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Image of Tomarus bituberculatus (Palisot de Beauvois 1805)
Creatures » … » Animal » … » Arthropods » … » Hexapods » Insects » Winged Insects » … » Endopterygotes » Beetles » … » Dynastidae

Carrot Beetles

Tomarus

Scientific Names

Preferred Names

Tomarus
Genus recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from Catalogue of Life in Species 2000 & ITIS Catalogue of Life
Tomarus
Genus recognized by wikipedia NO, EOL Dynamic Hierarchy 1.1, EOL Dynamic Hierarchy 1.1, Bolivia Species List, Brazil Species List, Canada Species List, Chile Species List, Colombia Species List, Costa Rica Species List, Dominican Republic Species List, El Salvador Species List, Guatemala Species List, Nicaragua Species List, Panama Species List, Paraguay Species List, The Bahamas Species List, Argentina Species List, Mexico Species List, North Atlantic Species List, North Pacific Species List, NCBI, United States Species List, wikidata_hierarchy tar gz, wikipedia EN, Wikidata Common Names, Wikimedia Commons, Barcode of Life Data Systems, Global Biotic Interactions, Global Biotic Interactions, iNaturalist data coverage, Wikipedia (inferred records), Dynamic Hierarchy Version 2.2.3, GGBN data coverage, and EOL Dynamic Hierarchy Version 2.2.4
Tomarus
Recognized by IABIN
Tomarus Erichson 1847
Genus recognized by GBIF classification, GBIF national node type records France, Catalogue of Life 26 March 2024, Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy, and GBIF data coverage

Alternative Names

Bothynus selanderi Cartwright
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Bothynus Hope, 1837)
Chalepus nasutus Burmeister 1847
Homotypic synonym according to GBIF classification (Originally found in sources as synonym of Oxyligyrus nasutus (Burmeister, 1847))
Ligyrus compositus Wickham 1911
Homotypic synonym according to GBIF classification (Originally found in sources as doubtful taxon within Tomarus Erichson, 1847)
Oxyligyrus nasutus (Burmeister 1847)
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Oxyligyrus Arrow, 1908)
Tomarus laticaudus López-García & Deloya 2019
Doubtful according to GBIF classification
Tomarus maracaiboensis López-García & Deloya 2019
Doubtful according to GBIF classification
Tomarus moroni López-García & Deloya 2019
Doubtful according to GBIF classification
Tomarus paranaensis López-García & Deloya 2019
Doubtful according to GBIF classification
Tomarus pilcopataensis López-García & Deloya 2019
Doubtful according to GBIF classification
? cuniculus Fabricius, 1801
Homotypic synonym according to GBIF classification (Originally found in sources as doubtful taxon within Dynastidae)
Anagryllus Casey 1915
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Ligyrellus Casey 1915
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Ligyrodes Casey 1915
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Ligyrus Burmeister 1847
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Ligyrus subgen. Grylius Casey 1915
Synonym according to Catalogue of Life 26 March 2024
Tomarus
Harvest ancestor according to GBIF national node type records France, NMNH type specimens, iDigBio type specimen records, iNaturalist, Harvard Museum of Comparative Zoology, Harvard Museum of Comparative Zoology, TreatmentBank, Global Biotic Interactions, and Global Biotic Interactions

Common Names

English

Carrot Beetles preferred
Recognized by Wikidata Common Names

Persian

توماروس preferred
Recognized by Wikidata Common Names

Curated hierarchies for Tomarus

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Dynastidae
Tomarus (this page)
Tomarus adoceteus Ratcliffe & Cave 2010
Tomarus bidentulus (Fairmaire 1892)
Tomarus bituberculatus (Palisot de Beauvois 1805)
Tomarus burmeisteri (Steinheil 1872)
Tomarus cicatricosus (Prell 1937)
Tomarus columbianus López-García & Gasca-Álvarez 2014
Tomarus cuniculus (Fabricius 1801)
Tomarus discrepans (Escalona & Joly 2006)
Tomarus ebenus (Degeer 1774)
Tomarus fossor (Latreille 1833)
Tomarus gibbosus (Degeer 1774)
Tomarus gyas Erichson 1848
Tomarus laevicollis (Bates 1888)
Tomarus maimon Erichson 1847
Tomarus maternus (Prell 1937)
Tomarus nasutus (Burmeister 1847)
Tomarus neglectus (Le Conte 1847)
Tomarus peruvianus (Endrödi 1970)
Tomarus pullus (Prell 1937)
Tomarus pumilus (Prell 1937)
Tomarus relictus (Say 1825)
Tomarus roigjunenti Neita & Ratcliffe 2017
Tomarus rostratus Dupuis 2014
Tomarus rubripes (Boheman 1858)
Tomarus sallaei (Bates 1888)
Tomarus selanderi (Cartwright 1959)
Tomarus similis (Endrödi 1968)
Tomarus spinipenis Neita & Ratcliffe 2017
Tomarus subtropicus (Blatchley 1922)
Tomarus villosus (Burmeister 1847)
Aceratus
Adoryphorus
Aegopsis
Agaocephala
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
116 additional siblings truncated for brevity.
See the resource file for a full list.

Catalogue of Life 26 March 2024

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Dynastinae MacLeay 1819
Pentodontini Mulsant 1842
Pentodontina Mulsant 1842
Tomarus Erichson 1847 (this page)
Tomarus adoceteus (Ratcliffe & Cave 2010)
Tomarus amazonicus (Arrow 1914)
Tomarus bituberculatus (Palisot de Beauvois 1811)
Tomarus colombianus López-García & Gasca-Álvarez 2014
Tomarus compositus (Wickham 1911)
Tomarus discrepans (Escalona & Joly 2006)
Tomarus effetus (Wickham 1914)
Tomarus gyas Erichson 1849
Tomarus laevicollis (Bates 1888)
Tomarus laticaudus López-García & Deloya 2019
Tomarus maimon Erichson 1847
Tomarus maracaiboensis López-García & Deloya 2019
Tomarus maternus (Prell 1937)
Tomarus pilcopataensis López-García & Deloya 2019
Tomarus pullus (Prell 1937)
Tomarus pumilus (Prell 1937)
Tomarus roigjunenti Neita & Ratcliffe 2017
Tomarus rostratus Dupuis 2014
Tomarus selanderi (Cartwright 1959)
Tomarus subtropicus (Blatchley 1922)
Aceratus Prell 1936
Adelaeus Ratcliffe 2014
Alissonotum Arrow 1908
Ampotis Dechambre & Madge 1979
Anoplognatho Rivers 1889
Anoronotum Arrow 1939
Aphonides Rivers 1889
Aphonodelus Kolbe 1905
Aphonus Le Conte 1856
Barutus Ratcliffe 1981
Bothynus Hope 1837
Calcitoryctes Krell 2011
Calicnemis Laporte 1832
Callistemonus Péringuey 1901
Carneiola Endrödi 1974
Carneodon Özdikmen 2009
Collagenus Ratcliffe & Hardy 2005
Coptognathus Burmeister 1847
Coscinocephalus Prell 1936
Dalgopus Dechambre 2001
Denhezia Dechambre 2006
Diloboderus Reiche 1859
Ebolowanius Dechambre 1979
Enarotadius Endrödi 1971
Eremobothynus Ohaus 1910
Eucopidocaulus Prell 1912
Euetheola Bates 1888
Euligyrus Casey 1915
Eutyctus Semenov 1890
Gillaspytes Howden 1980
Gorditus Ratcliffe 2010
Haplosoma Semenov 1890
Heteroconus Kolbe 1900
Heteroglobus Dupuis & Dechambre 2008
Heteroligus Kolbe 1900
Heteronychus Dejean 1833
Heterostriatus Dupuis 2006
Hiekeianus Endrödi 1978
Homoeomorphus Burmeister 1847
Hyboschema Péringuey 1901
Hylobothynus Ohaus 1910
Hyphoryctes Blackburn 1895
Ligyrus Burmeister 1847
Lonchotus Burmeister 1847
Marronus Coquerel 1866
Mellissius Wollaston 1869
Metanastes Arrow 1911
Microryctes Arrow 1908
Musurgus Vauloger 1898
Neocorynophyllus Carne 1985
Neometanastes Yamaya 2010
Neonastes Carne 1957
Neoryctes Arrow 1908
Nimbacola Paulian 1970
Orizabus Fairmaire 1878
Orsilochus Burmeister 1847
Oxyligyrus Arrow 1908
Papuana Arrow 1911
Paranodon Cockerell 1905
Pareteronychus Prell 1934
Parisomorphus Schaufuss 1890
Pentodina Endrödi 1968
Pentodon Hope 1837
Pentodontoschema Péringuey 1901
Pericoptus Burmeister 1847
Philoscaptus Brèthes 1919
Phyllognathus Eschscholtz 1830
Pimelopus Erichson 1842
Piscoperus Ratcliffe & Giraldo 2014
Podalgus Burmeister 1847
Prionoryctes Arrow 1911
Proculigyrus López-García & Deloya 2022
Pseudohomonyx Arrow 1908
Pycnoschema Thomson 1859
Saccharoscaptus Morón & Grossi 2015
Siralus Dechambre 1998
Temnorhynchus Hope 1837
Thronistes Burmeister 1847
Tribialus Dupuis & Perrin 2022
Wernoryctes Dechambre 1999

Dynamic Hierarchy Version 2.2.3

Tomarus (this page)
Tomarus adoceteus Ratcliffe & Cave 2010
Tomarus bidentulus (Fairmaire 1892)
Tomarus bituberculatus (Palisot de Beauvois 1805)
Tomarus burmeisteri (Steinheil 1872)
Tomarus cicatricosus (Prell 1937)
Tomarus columbianus López-García & Gasca-Álvarez 2014
Tomarus cuniculus (Fabricius 1801)
Tomarus discrepans (Escalona & Joly 2006)
Tomarus ebenus (Degeer 1774)
Tomarus fossor (Latreille 1833)
Tomarus gibbosus (Degeer 1774)
Tomarus gyas Erichson 1848
Tomarus laevicollis (Bates 1888)
Tomarus maimon Erichson 1847
Tomarus maternus (Prell 1937)
Tomarus nasutus (Burmeister 1847)
Tomarus neglectus (Le Conte 1847)
Tomarus peruvianus (Endrödi 1970)
Tomarus pullus (Prell 1937)
Tomarus pumilus (Prell 1937)
Tomarus relictus (Say 1825)
Tomarus roigjunenti Neita & Ratcliffe 2017
Tomarus rostratus Dupuis 2014
Tomarus rubripes (Boheman 1858)
Tomarus sallaei (Bates 1888)
Tomarus selanderi (Cartwright 1959)
Tomarus similis (Endrödi 1968)
Tomarus subtropicus (Blatchley 1922)
Tomarus villosus (Burmeister 1847)
Aceratus
Adoryphorus
Aegopsis
Agaocephala
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
116 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Dynastidae
Tomarus (this page)
Tomarus adoceteus Ratcliffe & Cave 2010
Tomarus bidentulus (Fairmaire 1892)
Tomarus bituberculatus (Palisot de Beauvois 1805)
Tomarus burmeisteri (Steinheil 1872)
Tomarus cicatricosus (Prell 1937)
Tomarus columbianus López-García & Gasca-Álvarez 2014
Tomarus compositus (Wickham 1911)
Tomarus cuniculus (Fabricius 1801)
Tomarus discrepans (Escalona & Joly 2006)
Tomarus ebenus (Degeer 1774)
Tomarus effetus (Wickham 1914)
Tomarus fossor (Latreille 1833)
Tomarus gibbosus (Degeer 1774)
Tomarus gyas Erichson 1848
Tomarus laevicollis (Bates 1888)
Tomarus maimon Erichson 1847
Tomarus maternus (Prell 1937)
Tomarus nasutus (Burmeister 1847)
Tomarus neglectus (Le Conte 1847)
Tomarus peruvianus (Endrödi 1970)
Tomarus pullus (Prell 1937)
Tomarus pumilus (Prell 1937)
Tomarus relictus (Say 1825)
Tomarus roigjunenti Neita & Ratcliffe 2017
Tomarus rostratus Dupuis 2014
Tomarus rubripes (Boheman 1858)
Tomarus sallaei (Bates 1888)
Tomarus selanderi (Cartwright 1959)
Tomarus similis (Endrödi 1968)
Tomarus spinipenis Neita & Ratcliffe 2017
Tomarus subtropicus (Blatchley 1922)
Tomarus villosus (Burmeister 1847)
Aceratus
Adoryphorus
Aegopsis
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calcitoryctes
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
118 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy Version 2.2.4

Tomarus (this page)
Tomarus adoceteus Ratcliffe & Cave 2010
Tomarus bidentulus (Fairmaire 1892)
Tomarus bituberculatus (Palisot de Beauvois 1805)
Tomarus burmeisteri (Steinheil 1872)
Tomarus cicatricosus (Prell 1937)
Tomarus columbianus López-García & Gasca-Álvarez 2014
Tomarus cuniculus (Fabricius 1801)
Tomarus discrepans (Escalona & Joly 2006)
Tomarus ebenus (Degeer 1774)
Tomarus fossor (Latreille 1833)
Tomarus gibbosus (Degeer 1774)
Tomarus gyas Erichson 1848
Tomarus laevicollis (Bates 1888)
Tomarus maimon Erichson 1847
Tomarus maternus (Prell 1937)
Tomarus nasutus (Burmeister 1847)
Tomarus neglectus (Le Conte 1847)
Tomarus peruvianus (Endrödi 1970)
Tomarus pullus (Prell 1937)
Tomarus pumilus (Prell 1937)
Tomarus relictus (Say 1825)
Tomarus roigjunenti Neita & Ratcliffe 2017
Tomarus rostratus Dupuis 2014
Tomarus rubripes (Boheman 1858)
Tomarus sallaei (Bates 1888)
Tomarus selanderi (Cartwright 1959)
Tomarus similis (Endrödi 1968)
Tomarus subtropicus (Blatchley 1922)
Tomarus villosus (Burmeister 1847)
Aceratus
Adoryphorus
Aegopsis
Agaocephala
Allomyrina
Ancognatha
Anomacaulus
Anomalomorpha
Aphonides
Aphonus
Archophanes
Arriguttia
Aspidolea
Augoderia
Augosoma
Barutus
Blabephorus
Bothynus
Calicnemis
Callistemonus
Caymania
Ceratoryctoderus
Chalcasthenes
Chalcocrates
Chalepides
Cheiroplatys
Clyster
Cnemidophileurus
Colacus
Corynophyllus
Cryptodus
Dalgopus
Dasygnathus
Denhezia
Ebolowanius
Endebius
Endroedianibe
Enracius
Erioscelis
Eucopidocaulus
Eutyctus
Gillaspytes
Goniophileurus
Gorditus
Haplophileurus
Haploscapanes
Hatamus
Hemicyrthus
Hemiphileurus
Heteroconus
Heteroglobus
Heterogomphus
Heterostriatus
Hispanioryctes
Horridocalia
Hyboschema
Hyphoryctes
Irazua
Licnostrategus
Lycomedes
Marronus
Megaceras
Megasoma
Mellissius
Metanastes
Microryctes
Mitracephala
Musurgus
Neocnecus
Neonastes
Neoryctes
Nimbacola
Orizabus
Orthocavonus
Oryctes
Oxyligyrus
Parisomorphus
Pentodina
Pentodon
Pericoptus
Phileucourtus
Phylliocephala
Pimelopus
Piscoperus
Planophileurus
Platyphileurus
Podalgus
Prionoryctes
Prosphileurus
Pseudocavonus
Pseudosyrichthus
Ruteloryctes
Saccharoscaptus
Semanopterus
Talautoclyster
Temnorhynchus
Thronistes
Trichogomphus
Trioplus
Trissodon
116 additional siblings truncated for brevity.
See the resource file for a full list.

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Dynastidae
Tomarus Erichson 1847 (this page)
Tomarus adoceteus Ratcliffe & Cave 2010
Tomarus bisignatus Horn 1895
Tomarus bituberculatus (Palisot de Beauvois 1805)
Tomarus burmeisteri (Steinheil 1872)
Tomarus chamaeropis Schaeffer
Tomarus cicatricosus (Prell 1937)
Tomarus colombianus Lopez-Garcia & Gasca-Alvarez 2014
Tomarus columbianus López-García & Gasca-Álvarez 2014
Tomarus compositus (Wickham 1911)
Tomarus cuniculus (Fabricius 1801)
Tomarus discrepans (Escalona & Joly 2006)
Tomarus ebenus (Degeer 1774)
Tomarus effetus (Wickham 1914)
Tomarus formosianus Grouvelle 1913
Tomarus fossor (Latreille 1833)
Tomarus gibbosus (Degeer 1774)
Tomarus gyas Erichson 1848
Tomarus hirtellus Schwarz
Tomarus laevicollis (Bates 1888)
Tomarus maimon Erichson 1847
Tomarus maternus (Prell 1937)
Tomarus nasutus (Burmeister 1847)
Tomarus neglectus (Le Conte 1847)
Tomarus obsoletus Casey
Tomarus pullus (Prell 1937)
Tomarus pumilus (Prell 1937)
Tomarus quadrinotatus Casey
Tomarus relictus (Say 1825)
Tomarus roigjunenti Neita & Ratcliffe 2017
Tomarus rostratus Dupuis 2014
Tomarus rubripes (Boheman 1858)
Tomarus sallaei (Bates 1888)
Tomarus selanderi (Cartwright 1959)
Tomarus similis (Endrödi 1968)
Tomarus spinipenis Neita & Ratcliffe 2017
Tomarus subtropicus (Blatchley 1922)
Tomarus ventricosus Casey
Tomarus villosus (Burmeister 1847)
Aceratus Prell 1936
Acrobolbia Ohaus 1912
Aegopsis Burmeister 1847
Agaocephala Le Peletier & Serville 1828
Allophileurinus Dupuis & Dechambre 2001
Amblyodus Westwood 1878
Amblyoproctus Kolbe 1910
Ancognatha Erichson 1847
Aphonodelus Kolbe 1905
Archophileurus Kolbe 1910
Aspidolea Bates 1888
Augosoma Burmeister 1841
Blabephorus Fairmaire 1898
Bothynus Hope 1837
Calicnemis Laporte de Castelnau 1832
Carneiola Endrödi 1974
Carneoryctes Özdikmen 2009
Cavonus Sharp 1875
Ceratophileurus Ohaus 1911
Ceratoryctoderus Arrow 1908
Chalcasthenes Arrow 1937
Chalcosoma Hope 1837
Cheiroplatys Kirby 1837
Chiliphileurus Endrodi 1977
Cnemidophileurus Kolbe 1910
Coenoryctoderus Prell 1933
Colacus Ohaus 1910
Collagenus Ratcliffe & Hardy 2005
Coptognathus Burmeister 1847
Coscinocephalus Prell 1936
Cryptodus Mac Leay 1819
Cyphonistes Burmeister 1847
Dalgopus Dechambre 2001
Dasygnathus Macleay 1819
Democrates Burmeister 1847
Denhezia Dechambre 2006
Dichodontus Burmeister 1847
Dyscinetus Harold 1869
Endroedianibe Chalumeau 1981
Enema Hope 1837
Eophileurus Arrow 1908
Epironastes Carne 1957
Erioscelis Burmeister 1847
Eupatorus Burmeister 1847
Eutyctus Semenov 1889
Gibboryctes Endrödi 1974
Golofa Hope 1837
Haplophileurus Kolbe 1910
Haplosoma Semenov 1889
Harposceles Burmeister 1847
Hatamus Sharp 1877
Hemicyrthus Reiche 1860
Hemiphileurus Kolbe 1910
Heteroglobus Dupuis & Dechambre 2008
Heterostriatus Dupuis 2006
Hexodon Olivier 1789
Homoeomorphus Burmeister 1847
Horridocalia Endrodi 1974
Hovophileurus Arrow 1911
Indieraligus Dechambre 1979
Irazua Ratcliffe 2003
Licnostrategus Prell 1933
Lonchotus Burmeister 1847
Lycomedes Brême 1845
Macrocyphonistes Ohaus 1910
Megaceras Hope 1837
Megasoma Kirby 1825
Mellissius Wollaston 1869
Microphileurus Kolbe 1910
Microryctes Arrow 1908
Musurgus Vauloger 1898
Neocorynophyllus Carne 1985
Neodasygnathus Carne 1957
Neohyphus Heller 1896
Novapus Sharp 1875
Onychionyx Arrow 1914
Oryctoderus Boisduval 1835
Oryctophileurus Kolbe 1878
Papuana Arrow 1911
Parapucaya Prell 1934
Parisomorphus Schaufuss 1890
Paroryctoderus Dechambre 1994
Pentodontoschema Péringuey 1901
Phileurus Latreille 1807
Philoscaptus Brèthes 1919
Platyphileurus Ohaus 1910
Prionoryctes Arrow 1911
Pseudocavonus Blackburn 1910
Pseudosyrichthus Péringuey 1901
Pucaya Ohaus 1910
Ruteloryctes Arrow 1908
Scapanes Burmeister 1847
Siralus Dechambre 1998
Syrichthodontus Péringuey 1901
Syrichthoschema Janssens 1942
Talautoclyster Yamaya 2001
Trichogomphus Burmeister 1847
Trioplus Burmeister 1847
Wernoryctes
Xyloryctes Hope 1837
116 additional siblings truncated for brevity.
See the resource file for a full list.

Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Coleoptera Linnaeus 1758
Polyphaga Emery 1886
Scarabeiformia Crowson 1960
Scarabaeoidea Latreille 1802
Scarabaeidae Latreille 1802
Dynastinae MacLeay 1819
Pentodontini Mulsant 1842
Tomarus Erichson 1847 (this page)
Tomarus cuniculus (Fabricius 1801)
Tomarus gibbosus (De Geer 1774)
Tomarus laevicollis (Bates 1888)
Tomarus nasutus (Burmeister 1847)
Tomarus neglectus (Le Conte 1847)
Tomarus relictus (Say 1825)
Tomarus sallaei (Bates 1888)
Tomarus selanderi (Cartwright 1959)
Tomarus subtropicus (Blatchley 1922)
Anoplognatho Rivers 1888
Aphonus Le Conte 1856
Coscinocephalus Prell 1936
Euetheola Bates 1888
Gillaspytes Howden 1980
Orizabus Fairmaire 1878
Oxygrylius Casey 1915

NCBI

root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Dynastinae
Tomarus (this page)
Tomarus gibbosus
Tomarus sallaei
Agaocephala
Alissonotum
Allomyrina
Amblyoproctus
Ancognatha
Aspidolea
Augosoma
Beckius
Bothynus
Calicnemis
Carneodon
Chalcosoma
Cheiroplatys
Coelosis
Cryptodus
Cyclocephala
Cyphonistes
Diloboderus
Dynastes
Dyscinetus
Enema
Eophileurus
Euetheola
Eupatorus
Eutyctus
Golofa
Haploscapanes
Hemiphileurus
Heterogomphus
Hexodon
Homophileurus
Lycomedes
Marronus
Megaceras
Megasoma
Metanastes
Mimeoma
Neocorynophyllus
Novapus
Orizabus
Oryctes
Oryctoderus
Pachyoryctes
Papuana
Parapucaya
Peltonotus
Pentodon
Phileurus
Phyllognathus
Pimelopus
Pucaya
Pycnoschema
Semanopterus
Spodistes
Stenocrates
Strategus
Syrictes
Temnorhynchus
Trichogomphus
Trissodon
Trypoxylus
unclassified Dynastinae
Xyloryctes
Xylotrupes

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Dynastinae
Pentodontini
Tomarus (this page)
Tomarus adoceteus
Tomarus bidentulus
Tomarus bituberculatus
Tomarus burmeisteri
Tomarus cicatricosus
Tomarus colombianus
Tomarus columbianus
Tomarus compositus
Tomarus cuniculus
Tomarus discrepans
Tomarus ebenus
Tomarus effetus
Tomarus fossor
Tomarus gianucai
Tomarus gibbosus
Tomarus gyas
Tomarus laevicollis
Tomarus maimon
Tomarus maternus
Tomarus nasutus
Tomarus neglectus
Tomarus peruvianus
Tomarus pullus
Tomarus pumilus
Tomarus relictus
Tomarus rosettae
Tomarus rostratus
Tomarus rubripes
Tomarus sallaei
Tomarus selanderi
Tomarus similis
Tomarus subtropicus
Tomarus villosus
Ampotis
Anaronotum
Anoplognatho
Australocus
Barutus
Callistemonus
Callistemonus intrusus
Cheiroplatina
Coptognatus
Dasygnathus
Dipelicina
Dipelicus
Eutheola
Heikeianus
Heteronychus
Heterostriatus
Hylobothynus
Idioschema karruense
Marronus
Microryctes
Neobothynus
Neoryctes
Nimbacola
Orizabus
Oxyligyrus
Papuana
Parabothynus
Paranodon
Pentodon
Pentodontina
Pericoptus
Piscoperus
Podalgus
Pseudocavonus
Pseudoryctina
Pycnoschema
Temnorrhynchus
Wernoryctes
Wernoryctes werneri
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