Ademosynidae

Ademosynidae is an extinct family of beetles, known from the Late Triassic to Early Cretaceous. There are at least six genera and 11 species in Ademosynidae.^[1][2][3][4] Members of the family were small oval beetles, with a length generally smaller than 1 cm. Characteristics of the family include a pronotum without anterior angles and a rounded anterior margin, and elytra with 9–12 punctate striae. The systematic position of the family within the order Coleoptera is currently uncertain: they can be considered early examples of the suborder Polyphaga, or unusual examples of Archostemata.^[3]

Genera

The family Ademosynidae is restricted to the following six genera and 11 species in Yan, Beutel and Ponomarenko (2017):^[3]

• † Ademosyne Handlirsch, 1906
  • †Ademosyne bacca Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
  • †Ademosyne elliptica Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
  • †Ademosyne kirghizica Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
  • †Ademosyne major Handlirsch, 1906 – Blackstone Formation, Australia, Late Triassic (Norian)
• † Cephalosyne Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
  • †Cephalosyne capitata Ponomarenko, 1969
• † Dolichosyne Ponomarenko, 1969 – Madygen Formation, Kyrgyzstan, Late Triassic (Carnian)
  • †Dolichosyne confragosa Ponomarenko, 1969
  • †Dolichosyne rostrata Ponomarenko, 1969
  • †Dolichosyne sulcata Ponomarenko, 1969
• † Gnathosyne Ponomarenko, 1969 – Tologoi Formation, Kazakhstan, Late Triassic (Norian)
  • †Gnathosyne akkolkensis Ponomarenko, 1969
• † Petrosyne Ponomarenko, 1969 – Kyrgyzstan, Early Jurassic (Pliensbachian)
  • †Petrosyne liassica Ponomarenko, 1969
• † Sphaerosyne Ponomarenko, 1969 – Zaza Formation, Russia, Early Cretaceous (Aptian)
  • †Sphaerosyne globosa Ponomarenko, 1969

The following genera and species are also included in the family according to Fossilworks, but are excluded or not mentioned in Yan, Beutel and Ponomarenko (2017):^[2]

• † Atalosyne Ren, 1995 – Lushangfen Formation, China, Early Cretaceous (Aptian)
  • †Atalosyne sinuolata Ren, 1995
• † Chaocoleus Ponomarenko, Yan & Huang, 2014 – Yinping Formation, China, Permian (Capitanian)
  • †Chaocoleus limnebius Ponomarenko, Yan & Huang, 2014
• † Eremisyne Wang, 1998 – Xiazhuang Formation, China, Early Cretaceous (Aptian/Albian)
  • †Eremisyne xiazhuangensis Wang, 1998
• † Grammositus Dunstan, 1923 (also spelled Grammositum) – Blackstone Formation, Australia, Late Triassic (Norian)
  • †Grammositus bilineatus Dunstan, 1923
• †Ademosyne adunca Dunstan, 1923
• †Ademosyne arcucciae Martins-Neto and Gallego, 2006
• †Ademosyne australiensis Tillyard, 1916
• †Ademosyne brevis Dunstan, 1923
• †Ademosyne cameroni Tillyard, 1916
• †Ademosyne congener Tillyard, 1916
• †Ademosyne curvata Dunstan, 1923
• †Ademosyne elongatus Martins-Neto and Gallego, 2006
• †Ademosyne hexacostata Martins-Neto and Gallego, 2006
• †Ademosyne intermedia Dunstan, 1923
• †Ademosyne lata Dunstan, 1923
• †Ademosyne llantenesensis Brauckmann et al., 2010
• †Ademosyne olliffi (Handlirsch, 1906)
• †Ademosyne parva Dunstan, 1923
• †Ademosyne polyzetete Ponomarenko, 2011
• †Ademosyne prisca Riek, 1974
• †Ademosyne punctata Tillyard, 1916
• †Ademosyne punctuada Martins-Neto and Gallego, 2006
• †Ademosyne ramocostata Dunstan, 1923
• †Ademosyne reducta Riek, 1976
• †Ademosyne rosenfeldi Brauckmann et al., 2010
• †Ademosyne rugulosa Dunstan, 1923
• †Ademosyne sibirica Rohdendorf, 1961
• †Ademosyne speciosa Riek, 1974
• †Ademosyne umutu Martins-Neto and Gallego, 2009
• †Ademosyne vittamargina Dunstan, 1923
• †Ademosyne wianamattensis Tillyard, 1918

References