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Life Expectancy

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The average lifespan of coppery brushtails is currently unknown. Closely related silver-gray brushtail possums, Trichosurus vulpecula, have an average lifespan of 7 years in the wild. The oldest known silver-gray brushtail lived 15.9 years in captivity.

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Associations

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Coppery brushtails may act as seed dispersers, as they consume some fruits.

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Benefits

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Coppery brushtails have small home ranges and rarely come into contact with humans. Because they are similar in appearance to other brushtails, they may be accidentally harvested for their fur.

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Benefits

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Negative effects of coppery brushtails on humans have not been recorded. A similar species Trichosurus vulpecula, however, damages crops, gardens, and plantations and are considered a pest species.

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Conservation Status

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Because Trichosurus johnstonii was only recently declared a separate species from Trichosurus vulpecula, little is know about its distribution. Trichosurus vulpecula is considered a species of least concern by the IUCN. Although they occur in some protected areas, T. vulpecula is considered a pest species to plantations and are frequently removed from agricultural areas. Changes in fire regime negatively affect these species.

US Federal List: no special status

CITES: no special status

State of Michigan List: no special status

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Behavior

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Coppery brushtails are nocturnal and have large eyes. A similar species, Trichosurus vulpecula, makes a variety of vocalizations, including alarm calls, screeches, hisses, grunts, clicks, and guttural coughs.

Communication Channels: acoustic

Perception Channels: visual ; tactile ; acoustic ; chemical

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Trophic Strategy

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The diet of Trichosurus johnstonii consists mainly of leaves and other plant material. They primarily feed upon the fruit of Solanum mauritianum, the leaves of Ipomoea, and the leaves of Cape Lilac (Melia azedarach).

Plant Foods: leaves; fruit; flowers

Primary Diet: herbivore (Folivore , Frugivore )

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Distribution

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Coppery brushtail, Trichosurus johnstonii, are found in Koombooloomba and Kuranda in northeastern Queensland, Australia. Little information is available regarding this species, as it was once considered a subspecies of silver-gray brushtail possums (common brushtails, Trichosurus vulpecula).

Biogeographic Regions: australian (Native )

Other Geographic Terms: island endemic

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Habitat

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Coppery brushtails mainly inhabit rainforest edges and tall open forests in the Atherton rainforests.

Habitat Regions: tropical

Terrestrial Biomes: forest ; rainforest

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Morphology

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Coppery brushtail possums get their name from the coppery, reddish colored fur that covers the majority of their medium sized body. The underside is covered in a creamy, lighter colored fur than the rest of the body. The head and body size of females ranges from 400 to 470 mm, while males tend to be slightly larger, reaching lengths of 490 mm. Tail length ranges from 300 to 380 mm in females and averages 400 mm in males. The hind feet of females and males measure 53.1 to 55.7 mm and about 59.3 mm respectively. Coppery brushtails have relatively large ears from 46.3 to 49.6 mm in length. Females weigh from 1200 to 1800 g, and males average 1800 g.

Scientists use differences in skull shape to distinguish this species from silver-gray brushtail possums.

Range mass: 1200 to 1800 g.

Range length: 400 to 490 mm.

Sexual Dimorphism: male larger

Other Physical Features: endothermic ; bilateral symmetry

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Meyer, T. 2012. "Trichosurus johnstonii" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Trichosurus_johnstonii.html
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Associations

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Known predators of Trichosurus johnstonii include barking owls, powerful owls, red foxs, cats, and domestic dogs. Coppery brushtails evade terrestrial predators by fleeing up into trees.

Known Predators:

  • cat (Felis catus)
  • red fox (Vulpes vulpes)
  • domestic dog (Canis lupis familiaris)
  • barking owl (Ninox connivens)
  • powerful owl (Ninox strenua)
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Reproduction

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Little is known regarding the mating systems of coppery brushtails.

Little data is currently available regarding the reproduction of coppery brushtails. Their reproductive habits are expected to be similar to those of silver-gray brushtail possums (Trichosurus vulpecula). Trichosurus vulpecula typically breeds twice a year, only giving birth once. Female silver-gray brushtails reach sexual maturity at 24 to 36 months of age, and males at around 48 months.

Female silver-gray brushtail possums have an estrous cycle of about 25 days. Gestation lasts an average of 17.5 days, after which a single young is born. After 4 to 5 months, the young leaves the pouch but continues to remain with the mother. At 6 to 7 months, the young is weaned, and it leaves the mother at 8 to 18 years of age.

Breeding interval: Coppery brushtails likely breed twice a year.

Key Reproductive Features: gonochoric/gonochoristic/dioecious (sexes separate); sexual

Parental investment of coppery brushtails is likely similar to that of silver-gray brushtail possums. Female silver-gray brushtails carry their young in their pouch for 4 to 5 months. Young are weaned at around 6 or 7 months, but they remain with their mother until about 8 to 18 years of age. Mothers likely provide protection or nourishment for a portion of this time.

Parental Investment: female parental care ; pre-hatching/birth (Provisioning: Female, Protecting: Female); pre-weaning/fledging (Provisioning: Female, Protecting: Female); post-independence association with parents

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Pòssum de Johnston ( Catalan; Valencian )

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El pòssum de Johnston (Trichosurus johnstonii) és una espècie de marsupial de la família dels falangèrids.[1] És originari d'Austràlia, on viu al nord-est de Queensland.

Referències

  1. Fitxa de l'animal al web de Mammal Species of the World. (anglès)


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Pòssum de Johnston: Brief Summary ( Catalan; Valencian )

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El pòssum de Johnston (Trichosurus johnstonii) és una espècie de marsupial de la família dels falangèrids. És originari d'Austràlia, on viu al nord-est de Queensland.

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Queensland-Kusu ( German )

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Der Queensland-Kusu (Trichosurus johnstonii) ist ein australisches Beuteltier, das im nördlichen Queensland an der westlichen Seite der Atherton Tablelands vom Koombooloomba-Nationalpark bis Kuranda endemisch vorkommt.

Merkmale

Die Tiere erreichen eine Kopfrumpflänge von 35,5 bis 43,8 cm, haben einen 33 bis 42 cm langen Schwanz und erreichen ein Gewicht von 1,2 bis 2 kg. Im Unterschied zum Fuchskusu, der im gleichen Gebiet vorkommt, zeigen sie keinen ausgeprägten Geschlechtsdimorphismus, lediglich der Schädel der Männchen ist ein klein wenig größer. Vom grauen Fuchskusu kann der Queensland-Kusu durch sein orangebraunes bis rötlichbraunes Fell leicht unterschieden werden. Sein buschiger Schwanz ist schwärzlich und proportional länger, die Ohren sind kleiner und die Backenzähne sind größer als beim Fuchskusu.[1]

Lebensraum und Lebensweise

Der Queensland-Kusu kommt vor allem an den Rändern der Regenwälder und den dort anschließenden hohen, offenen Wäldern vor. Bei einer Untersuchung aus einem Regenwald wurde eine Populationsdichte von drei bis vier Individuen pro Hektar festgestellt. Die Tiere sind nachtaktiv und verbringen den Tag in Baumhöhlen. Sie ernähren sich bevorzugt von Blättern, andere Pflanzenteile machen aber etwa ein Drittel ihrer Ernährung aus, vor allem die grünen Früchte verschiedener Nachtschattengewächse (Solanum). Insgesamt wurden 33 Pflanzenarten festgestellt, die zur Ernährung des Queensland-Kusu beitragen. Die meisten Jungtiere werden im Juni geboren, wenn auch der Fuchskusu seine Jungen zur Welt bringt. Beutegreifer, die dem Queensland-Kusu nachstellen, sind vor allem Schlangen, Eulen und Greifvögel.[1]

Gefährdung

Die IUCN listet den Queensland-Kusu bisher nicht, da er früher als Unterart des Fuchskusu angesehen wurde. Wahrscheinlich hat der Queensland-Kusu in der Vergangenheit durch Rodungen einen großen Teil seines Lebensraumes verloren. Der heutige Bestand gilt jedoch als stabil.[1]

Belege

  1. a b c Kristofer Helgen & Stephen Jackson: Family Phalangeridae (Cuscuses, Brush-tailed Possums and Scaly-tailed Possum). In: Don E. Wilson, Russell A. Mittermeier: Handbook of the Mammals of the World – Volume 5. Monotremes and Marsupials. Lynx Editions, 2015, ISBN 978-84-96553-99-6, S. 483
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Queensland-Kusu: Brief Summary ( German )

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Der Queensland-Kusu (Trichosurus johnstonii) ist ein australisches Beuteltier, das im nördlichen Queensland an der westlichen Seite der Atherton Tablelands vom Koombooloomba-Nationalpark bis Kuranda endemisch vorkommt.

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Coppery brushtail possum

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Coppery Brush-tailed Possum
This is a sub-species of the Brush-tailed Possum and is only found in the Atherton Tablelands area of Australia.

The coppery brushtail possum (Trichosurus johnstonii) is a species of marsupial possum in the family Phalangeridae.[2] Coppery brushtails are found within the Atherton Tablelands area of Queensland, in northeastern Australia.[1][2] These mammals inhabit rainforest ecosystems, living within the tree canopy. Though they have a restricted distribution, they are locally common.[2] This population is often considered a subspecies of T. vulpecula.[1]

Description

Coppery brushtail possums have a typical length of 400-490 mmm and weigh 1200-1800 g, with males being larger and heavier than females.[3][4]

Ecology

Like the common brushtail possum, coppery brushtails are nocturnal, and live in dens, which are usually tree hollows. At night, they still spend half of their time resting to conserve energy, and the other half in foraging.[5] In feeding experiments, in selecting their food, these possums may tend to select a mix of plant materials with detoxification requirements that are correlated or independent, rather than contradictory, thus maximizing their ability to process harmful plant byproducts.[6]

Dominance among individuals tends to place females above males, and larger over smaller individuals.[7]

References

  1. ^ a b c Morris, K.; Woinarski, J.; Friend, T.; Foulkes, J.; Kerle, A. & Ellis, M. (2008). "Trichosurus vulpecula". IUCN Red List of Threatened Species. 2008. Retrieved 11 July 2012.old-form url
  2. ^ a b c Groves, C. P. (2005). "Order Diprotodontia". In Wilson, D. E.; Reeder, D. M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 50. ISBN 978-0-8018-8221-0. OCLC 62265494.
  3. ^ Kerr, Sarah Emily (2011). Divergence of a mammal along a habitat gradient: a study of the coppery brushtail possum, Trichosurus vulpecula johnsonii (PhD Thesis). James Cook University.
  4. ^ Meyer, T. (2012). "Trichosurus johnstonii". Animal Diversity Web. Retrieved 29 January 2015.
  5. ^ Vandenbeld, John (1998). Nature of Australia: a portrait of the island continent. London: BBC Books.
  6. ^ Marsh, Karen J; Wallis, Ian R; McLean, Stuart; Sorenson, Jennifer S; Foley, William J (2006). "Conflicting Demands on Detoxification Pathways Influence How Common Brushtail Possums Choose Their Diets". Ecology. 87 (8): 2103–2112. doi:10.1890/0012-9658(2006)87[2103:cdodpi]2.0.co;2. JSTOR 20069195. PMID 16937649.
  7. ^ Blackie, Helen M; Russel, James C; Clout, Mick N (2011). "Maternal influence on philopatry and space use by juvenile brushtail possums (Trichosurus vulpecula)". Journal of Animal Ecology. 80 (2): 477–483. doi:10.1111/j.1365-2656.2010.01781.x. PMID 21155769.
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Coppery brushtail possum: Brief Summary

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Coppery Brush-tailed Possum This is a sub-species of the Brush-tailed Possum and is only found in the Atherton Tablelands area of Australia.

The coppery brushtail possum (Trichosurus johnstonii) is a species of marsupial possum in the family Phalangeridae. Coppery brushtails are found within the Atherton Tablelands area of Queensland, in northeastern Australia. These mammals inhabit rainforest ecosystems, living within the tree canopy. Though they have a restricted distribution, they are locally common. This population is often considered a subspecies of T. vulpecula.

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Trichosurus johnstonii ( Basque )

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Trichosurus johnstonii Trichosurus generoko animalia da. Martsupialen barruko Diprotodontia ordeneko animalia da. Phalangerinae azpifamilia eta Phalangeridae familian sailkatuta dago.

Erreferentziak

  1. Ramsay (1888) 8 Proc. Linn. Soc. N.S.W. 1297. or..

Ikus, gainera

(RLQ=window.RLQ||[]).push(function(){mw.log.warn("Gadget "ErrefAurrebista" was not loaded. Please migrate it to use ResourceLoader. See u003Chttps://eu.wikipedia.org/wiki/Berezi:Gadgetaku003E.");});
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Trichosurus johnstonii: Brief Summary ( Basque )

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Trichosurus johnstonii Trichosurus generoko animalia da. Martsupialen barruko Diprotodontia ordeneko animalia da. Phalangerinae azpifamilia eta Phalangeridae familian sailkatuta dago.

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Trichosurus johnstonii ( French )

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Trichosurus johnstonii est une espèce de marsupiaux de la famille des Phalangeridae.

Distribution

Cette espèce est endémique d'Australie, elle se rencontre sur le plateau d'Atherton au Queensland.

Publication originale

  • Ramsay, 1888 : Notes on the fauna of the Bellenden-Ker Ranges. Proceedings of the Linnaean Society of New South Wales, ser. 2, vol. 3, p. 1295–1299.

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Trichosurus johnstonii: Brief Summary ( French )

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Trichosurus johnstonii est une espèce de marsupiaux de la famille des Phalangeridae.

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Queenslandvoskoesoe ( Dutch; Flemish )

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De Queenslandvoskoesoe (Trichosurus johnstonii) is een klimbuideldier uit het geslacht der koesoes (Trichosurus).

Kenmerken

De Queenslandvoskoesoe heeft een korte vacht. Het lichaam is grotendeels lichtoranje van kleur. De staart is zwart.

Verspreiding

Deze soort komt voor in de regenwouden van Noordoost-Queensland.

Verwantschap

Deze soort is nauw verwant aan de voskoesoe (Trichosurus vulpecula) en wordt daar vaak toe gerekend als een ondersoort.

Literatuur

  • Groves, C.P. 2005. Order Diprotodontia. Pp. 43-70 in Wilson, D.E. & Reeder, D.M. (eds.). Mammal Species of the World: A taxonomic and geographic reference. 3rd ed. Baltimore: The Johns Hopkins University Press, Vol. 1: pp. i-xxxv+1-743; Vol. 2: pp. i-xvii+745-2142. ISBN 0 8018 8221 4
  • Menkhorst, P. & Knight, F. 2001. A Field Guide to the Mammals of Australia. South Melbourne: Oxford University Press, x+269 pp. ISBN 0 19 550870 X
Bronnen, noten en/of referenties
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Queenslandvoskoesoe: Brief Summary ( Dutch; Flemish )

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De Queenslandvoskoesoe (Trichosurus johnstonii) is een klimbuideldier uit het geslacht der koesoes (Trichosurus).

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Trichosurus johnstonii ( Portuguese )

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Trichosurus johnstonii é uma espécie de marsupial da família Phalangeridae. Endêmica da Austrália.

Referências

  • GROVES, C. P. Order Diprotodontia. In: WILSON, D. E.; REEDER, D. M. (Eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. 3. ed Baltimore: Johns Hopkins University Press, 2005. v. 1, p. 43-70.
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Trichosurus johnstonii: Brief Summary ( Portuguese )

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Trichosurus johnstonii é uma espécie de marsupial da família Phalangeridae. Endêmica da Austrália.

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Trichosurus johnstonii ( Swedish )

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Trichosurus johnstonii[1] är en pungdjursart som först beskrevs av Ramsay 1888. Trichosurus johnstonii ingår i släktet pungrävar och familjen klätterpungdjur.[2][3] Inga underarter finns listade.[2]

Pungdjuret förekommer i nordöstra Queensland, Australien.[1] Djuret betraktas av IUCN som population av vanlig pungräv (Trichosurus vulpecula) och listas därför inte extra.[4]

Källor

  1. ^ [a b] Wilson, Don E., and DeeAnn M. Reeder, eds. (2005) , Mammal Species of the World: A Taxonomic and Geographic Reference, 3rd ed., vols. 1 & 2, Trichosurus johnstonii
  2. ^ [a b] Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (red.) (28 april 2011). ”Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist.”. Species 2000: Reading, UK. Arkiverad från originalet den 18 juni 2012. https://web.archive.org/web/20120618223324/http://www.catalogueoflife.org/services/res/2011AC_26July.zip. Läst 24 september 2012.
  3. ^ ITIS: The Integrated Taxonomic Information System. Orrell T. (custodian), 2011-04-26
  4. ^ Morris, K., Woinarski, J., Friend, T., Foulkes, J., Kerle, A. & Ellis, M. 2008 Trichosurus vulpecula Från: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.2 <www.iucnredlist.org>. Läst 2013-03-26.

Externa länkar

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Trichosurus johnstonii: Brief Summary ( Swedish )

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Trichosurus johnstonii är en pungdjursart som först beskrevs av Ramsay 1888. Trichosurus johnstonii ingår i släktet pungrävar och familjen klätterpungdjur. Inga underarter finns listade.

Pungdjuret förekommer i nordöstra Queensland, Australien. Djuret betraktas av IUCN som population av vanlig pungräv (Trichosurus vulpecula) och listas därför inte extra.

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Trichosurus johnstonii ( Vietnamese )

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Trichosurus johnstonii là một loài động vật có vú trong họ Phalangeridae, bộ Hai răng cửa. Loài này được Ramsay mô tả năm 1888.[2]

Chú thích

  1. ^ Morris, K., Woinarski, J., Friend, T., Foulkes, J., Kerle, A. & Ellis, M. (2008). Trichosurus vulpecula. Sách Đỏ IUCN các loài bị đe dọa. Phiên bản 2012.1. Liên minh Bảo tồn Thiên nhiên Quốc tế. Truy cập ngày 11 tháng 7 năm 2012.
  2. ^ a ă Wilson, D. E.; Reeder, D. M. biên tập (2005). “Trichosurus johnstonii”. Mammal Species of the World . Baltimore: Nhà in Đại học Johns Hopkins, 2 tập (2.142 trang). ISBN 978-0-8018-8221-0. OCLC 62265494.

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Trichosurus johnstonii: Brief Summary ( Vietnamese )

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Trichosurus johnstonii là một loài động vật có vú trong họ Phalangeridae, bộ Hai răng cửa. Loài này được Ramsay mô tả năm 1888.

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Медный кузу ( Russian )

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Царство: Животные
Подцарство: Эуметазои
Без ранга: Вторичноротые
Подтип: Позвоночные
Инфратип: Челюстноротые
Надкласс: Четвероногие
Подкласс: Звери
Инфракласс: Сумчатые
Подотряд: Phalangeriformes
Надсемейство: Phalangeroidea
Семейство: Кускусовые
Род: Кузу
Вид: Медный кузу
Международное научное название

Trichosurus johnstonii Ramsay, 1888

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ITIS 709436NCBI 596272EOL 311986

Медный кузу[источник не указан 340 дней] (лат. Trichosurus johnstonii) — сумчатое млекопитающие семейства кускусовых.

Описание

Длина тела составляет от 40 до 49 см. Вес взрослой особи — от 1,2 до 1,8 кг. Самцы крупнее и тяжелее самок.[1][2] Мех серо-рыжий или полностью рыжий. Медный кузу часто считается подвидом лисьего кузу.

Распространение

Обитает во влажных тропических лесах Квинсленда, в северо-восточной Австралии. Хотя в целом вид имеет ограниченное географическое распространение, на данной территории встречается часто.

Образ жизни

Как и другие кузу, ведет ночной образ жизни, в дневное время укрывается в дуплах деревьев. Но в отличие от других кузу, медный кузу и ночью часть времени тратит на сон и отдых.[3] Питается преимущественно листьями, фруктами, семенами, беспозвоночными и мелкими позвоночными.

Примечания

  1. Kerr, Sarah Emily (2011). Divergence of a mammal along a habitat gradient: a study of the coppery brushtail possum, Trichosurus vulpecula johnsonii (PhD Thesis). James Cook University.
  2. Meyer, T. Trichosurus johnstonii (неопр.). Animal Diversity Web (2012). Проверено 29 января 2015.
  3. Vandenbeld, John. Nature of Australia: a portrait of the island continent. — London : BBC Books, 1998.
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Медный кузу: Brief Summary ( Russian )

provided by wikipedia русскую Википедию

Медный кузу[источник не указан 340 дней] (лат. Trichosurus johnstonii) — сумчатое млекопитающие семейства кускусовых.

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구릿빛붓꼬리주머니쥐 ( Korean )

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구릿빛붓꼬리주머니쥐(Trichosurus johnstonii)는 쿠스쿠스과에 속하는 유대류의 일종이다.[3] 오스트레일리아 북동부 퀸즐랜드주 애서톤 테이블랜드 지역에서 발견된다.[3][2] 우림 생태계에서 서식하며 나무 캐노피에서 생활한다. 제한된 지역에 분포하지만 현지에서는 흔하게 발견된다.[3] 한때는 주머니여우(T. vulpecula)의 아종으로 간주되기도 하였다.[2]

특징

몸길이는 평균 400~490mm, 몸무게는 1200~1800g 정도이고 수컷이 암컷보다 크고 무겁다.[4][5]

생태

주머니여우처럼 구릿빛붓꼬리주머니쥐는 야행성 동물이며, 보통 나무 구멍 속에 만든 굴 속에서 산다. 밤에 에너지를 비축하기 위하여 휴식 시간의 절반을 쓰고, 나머지 반은 먹이를 구하는 데 쓴다.[6]

각주

  1. Groves, C.P. (2005). Wilson, D.E.; Reeder, D.M., 편집. 《Mammal Species of the World: A Taxonomic and Geographic Reference》 (영어) 3판. 존스 홉킨스 대학교 출판사. 49쪽. ISBN 0-801-88221-4. OCLC 62265494.
  2. Morris, K., Woinarski, J., Friend, T., Foulkes, J., Kerle, A. & Ellis, M. (2008). Trichosurus vulpecula. 《멸종 위기 종의 IUCN 적색 목록. 2012.1판》 (영어). 국제 자연 보전 연맹. 2012년 7월 11일에 확인함.
  3. Groves, C.P. (2005). 〈Order Diprotodontia〉 [캥거루목]. Wilson, D.E.; Reeder, D.M. 《Mammal Species of the World: A Taxonomic and Geographic Reference》 (영어) 3판. 존스 홉킨스 대학교 출판사. 50쪽. ISBN 978-0-8018-8221-0. OCLC 62265494.
  4. Kerr, Sarah Emily (2011). 《Divergence of a mammal along a habitat gradient: a study of the coppery brushtail possum, Trichosurus vulpecula johnsonii》 (PhD Thesis). James Cook University.
  5. Meyer, T. (2012). “Trichosurus johnstonii”. Animal Diversity Web. 2015년 1월 29일에 확인함.
  6. Vandenbeld, John (1998). 《Nature of Australia: a portrait of the island continent》. London: BBC Books.
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