Pseudopogonatherum quadrinerve (Hack.) Ohwi

Eulalia quadrinervis is very close to E. villosa (Sprengel) Nees (E. wightii (J. D. Hooker) Bor). The latter species is distributed from Africa to S India and differs in its acute leaf blades and leathery, broader lower glume, which is concave down the whole midline and usually hairy only on the flanks. A few specimens from Yunnan with acute leaf blades are better placed in E. quadrinervis.

Culm stout erect. Ligule about 1 mm long, chartaceous, hispid on the back. inflorescence of digitately arranged racemes. Spikelets paired monomorphic the upper pedicellate, the lower sessile, about 6 mm long. Lower glume subcoriaceous broadly lanceolate, as long as the spikelet, margins inrolled, upper part 2-keeled keels ciliate, nerves anastomosing; upper glume 3-nerved, of the same texture as the lower glume, margins fimbriate; lower lemma nearly of the same texture and length as the upper glume; upper lemma about 3 mm long, 2-cleft, fimbriate, aristate, awn flexuous and hispid; upper palea hyaline, about 2 mm long, margins inrolled, fimbriate.

Perennial, tufted from a short scaly rhizome; basal sheaths glabrous, rarely thinly appressed pilose between veins. Culms 0.6–1.2 m tall, 2–4 mm in diam., pilose below inflorescence, nodes glabrous. Leaf sheaths densely villous to hirsute with tubercle-based hairs, or subglabrous; leaf blades linear, 10–40 × 0.4–0.6 cm, glabrous to tuberculate-villous, abaxial surface glaucous, apex acuminate-filiform; ligule 1–1.5 mm. Racemes 2–7, subdigitate, 10–18 cm, hairs white or pale mauve; rachis internodes and pedicels 1/2–2/3 spikelet length, silky villous. Sessile spikelet 5–6.5 mm, golden brown; callus hairs 1/3 spikelet length, white; lower glume narrowly elliptic, back slightly concave and villous below middle, flat and glabrous above, 2–4 green veins between keels, connected by veinlets below apex and between lateral veins, upper keels shortly pectinate-ciliate, apex membranous, subacute; upper lemma ovate-oblong, 2-lobed in upper 1/3; awn 1.2–2 cm. Anthers 2.7–3.4 mm. Fl. and fr. Sep–Nov.

Himalaya, Assam (Khasia & Naga Hills), Manipur, Burma, Indo-China.

Distributed in Himalaya, India, Burma, Thailand, China, Japan and extending to the Philippines.

1500-2700 m

Dry mountain slopes, grassy places. Anhui, Fujian, Guangdong, Henan, Sichuan, Taiwan, Yunnan, Zhejiang [Bhutan, N India, Japan, Korea, Myanmar, Nepal, Philippines, Thailand, Vietnam].

Pollinia quadrinervis Hack., Andropog. 158 & in Bull. Herb. Boiss. 7: 640. 721. 1899.

Pollinia quadrinervis Hackel in A. Candolle & C. Candolle, Monogr. Phan. 6: 158. 1889; Eulalia quadrinervis var. latigluma B. S. Sun & S. Wang; P. villosa var. chefuensis Franchet; Pseudopogonatherum quadrinerve (Hackel) Ohwi.