorius) subsapaensis Kabakov 1994
Onthophagus (Matashia) fossor Arrow 1931
Onthophagus (Micronthophagus) collinsi Krikken & Huijbregts 1987
Onthophagus (Micronthophagus) liui Masumoto, Ochi & Lee 2014
Onthophagus (Micronthophagus) vanofwegeni Krikken & Huijbregts 2008
Onthophagus (Onthophagiellus) thanwaakhomus Masumoto 1992
Onthophagus (Palaeonthophagus) clitellifer Reitter 1894
Onthophagus (Palaeonthophagus) kolenatii Reitter 1892
Onthophagus (Palaeonthophagus) nikolajevi Kabakov 2006
Onthophagus (Palaeonthophagus) parmatus Reitter 1892
Onthophagus (Palaeonthophagus) pygargus Motschulsky 1845
Onthophagus (Palaeonthophagus) rachelis Martin-Piera 1985
Onthophagus (Palaeonthophagus) vacca (Linnaeus 1767)
Onthophagus (Paraphanaeomorphus) jeannelianus Paulian 1945
Onthophagus (Parentius) nigellus (Illiger 1803)
Onthophagus (Phanaeomorphus) gagatoides Kabakov 2006
Onthophagus (Sinonthophagus) rugulosus Harold 1886
Onthophagus (Trichonthophagus) mediofuscatus D' Orbigny 1908
Onthophagus (Trichonthophagus) triacanthus Castelnau 1840
Onthophagus abyssinicus Gillet 1925
Onthophagus annulopunctatus Krikken & Huijbregts 2017
Onthophagus atridorsis D' Orbigny 1902
Onthophagus atriglabrus Howden & Gill 1987
Onthophagus axillaris Boheman 1860
Onthophagus baiyericus Krikken & Huijbregts 2013
Onthophagus batesi Howden & Cartwright 1963
Onthophagus birugifer D' Orbigny 1908
Onthophagus bituberifrons Josso 2018
Onthophagus bocandei D' Orbigny 1904
Onthophagus carpophilus Pereira & Halffter 1961
Onthophagus centurio Lansberge 1885
Onthophagus ceylonicus Harold 1859
Onthophagus chryses Bates 1887
Onthophagus civettae Cambefort 1984
Onthophagus convexicollis Boheman 1858
Onthophagus cribratus Lansberge 1883
Onthophagus crinitus Harold 1869
Onthophagus cruciger Macleay 1888
Onthophagus cuniculus Mac Leay 1864
Onthophagus diversus Reiche 1847
Onthophagus ebenicolor D' Orbigny 1902
Onthophagus egenus Harold 1877
Onthophagus evae Balthasar 1944
Onthophagus excisiceps D' Orbigny 1902
Onthophagus flavimargo D' Orbigny 1902
Onthophagus foulliouxi Cambefort 1971
Onthophagus gabonensis Walter 1989
Onthophagus gibbus D' Orbigny 1913
Onthophagus grandivigilans Masumoto 1995
Onthophagus gurburra Storey & Weir 1990
Onthophagus hemipygus Frey 1964
Onthophagus hircus Billberg 1815
Onthophagus hulstaerti Boucomont 1932
Onthophagus imbutus Sharp 1875
Onthophagus impunctatus D' Orbigny 1904
Onthophagus impurus Harold 1868
Onthophagus inermicollis D' Orbigny 1908
Onthophagus juncticornis D' Orbigny 1908
Onthophagus koebelei Blackburn 1903
Onthophagus mexicanus Bates 1887
Onthophagus miles D' Orbigny 1905
Onthophagus mimikanus Balthasar 1969
Onthophagus mongana Storey & Weir 1990
Onthophagus naevuliger D' Orbigny 1908
Onthophagus nigriceps Raffray 1877
Onthophagus novaeirlandiae Balthasar 1969
Onthophagus orientalis Harold 1868
Onthophagus palamoui Biswas & Chatterjee 1986
Onthophagus paravinctus Tagliaferri & Moretto 2012
Onthophagus pseudoconvexicollis Bai & Yang 2016
Onthophagus puberulus D' Orbigny 1902
Onthophagus pugnax Harold 1868
Onthophagus pullus Roth 1851
Onthophagus pupillatus Kolbe 1886
Onthophagus pygidialis Lansberge 1883
Onthophagus sansibaricus Harold 1878
Onthophagus savanicola Cambefort 1984
Onthophagus sculptilis Gerstaecker 1871
Onthophagus solomonensis Krikken & Huijbregts 2012
Onthophagus spiculatus Boucomont 1914
Onthophagus tamworthi Blackburn 1903
Onthophagus tatsienluensis Balthasar 1942
Onthophagus tessulatus Harold 1871
Onthophagus thainuaensis Masumoto, Ochi & Hanboonsong 2008
Onthophagus tirapensis Biswas & Chatterjee 1985
Onthophagus toraut Krikken & Huijbregts 2011
Onthophagus tricariniger D' Orbigny 1902
Onthophagus tripolitanus Heyden 1890
Onthophagus tshuapae Balthasar 1964
Onthophagus undaticeps D' Orbigny 1902
Onthophagus viridivinosus Kohlmann & Solis 2001
2071 additional siblings truncated for brevity.
See the
resource file
for a full list.
GBIF classification
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Onthophagus Latreille 1802
Onthophagus marginalis (Gebler 1817)
(this page)
Onthophagus marginalis andalusicus Waltl 1835
Onthophagus marginalis marginalis
Onthophagus marginalis przewalski Kabakov 2006
Onthophagus akhaus Masumoto, Ochi & Hanboonsong 2007
Onthophagus amicus (Gillet 1925)
Onthophagus anthracinus Harold 1873
Onthophagus antoinei Walter 1990
Onthophagus apunneea Masumoto, Ochi & Hanboonsong 2007
Onthophagus aztecus Zunino & Halffter 1988
Onthophagus babaulti D' Orbigny 1915
Onthophagus babirussa (Eschscholtz 1822)
Onthophagus bambra Matthews 1972
Onthophagus basilewskyi Frey 1961
Onthophagus bassariscus Zunino & Halffter 1988
Onthophagus batesi Howden & Cartwright 1963
Onthophagus bayeri Balthasar 1942
Onthophagus bennigseni Paulian 1937
Onthophagus bidentatus Drapiez 1819
Onthophagus bilingula Balthasar 1965
Onthophagus bituberifrons Josso 2018
Onthophagus campestris Masumoto, Kiuchi & Ho 2018
Onthophagus canescens Zunino & Halffter 1988
Onthophagus chilapensis Gasca-Álvarez, Zunino & Deloya 2018
Onthophagus curvicornis Latreille 1812
Onthophagus curvifrons D' Orbigny 1906
Onthophagus curvispina Reitter 1892
Onthophagus cyaneiceps D' Orbigny 1913
Onthophagus cyanellus Bates 1887
Onthophagus cyaneoniger D' Orbigny 1902
Onthophagus cyanochlorus D' Orbigny 1902
Onthophagus dorsofasciatus Fairmaire 1893
Onthophagus flaviclava D' Orbigny 1902
Onthophagus formosanus Gillet
Onthophagus fukuyamai Ochi, Kon & Ueda 2018
Onthophagus gagates Hope 1831
Onthophagus genuinus Kohlmann & Solis 2001
Onthophagus gibbicollis Lansberge 1885
Onthophagus ginyunensis Všetečka 1942
Onthophagus glasunowi Koshantschikov 1894
Onthophagus graniceps D' Orbigny 1908
Onthophagus granosus D' Orbigny 1913
Onthophagus granulatus Boheman 1858
Onthophagus granulifer Harold 1886
Onthophagus granulifrons Frey 1961
Onthophagus granulum D' Orbigny 1904
Onthophagus granum Lansberge 1885
Onthophagus graphicus Wallengren 1881
Onthophagus grataehelenae Kohlmann & Solis 2001
Onthophagus hingstoni Arrow 1931
Onthophagus hiroyukii Ochi 2007
Onthophagus inediapterus Kohlmann & Solis 2001
Onthophagus iumienus Masumoto, Ochi & Hanboonsong 2007
Onthophagus japonicus Harold 1875
Onthophagus kachinicus Kabakov 2006
Onthophagus kawaharai Ochi & Kon 2007
Onthophagus khonkaenus Masumoto, Ochi & Hanboonsong 2008
Onthophagus kivuensis Walter 2018
Onthophagus kuraruanus Matsumura 1938
Onthophagus lenzii Harold 1875
Onthophagus limonensis Kohlmann & Solis 2001
Onthophagus longipilis D' Orbigny 1905
Onthophagus luctuosus Boucomont 1914
Onthophagus ludio Boucomont 1914
Onthophagus lugens Fahraeus 1857
Onthophagus lugubris Fahraeus 1857
Onthophagus lunulifer Boucomont 1914
Onthophagus marginipennis Josso 2018
Onthophagus millingeni D' Orbigny 1898
Onthophagus monforti Josso & Prévost 2006
Onthophagus mycetorum Zunino & Halffter 1988
Onthophagus nemorivagus Kohlmann & Solis 2001
Onthophagus nubilus Kohlmann & Solis 2001
Onthophagus obenbergeri Balthasar 1965
Onthophagus oberthueri d'Orbigny 1898
Onthophagus obliquus (Olivier 1789)
Onthophagus papulatorius Kabakov 2008
Onthophagus parachandrai Ochi, Kon & Tsubaki 2009
Onthophagus pexatus Harold 1869
Onthophagus philippinensis Boucomont 1919
Onthophagus phoenicocerus Lea 1923
Onthophagus picatus D' Orbigny 1902
Onthophagus piceiceps D' Orbigny 1904
Onthophagus picipennis Hope 1841
Onthophagus pictus Reitter 1892
Onthophagus pseudoarai Ochi & Kon 2018
Onthophagus rojkoffianus Josso 2018
Onthophagus schoolmeestersi Ochi & Kon 2007
Onthophagus seniculus (Fabricius 1781)
Onthophagus shimba Cambefort 1980
Onthophagus shinichii Ochi & Kon 2018
Onthophagus simplex Raffray 1877
Onthophagus simpliciceps D' Orbigny 1905
Onthophagus simplicifrons Reitter 1892
Onthophagus simulator D' Orbigny 1905
Onthophagus sinuosicollis D' Orbigny 1905
Onthophagus sinuosus D' Orbigny 1913
Onthophagus sisyphoides Krikken 1977
Onthophagus skelleyi Sanchez-Huerta, Zunino & Halffter 2018
Onthophagus tigrinus D' Orbigny 1908
Onthophagus tungkamangensis Masumoto, Ochi & Hanboonsong 2007
Onthophagus ulula Balthasar 1966
Onthophagus zimmermanni Balthasar 1959
Onthophagus zumpti Frey 1954
2113 additional siblings truncated for brevity.
See the
resource file
for a full list.
NCBI
root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Scarabaeinae
Scarabaeinae incertae sedis
Onthophagus
Onthophagus marginalis
(this page)
Onthophagus alcyon
Onthophagus andalusicus
Onthophagus angustatus
Onthophagus anthracinus
Onthophagus apterus
Onthophagus atripennis
Onthophagus babirussa
Onthophagus babirussoides
Onthophagus baolocensis
Onthophagus batesi
Onthophagus bicallosus
Onthophagus bidentatus
Onthophagus bivertex
Onthophagus bonasus
Onthophagus borneensis
Onthophagus capella
Onthophagus carbonarius
Onthophagus cervicapra
Onthophagus cervus
Onthophagus championi
Onthophagus clypeatus
Onthophagus coenobita
Onthophagus consentaneus
Onthophagus crassicollis
Onthophagus cuniculus
Onthophagus curvicornis
Onthophagus dama
Onthophagus damaki
Onthophagus diabolicus
Onthophagus dunningi
Onthophagus falculatus
Onthophagus fimetarius
Onthophagus fodiens
Onthophagus furcaticeps
Onthophagus furcatus
Onthophagus gibbulus
Onthophagus giraffa
Onthophagus glabratus
Onthophagus gracilipes
Onthophagus grossepunctatus
Onthophagus haematopus
Onthophagus hirtus
Onthophagus illyricus
Onthophagus incensus
Onthophagus interstitialis
Onthophagus japonicus
Onthophagus joannae
Onthophagus laminatus
Onthophagus laticollis
Onthophagus latigena
Onthophagus lemur
Onthophagus lenzi
Onthophagus longimanus
Onthophagus maki
Onthophagus maniti
Onthophagus medius
Onthophagus melitaeus
Onthophagus merdarius
Onthophagus mije
Onthophagus minutus
Onthophagus mulgravei
Onthophagus muticus
Onthophagus nebulosus
Onthophagus nitidior
Onthophagus nutans
Onthophagus obscurior
Onthophagus ochreatus
Onthophagus ochromerus
Onthophagus ochropygus
Onthophagus opacicollis
Onthophagus orientalis
Onthophagus ovatus
Onthophagus pedator
Onthophagus phanaeides
Onthophagus piyawati
Onthophagus pullus
Onthophagus punctatus
Onthophagus quadripustulatus
Onthophagus rhinolophus
Onthophagus rorarius
Onthophagus rubicundulus
Onthophagus ruficapillus
Onthophagus rugicollis
Onthophagus schwaneri
Onthophagus semiareus
Onthophagus similis
Onthophagus sinicus
Onthophagus solivagus
Onthophagus stylocerus
Onthophagus taurinus
Onthophagus taurus
Onthophagus tragus
Onthophagus tricornis
Onthophagus vacca
Onthophagus variolaris
Onthophagus verticicornis
Onthophagus viduus
Onthophagus vinctus
Onthophagus vulpes
Onthophagus yukae
66 additional siblings truncated for brevity.
See the
resource file
for a full list.
wikidata_hierarchy tar gz
Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Scarabaeinae
Onthophagini
Onthophagus
Onthophagus marginalis
(this page)
Onthophagus marginalis andalusicus
Onthophagus marginalis marginalis
Onthophagus angularis
Onthophagus asper
Onthophagus aspericollis
Onthophagus asperipennis
Onthophagus aspernatus
Onthophagus asperrimus
Onthophagus asperulus
Onthophagus astigma
Onthophagus athiensis
Onthophagus atricolor
Onthophagus atridorsis
Onthophagus atroaereus
Onthophagus atrofasciatus
Onthophagus atronitidus
Onthophagus atrosericeus
Onthophagus aureofuscus
Onthophagus aureopilosus
Onthophagus auriculatus
Onthophagus aurifex
Onthophagus australis
Onthophagus axillaris
Onthophagus aztecus
Onthophagus comperei
Onthophagus compositus
Onthophagus costatus
Onthophagus costiger
Onthophagus creber
Onthophagus cribellum
Onthophagus cribratus
Onthophagus cribripennis
Onthophagus curvispina
Onthophagus cyaneiceps
Onthophagus cyanellus
Onthophagus cyaneoniger
Onthophagus cyanochlorus
Onthophagus cyclographus
Onthophagus cynomysi
Onthophagus cyobioides
Onthophagus dama
Onthophagus dandalu
Onthophagus dapcauensis
Onthophagus dapitanensis
Onthophagus decedens
Onthophagus digitifer
Onthophagus discrepans
Onthophagus endroedii
Onthophagus epilamprus
Onthophagus frontalis
Onthophagus gaillardi
Onthophagus gajo
Onthophagus galeatus
Onthophagus ganalensis
Onthophagus gandju
Onthophagus gangeticus
Onthophagus gangulu
Onthophagus gazellinus
Onthophagus geelongensis
Onthophagus gemellatus
Onthophagus gemmatus
Onthophagus gestroi
Onthophagus gibber
Onthophagus gibberosus
Onthophagus gibbidorsis
Onthophagus gibbus
Onthophagus gidju
Onthophagus giraffa
Onthophagus glabratus
Onthophagus glasunovi
Onthophagus gonopygus
Onthophagus gracilipes
Onthophagus humboldti
Onthophagus illyricus
Onthophagus indosinicus
Onthophagus indutus
Onthophagus inelegans
Onthophagus ineptus
Onthophagus inermiceps
Onthophagus inermicollis
Onthophagus inflaticollis
Onthophagus infucatus
Onthophagus inops
Onthophagus laborans
Onthophagus mocquerysi
Onthophagus modestus
Onthophagus noctis
Onthophagus pilauco
Onthophagus pluton
Onthophagus quadripustulatus
Onthophagus queenslandicus
Onthophagus quinquedens
Onthophagus quinquetuberculatus
Onthophagus ragazzii
Onthophagus rotundicollis
Onthophagus schultzei
Onthophagus sideki
Onthophagus simplex
Onthophagus simpliciceps
Onthophagus simulator
Onthophagus sinuosicollis
Onthophagus sinuosus
2369 additional siblings truncated for brevity.
See the
resource file
for a full list.
