dcsimg
 
  en  
names in breadcrumbs
Image of horned dung beetle
Creatures » … » Animal » … » Arthropods » … » Hexapods » Insects » Winged Insects » … » Endopterygotes » Beetles » … » Scarab Beetles

Copris

Scientific Names

Preferred Names

Copris
Genus recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from Catalogue of Life in Species 2000 & ITIS Catalogue of Life
Copris
Genus recognized by wikipedia 日本語, wikipedia русскую Википедию, wikipedia NL, wikipedia POL, wikipedia VI, wikipedia NO, wikipedia SV, EOL Dynamic Hierarchy 1.1, EOL Dynamic Hierarchy 1.1, Austria Species List, Belize Species List, Botswana Species List, Burundi Species List, Cameroon Species List, Canada Species List, China Species List, Colombia Species List, Costa Rica Species List, Ecuador Species List, El Salvador Species List, France Species List, Germany Species List, Ghana Species List, Greece Species List, Guatemala Species List, Guernsey Species List, India Species List, Israel Species List, Italy Species List, Ivory Coast Species List, Kenya Species List, Lesotho Species List, Malawi Species List, Namibia Species List, Nicaragua Species List, Panama Species List, Poland Species List, Portugal Species List, Rwanda Species List, Slovakia Species List, South Africa Species List, Spain Species List, Sweden Species List, Switzerland Species List, Tanzania Species List, The Netherlands Species List, Turkey Species List, United Kingdom Species List, Zambia Species List, Zimbabwe Species List, wikipedia ID, Japan Species List, Mexico Species List, Gulf of Guinea Species List, Indian Ocean Species List, North Atlantic Species List, North Pacific Species List, South Atlantic Species List, wikipedia AST, wikipedia ET, NCBI, wikipedia emerging languages, wikipedia MIN, United States Species List, wikidata_hierarchy tar gz, Paleobiology Database, wikipedia DE, wikipedia ES, wikipedia IT, wikipedia PT, wikipedia FR, wikipedia EN, Wikidata Common Names, Wikipedia (inferred records), Wikimedia Commons, Barcode of Life Data Systems, Dynamic Hierarchy Version 2.2 - Test, Global Biotic Interactions, Global Biotic Interactions, and Global Biotic Interactions
Copris
Subgenus recognized by wikidata_hierarchy tar gz
Copris
Recognized by Animal Diversity Web, OBIS environmental data records, and Animal Diversity Web Traits and images
Copris Geoffroy 1762
Genus recognized by GBIF classification, Catalogue of Life 7 February 2023, and Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Copris subgen. Copris Geoffroy 1762
Subgenus recognized by Catalogue of Life 7 February 2023
Litocopris Waterhouse 1891
Genus recognized by GBIF classification

Alternative Names

Ateuchus klugi (Harold 1868)
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Ateuchus Weber, 1801)
Cleptocaccobius youngai (Balthasar 1967)
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Cleptocaccobius Cambefort, 1984)
Copris amyntor Harold 1869
Heterotypic synonym according to GBIF classification
Copris belisama Schrank 1798
Heterotypic synonym according to GBIF classification
Copris castaneus Mulsant 1842
Heterotypic synonym according to GBIF classification
Copris colonica Say 1835
Doubtful according to GBIF classification
Copris corniculatus Mulsant 1842
Heterotypic synonym according to GBIF classification
Copris deletus Mulsant 1842
Heterotypic synonym according to GBIF classification
Copris gistelianus Gistel 1857
Heterotypic synonym according to GBIF classification
Copris integer Harold 1869
Heterotypic synonym according to GBIF classification
Copris jenisonianus Gistel 1857
Heterotypic synonym according to GBIF classification
Copris obliteratus Mulsant 1842
Heterotypic synonym according to GBIF classification
Copris quadridentatus Fabricius 1798
Doubtful according to GBIF classification
Copris victorini Péringuey 1901
Heterotypic synonym according to GBIF classification
Dichotomius laeviceps (Harold 1869)
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Dichotomius Hope, 1838)
Neosisyphus gladiator (Arrow 1927)
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Neosisyphus Müller, 1942)
Oryctes silenus (Fabricius 1775)
Proparte synonym according to GBIF classification (Originally found in sources as synonym of Phyllognathus excavatus (Forster, 1771))
Paragymnopleurus sinuatus szechouanicus (Balthasar 1934)
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Paragymnopleurus sinuatus (Olivier, 1789))
Paraphytus ritsemae (Harold 1880)
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Paraphytus Harold, 1877)
Proagoderus gladiator (Arrow 1907)
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Proagoderus Lansberge, 1883)
Scarabaeus ammon Fabricius 1781
Heterotypic synonym according to GBIF classification
Scarabaeus bifidus Poda 1761
Heterotypic synonym according to GBIF classification
Scarabaeus emarginatus Olivier 1789
Heterotypic synonym according to GBIF classification
Scarabaeus hispanus Linnaeus 1764
Homotypic synonym according to GBIF classification
Scarabaeus lar Fabricius 1787
Heterotypic synonym according to GBIF classification
Scarabaeus lunaris Linnaeus 1758
Homotypic synonym according to GBIF classification (Originally found in sources as accepted taxon within Scarabaeus)
Scarabaeus lunus Schrank 1798
Heterotypic synonym according to GBIF classification
Scarabaeus marianus Petiver 1764
Heterotypic synonym according to GBIF classification
Scarabaeus minutus Drury 1773
Homotypic synonym according to GBIF classification (Originally found in sources as doubtful taxon within Scarabaeus)
Scarabaeus quadridentatus Degeer 1778
Heterotypic synonym according to GBIF classification
Scarabaeus silenus Fabricius 1775
Proparte synonym according to GBIF classification
Afrontherus Marchisio & Zunino 2012
Synonym according to Catalogue of Life 7 February 2023
Copris
Harvest ancestor according to GBIF national node type records France, Mozambique Species List, NMNH type specimens, Flickr Group, iNaturalist, Harvard Museum of Comparative Zoology, Harvard Museum of Comparative Zoology, Simpson et al 2020, TreatmentBank, and Global Biotic Interactions
Copris subgen. Insulicopris Tsukamoto 1974
Synonym according to Catalogue of Life 7 February 2023
Corpus Geoffroy 1762
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Litocopris Waterhouse 1891
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Microcopris Balthasar 1958
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Paracopris Balthasar 1939
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Common Names

Bulgarian

копри preferred
Recognized by Wikidata Common Names

Catalan; Valencian

escarabat banyut preferred
Recognized by Wikidata Common Names

Chinese

粪蜣螂属 preferred
Recognized by Wikidata Common Names

Czech

výkalník preferred
Recognized by Wikidata Common Names

Esperanto

Kopriso preferred
Recognized by Wikidata Common Names

Kazakh

Копрлар preferred
Recognized by Wikidata Common Names

Kirghiz; Kyrgyz

Жампачылдар preferred
Recognized by Wikidata Common Names

Navajo; Navaho

Chąąneiłmaasii bideeʼ hólóní preferred
Recognized by Wikidata Common Names

Russian

Копры preferred
Recognized by Wikidata Common Names

Zh Cn

粪蜣螂属 preferred
Recognized by Wikidata Common Names

Zh Tw

糞蜣螂屬 preferred
Recognized by Wikidata Common Names
show 8 more languages
hide 8 languages

Curated hierarchies for Copris

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Copris (this page)
Copris (Paracopris) andrewesi Waterhouse 1891
Copris (Paracopris) anomiopseoides Boucomont 1924
Copris (Paracopris) cariniceps Felsche 1910
Copris (Paracopris) compressipennis Gillet 1910
Copris (Paracopris) cribratus Gillet 1927
Copris (Paracopris) davisoni Waterhouse 1891
Copris (Paracopris) excisus Waterhouse 1891
Copris (Paracopris) furciceps Felsche 1910
Copris (Paracopris) imitans Felsche 1910
Copris (Paracopris) kasagii Ochi & Kon 1996
Copris (Paracopris) pedarioides Lansberge 1886
Copris (Paracopris) punctulatus Wiedemann 1823
Copris (Paracopris) punjabensis Gillet 1921
Copris (Paracopris) ramosiceps Gillet 1921
Copris (Paracopris) signatus Walker 1858
Copris (Paracopris) surdus Arrow 1931
Copris (Sinocopris) barclayi Ochi, Kon & Bai 2009
Copris (Sinocopris) basipunctatus Balthasar 1942
Copris (Sinocopris) cheni Ochi, Kon & Bai 2007
Copris (Sinocopris) kiuchii Masumoto 1989
Copris (Sinocopris) kusakabei Ochi, Kon & Kawahara 2005
Copris (Sinocopris) laevigatus Gillet 1927
Copris (Sinocopris) obenbergeri Balthasar 1933
Copris (Sinocopris) ochus (Motschulsky 1860)
Copris (Sinocopris) parapecuarius Ochi, Kon & Kawahara 2008
Copris (Sinocopris) pecuarius Lewis 1884
Copris (Sinocopris) pseudochus Ochi & Kon 2004
Copris (Sinocopris) quasilaevigatus Ochi, Kon & Bai 2007
Copris (Sinocopris) sabinus Gillet 1910
Copris (Sinocopris) sacontala Redtenbacher 1848
Copris (Sinocopris) tsukamotoi Ochi, Kon & Kawahara 2008
Copris (Sinocopris) uenoi Hanboonsong, Masumoto & Ochi 2003
Copris (Sinocopris) yangi Ochi, Kon & Bai 2007
Copris acutidens Motschulsky 1860
Copris afgoi Cambefort 1992
Copris agnus Sharp 1875
Copris algol Nguyen-phung 1989
Copris amabilis Kolbe 1914
Copris amazonicus Darling & Génier 2018
Copris amyntor Klug 1855
Copris anceus (Olivier 1789)
Copris angusticornis Arrow 1933
Copris angustus Nguyen-phung & Cambefort 1986
Copris antares Ferreira 1958
Copris arcturus Gillet 1907
Copris ares Zelenka 1993
Copris arimotoi Ochi & Araya 1992
Copris arizonensis Schaeffer 1906
Copris armatus Harold 1869
Copris armeniacus Faldermann 1835
Copris armiger Gillet 1910
Copris arrowi Felsche 1910
Copris aspericollis Gillet 1910
Copris atropolitus Gillet 1910
Copris basilewskyi Ferreira 1962
Copris bellator Chevrolat 1844
Copris bengalensis Gillet 1911
Copris bidens Kolbe 1893
Copris bootes Klug 1855
Copris boucardi Harold 1869
Copris bovinus Gillet 1908
Copris brachypterus Nomura 1964
Copris brevicornis Darling & Génier 2018
Copris brittoni Ferreira 1962
Copris caelatus (Fabricius 1794)
Copris caliginosus Kohlmann, Cano & Delgado 2003
Copris cambeforti Nguyen-phung 1988
Copris cambodiensis Ochi, Kon & Kawahara 2008
Copris camerunus Felsche 1904
Copris caobangensis Bui, Dumack & Bonkowski 2018
Copris capensis Waterhouse 1891
Copris carinicus Gillet 1910
Copris carmelita Fabricius 1801
Copris cassius Péringuey 1901
Copris celebensis Lansberge 1886
Copris chimalapensis Mora-Aguilar & Delgado 2015
Copris colmanti Gillet 1908
Copris complexus Nguyen-phung 1989
Copris confucius Harold 1877
Copris coriarius Gillet 1907
Copris cornifrons Boheman 1860
Copris corniger Sahlberg 1823
Copris corpulentus Gillet 1910
Copris costaricensis Gahan 1894
Copris crassus Deschodt & Davis 2015
Copris cribricollis Gillet 1910
Copris curlettii Moretto 2011
Copris davidi Darling & Génier 2018
Copris davisi Nguyen-phung & Cambefort 1986
Copris decellei Cambefort 1992
Copris delicatus Arrow 1931
Copris denticulatus Nguyen-phung 1988
Copris diversus Waterhouse 1891
Copris draco Arrow 1906
Copris dracunculus Ferreira 1959
Copris dudleyi Cambefort 1992
Copris eburneus Cambefort 1992
Copris elpheneroides Felsche 1910
Copris elphenor Klug 1855
Copris erratus Lansberge 1886
Copris evanidus Klug 1855
Copris fallaciosus Gillet 1907
Copris fallax Felsche 1910
Copris felschei Reitter 1892
Copris fidius (Olivier 1789)
Copris frankenbergerianus Balthasar 1958
Copris freyi Ferreira 1961
Copris fricator (Fabricius 1787)
Copris fukiensis Balthasar 1952
Copris garambae Cambefort 1992
Copris gazellarum Gillet 1918
Copris gibbulus Lansberge 1886
Copris gilleti Kolbe 1907
Copris gladiator Arrow 1933
Copris gopheri Hubbard 1894
Copris gracilis Waterhouse 1891
Copris graueri Kolbe 1914
Copris halffteri Matthews 1959
Copris harrisi Waterhouse 1891
Copris hartli Frey 1975
Copris hatayamai Ochi & Araya 1992
Copris hispanus (Linnaeus 1764)
Copris howdeni Matthews & Halffter 1959
Copris humilis Gillet 1908
Copris hybridus Gillet 1910
Copris igualensis Warner 1990
Copris illotus Balthasar 1942
Copris inaequabilis Zhang 1997
Copris incertus Say 1835
Copris inemarginatus Blatchley 1918
Copris inhalatus Quedenfeldt 1884
Copris insidiosus Péringuey 1901
Copris integer Reiche 1847
Copris interioris Kolbe 1897
Copris iris Sharp 1875
Copris jacchoides Nguyen-phung & Cambefort 1987
Copris jacchus (Fabricius 1775)
Copris jahi Nguyen-phung & Cambefort 1986
Copris johkii Ochi & Kon 2014
Copris juanmorai Mora-Aguilar & Delgado 2015
Copris jucundus Gillet 1932
Copris kachinensis Ochi, Kon & Kawahara 2008
Copris kasanka Josso & Prévost 2016
Copris katangae Cambefort & Nguyen-phung 1996
Copris keralensis Gill 1986
Copris klapperichi Balthasar 1942
Copris klugi Harold 1869
Copris laeviceps Harold 1869
Copris laiiformis Nguyen-phung & Cambefort 1987
Copris laioides Boucomont 1932
Copris laius Harold 1868
Copris lannathai Hanboonsong, Masumoto & Ochi 2003
Copris lecontei Matthews 1961
Copris leruei Josso & Prévost 2017
Copris lugubris Boheman 1858
Copris lunaris (Linnaeus 1758)
Copris macacus Lansberge 1886
Copris macclevei Warner 1990
Copris macer Péringuey 1901
Copris maesi Ratcliffe 1998
Copris magicus Harold 1881
Copris manni Ochi, Kon & Bai 2009
Copris marcus Gillet 1921
Copris martinae Nguyen-phung 1988
Copris martinezi Matthews & Halffter 1968
Copris matthewsi Delgado & Kohlmann 2001
Copris medogensis Zhang 1988
Copris megaceratoides Waterhouse 1891
Copris megasoma Matthews & Halffter 1959
Copris mesacanthus Harold 1878
Copris mexicanus Matthews & Halffter 1959
Copris minutus (Drury 1773)
Copris misellus Péringuey 1901
Copris moechus Le Conte 1854
Copris moffartsi Gillet 1907
Copris mongkhoni Hanboonsong, Masumoto & Ochi 2003
Copris montivagus Gillet 1908
Copris moroni Darling & Génier 2018
Copris morphaeus Gillet 1932
Copris mourgliai Zelenka 1993
Copris neglectus Moxey 1963
Copris neotridens Génier & Prévost 2017
Copris nepos Gillet 1908
Copris nevinsoni Waterhouse 1891
Copris novaki Zelenka 1992
Copris nubilosus Kohlmann, Cano & Delgado 2003
Copris numa Lansberge 1886
Copris obesus Boheman 1857
Copris orion Klug 1835
Copris orphanus Guérin-Méneville 1847
Copris paraszechouanicus Ochi, Kon & Pham 2018
Copris pauliani Nguyen-phung & Cambefort 1986
Copris phungae Cambefort 1992
Copris phylax Gillet 1908
Copris poggii Ochi & Kon 2005
Copris potanini Semenov 1891
Copris prevosti Josso 2011
Copris pseudobootes Ferreira 1962
Copris pseudomoffartsi Moretto 2011
Copris pseudosinicus Balthasar 1958
Copris pueli Mollandin De Boissy 1905
Copris punctatus Gillet 1910
Copris puncticollis Boheman 1857
Copris punctipennis Boucomont 1914
Copris rebouchei Harold 1869
Copris remotus Le Conte 1866
Copris renwarti Nguyen-phung & Cambefort 1986
Copris repertus Walker 1858
Copris ribbei Lansberge 1886
Copris riekoae Ochi & Kon 2003
Copris ritsemae Harold 1875
Copris rugosus Gillet 1908
Copris ruricola Balthasar 1933
Copris sallei Harold 1869
Copris sarpedon Harold 1868
Copris saundersi Harold 1869
Copris schoolmeestersi Ochi & Kon 2010
Copris scorpio Zelenka 1993
Copris serius Nguyen-phung 1987
Copris sexdentatus Thunberg 1818
Copris siamensis Gillet 1910
Copris siangensis Biswas 1980
Copris sierrensis Matthews 1961
Copris simonettai Cambefort 1992
Copris simulator Nguyen-phung & Cambefort 1986
Copris singularis Nguyen-phung & Cambefort 1986
Copris sinicus Hope 1842
Copris sinon (Olivier 1789)
Copris sodalis Walker 1858
Copris sonensis Bui, Dumack & Bonkowski 2018
Copris sorex Balthasar 1942
Copris sphaeropterus Harold 1877
Copris spinator Harold 1881
Copris subdolus Balthasar 1958
Copris subpunctatus Gillet 1910
Copris subsidens Péringuey 1901
Copris sumatrensis Gillet 1921
Copris susanae Darling & Génier 2018
Copris szechouanicus Balthasar 1958
Copris tetraodon Gillet 1910
Copris tridentatus Solis & Kohlmann 2003
Copris tripartitus Waterhouse 1875
Copris truncatus Felsche 1901
Copris typhoeus Gerstaecker 1884
Copris umbilicatus Abeille De Perrin 1901
Copris urus Boheman 1857
Copris usambaricus Gillet 1908
Copris vankhaii Nguyen-phung 1988
Copris victorini Boheman 1857
Copris vietnamicus Kabakov 1994
Copris vietnamimontanus Ochi, Kon & Pham 2018
Copris vilhenai Ferreira 1962
Copris vrydaghi Ferreira 1962
Copris warneri Mccleve & Kohlmann 2005
Copris wiesei Kolbe 1914
Copris youngai Balthasar 1967
Copris zhangi Ochi, Kon & Bai 2009
Acanthonitis
Aliuscanthoniola
Allonitis
Anoctus
Anomiopsoides
Anonychonitis
Anoplodrepanus
Apotolamprus
Attavicinus
Ausmontins
Bdelyrus
Besourenga
Byrrhidium
Cambefortantus
Cambefortius
Canthon
Canthonidia
Catharsiocopris
Catharsius
Chalcocopris
Cheironitis
Clypeodrepanus
Copridaspidus
Coprophanaeus
Delopleurus
Deltochilum
Deltorhinum
Dendropaemon
Digitonthophagus
Diorygopyx
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drogo
Dwesasilvasedis
Epidrepanus
Escarabaeus
Eudinopus
Euoniticellus
Eusaproecius
Frankenbergerius
Gilletellus
Glyphoderus
Gymnopleurus
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holocephalus
Homocopris
Homocorpis
Hypocanthidium
Isocopris
Jossonthophagus
Latodrepanus
Liatongus
Martinezidium
Megathopa
Melanocanthon
Mentophilus
Microcopris
Mimonthophagus
Mnematidium
Monoplistes
Morettius
Neosaproecius
Ochicanthon
Oniticellus
Onitis
Onthophagus
Oxysternon
Pachylomera
Paracanthon
Paracryptocanthon
Paragymnopleurus
Paronthobium
Pedaria
Phaedotrogus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Scaptocnemis
Scarabaeus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Silvinha
Streblopus
Sulcophanaeus
Temnoplectron
Theotimius
Trichillidium
Trichillum
Tropidonitis
178 additional siblings truncated for brevity.
See the resource file for a full list.

Catalogue of Life 7 February 2023

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Scarabaeinae Latreille 1802
Coprini Leach 1815
Copris Geoffroy 1762 (this page)
Copris subgen. Copris Geoffroy 1762
Copris subgen. Paracopris Balthasar 1939
Copris subgen. Sinocopris Ochi, Kon & Bai 2009
Canthidium Erichson 1847
Catharsiocopris Balthasar 1967
Catharsius Hope 1837
Copridaspidus Boucomont 1920
Coptodactyla Burmeister 1846
Cretonitis Nikolajev 2007
Heliocopris Hope 1837
Holocanthon Martinez & Pereira 1956
Homocopris Burmeister 1842
Litocopris Waterhouse 1891
Macroderes Westwood 1842
Metacatharsius Paulian 1939
Microcopris Balthasar 1958
Ontherus Erichson 1847
Pedaria Laporte 1832
Pseudocopris Ferreira 1960
Pseudopedaria Felsche 1904
Synapsis Bates 1868
Thyregis Blackburn 1904
Xinidium Harold 1869

Catalogue of Life 7 February 2023

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeoidea
Scarabaeidae Latreille 1802
Scarabaeinae Latreille 1802
Coprini Leach 1815
Copris Geoffroy 1762
Copris subgen. Copris Geoffroy 1762 (this page)
Copris (Copris) acutidens Motschulsky 1860
Copris (Copris) afgoi Cambefort 1992
Copris (Copris) agnus Sharp 1875
Copris (Copris) algol Nguyen-phung 1989
Copris (Copris) amabilis Kolbe 1914
Copris (Copris) amazonicus Darling & Génier 2018
Copris (Copris) amyntor Klug 1855
Copris (Copris) anceus (Olivier 1789)
Copris (Copris) angusticornis Arrow 1933
Copris (Copris) angustus Nguyen-phung & Cambefort 1986
Copris (Copris) antares Ferreira 1958
Copris (Copris) arcturus Gillet 1907
Copris (Copris) ares Zelenka 1993
Copris (Copris) arimotoi Ochi & Araya 1992
Copris (Copris) arizonensis Schaeffer 1906
Copris (Copris) armatus Harold 1869
Copris (Copris) armeniacus Faldermann 1836
Copris (Copris) armiger Gillet 1910
Copris (Copris) arrowi Felsche 1910
Copris (Copris) aspericollis Gillet 1910
Copris (Copris) atropolitus Gillet 1910
Copris (Copris) basilewskyi Ferreira 1962
Copris (Copris) bellator Chevrolat 1844
Copris (Copris) bengalensis Gillet 1911
Copris (Copris) bidens Kolbe 1893
Copris (Copris) bootes Klug 1855
Copris (Copris) boucardi Harold 1869
Copris (Copris) bovinus Gillet 1908
Copris (Copris) brachypterus Nomura 1964
Copris (Copris) brevicornis Darling & Génier 2018
Copris (Copris) brittoni Ferreira 1962
Copris (Copris) caelatus (Fabricius 1794)
Copris (Copris) caliginosus Kohlmann, Cano & Delgado 2003
Copris (Copris) cambeforti Nguyen-phung 1988
Copris (Copris) cambodiensis Ochi, Kon & Kawahara 2008
Copris (Copris) camerunus Felsche 1904
Copris (Copris) caobangensis Bui, Dumack & Bonkowski 2018
Copris (Copris) capensis Waterhouse 1891
Copris (Copris) carinicus Gillet 1910
Copris (Copris) carmelita Fabricius 1801
Copris (Copris) cassius Péringuey 1901
Copris (Copris) celebensis Lansberge 1886
Copris (Copris) chimalapensis Mora-Aguilar & Delgado 2015
Copris (Copris) colmanti Gillet 1908
Copris (Copris) complexus Nguyen-phung 1989
Copris (Copris) confucius Harold 1877
Copris (Copris) coriarius Gillet 1907
Copris (Copris) cornifrons Boheman 1860
Copris (Copris) corniger Sahlberg 1823
Copris (Copris) corpulentus Gillet 1910
Copris (Copris) costaricensis Gahan 1894
Copris (Copris) crassus Deschodt & Davis 2015
Copris (Copris) cribricollis Gillet 1910
Copris (Copris) curlettii Moretto 2011
Copris (Copris) davidi Darling & Génier 2018
Copris (Copris) davisi Nguyen-phung & Cambefort 1986
Copris (Copris) decellei Cambefort 1992
Copris (Copris) delicatus Arrow 1931
Copris (Copris) denticulatus Nguyen-phung 1988
Copris (Copris) diversus Waterhouse 1891
Copris (Copris) draco Arrow 1906
Copris (Copris) dracunculus Ferreira 1959
Copris (Copris) druidum Heer 1862
Copris (Copris) dudleyi Cambefort 1992
Copris (Copris) eburneus Cambefort 1992
Copris (Copris) elpheneroides Felsche 1910
Copris (Copris) elphenor Klug 1855
Copris (Copris) erratus Lansberge 1886
Copris (Copris) evanidus Klug 1855
Copris (Copris) fallaciosus Gillet 1907
Copris (Copris) fallax Felsche 1910
Copris (Copris) fidius (Olivier 1789)
Copris (Copris) frankenbergerianus Balthasar 1958
Copris (Copris) freyi Ferreira 1961
Copris (Copris) fricator (Fabricius 1787)
Copris (Copris) fukiensis Balthasar 1952
Copris (Copris) garambae Cambefort 1992
Copris (Copris) gazellarum Gillet 1918
Copris (Copris) gibbulus Lansberge 1886
Copris (Copris) gilleti Kolbe 1907
Copris (Copris) gladiator Arrow 1933
Copris (Copris) gopheri Hubbard 1894
Copris (Copris) gracilis Waterhouse 1891
Copris (Copris) graueri Kolbe 1914
Copris (Copris) halffteri Matthews 1959
Copris (Copris) harrisi Waterhouse 1891
Copris (Copris) hartli Frey 1975
Copris (Copris) hatayamai Ochi & Araya 1992
Copris (Copris) hispanus (Linnaeus 1764)
Copris (Copris) howdeni Matthews & Halffter 1959
Copris (Copris) humilis Gillet 1908
Copris (Copris) hybridus Gillet 1910
Copris (Copris) igualensis Warner 1990
Copris (Copris) illotus Balthasar 1942
Copris (Copris) inaequabilis Zhang 1997
Copris (Copris) incertus Say 1835
Copris (Copris) inemarginatus Blatchley 1918
Copris (Copris) inhalatus Quedenfeldt 1884
Copris (Copris) insidiosus Péringuey 1901
Copris (Copris) integer Reiche 1850
Copris (Copris) interioris Kolbe 1897
Copris (Copris) iris Sharp 1875
Copris (Copris) jacchoides Nguyen-phung & Cambefort 1987
Copris (Copris) jacchus (Fabricius 1775)
Copris (Copris) jahi Nguyen-phung & Cambefort 1986
Copris (Copris) johkii Ochi & Kon 2014
Copris (Copris) juanmorai Mora-Aguilar & Delgado 2015
Copris (Copris) jucundus Gillet 1932
Copris (Copris) kachinensis Ochi, Kon & Kawahara 2008
Copris (Copris) kartlinus Kabakov 1988
Copris (Copris) kasanka Josso & Prévost 2016
Copris (Copris) katangae Cambefort & Nguyen-phung 1996
Copris (Copris) keralensis Gill 1986
Copris (Copris) klapperichi Balthasar 1942
Copris (Copris) klugi Harold 1869
Copris (Copris) laeviceps Harold 1869
Copris (Copris) laiiformis Nguyen-phung & Cambefort 1987
Copris (Copris) laioides Boucomont 1932
Copris (Copris) laius Harold 1868
Copris (Copris) lannathai Hanboonsong, Masumoto & Ochi 2003
Copris (Copris) laosensis Ochi & Kon 2021
Copris (Copris) leakeyorum Paulian 1976
Copris (Copris) lecontei Matthews 1961
Copris (Copris) leruei Josso & Prévost 2017
Copris (Copris) lugubris Boheman 1858
Copris (Copris) lunaris (Linnaeus 1758)
Copris (Copris) macacus Lansberge 1886
Copris (Copris) macclevei Warner 1990
Copris (Copris) macer Péringuey 1901
Copris (Copris) maesi Ratcliffe 1998
Copris (Copris) magicus Harold 1881
Copris (Copris) manni Ochi, Kon & Bai 2009
Copris (Copris) marcoberrai Ochi, Kon & Pham 2019
Copris (Copris) marcus Gillet 1921
Copris (Copris) martinae Nguyen-phung 1988
Copris (Copris) martinezi Matthews & Halffter 1968
Copris (Copris) matthewsi Delgado & Kohlmann 2001
Copris (Copris) medogensis Zhang 1988
Copris (Copris) megaceratoides Waterhouse 1891
Copris (Copris) megasoma Matthews & Halffter 1959
Copris (Copris) mesacanthus Harold 1878
Copris (Copris) mexicanus Matthews & Halffter 1959
Copris (Copris) micros Ochi & Kon 2021
Copris (Copris) minutus (Drury 1773)
Copris (Copris) moechus Le Conte 1854
Copris (Copris) moffartsi Gillet 1907
Copris (Copris) mongkhoni Hanboonsong, Masumoto & Ochi 2003
Copris (Copris) montivagus Gillet 1908
Copris (Copris) morettoi Josso & Prévost 2020
Copris (Copris) moroni Darling & Génier 2018
Copris (Copris) morphaeus Gillet 1932
Copris (Copris) mourgliai Zelenka 1993
Copris (Copris) neglectus Moxey 1963
Copris (Copris) neotridens Génier & Prévost 2017
Copris (Copris) nepos Gillet 1908
Copris (Copris) nevinsoni Waterhouse 1891
Copris (Copris) novaki Zelenka 1992
Copris (Copris) nubilosus Kohlmann, Cano & Delgado 2003
Copris (Copris) numa Lansberge 1886
Copris (Copris) obesus Boheman 1857
Copris (Copris) orion Klug 1835
Copris (Copris) orphanus Guérin-Méneville 1847
Copris (Copris) parasonensis Ochi, Kon & Pham 2019
Copris (Copris) paraszechouanicus Ochi, Kon & Pham 2018
Copris (Copris) pauliani Nguyen-phung & Cambefort 1986
Copris (Copris) phungae Cambefort 1992
Copris (Copris) phylax Gillet 1908
Copris (Copris) poggii Ochi & Kon 2005
Copris (Copris) potanini Semenov 1891
Copris (Copris) prevosti Josso 2011
Copris (Copris) pristinus Pierce 1946
Copris (Copris) pseudobootes Ferreira 1962
Copris (Copris) pseudomoffartsi Moretto 2011
Copris (Copris) pseudosinicus Balthasar 1958
Copris (Copris) punctatus Gillet 1910
Copris (Copris) puncticollis Boheman 1857
Copris (Copris) punctipennis Boucomont 1914
Copris (Copris) rebouchei Harold 1869
Copris (Copris) remotus Le Conte 1866
Copris (Copris) renwarti Nguyen-phung & Cambefort 1986
Copris (Copris) repertus Walker 1858
Copris (Copris) ribbei Lansberge 1886
Copris (Copris) riekoae Ochi & Kon 2003
Copris (Copris) ritsemae Harold 1875
Copris (Copris) rugosus Gillet 1908
Copris (Copris) ruricola Balthasar 1933
Copris (Copris) sallei Harold 1869
Copris (Copris) sarpedon Harold 1868
Copris (Copris) saundersi Harold 1869
Copris (Copris) schoolmeestersi Ochi & Kon 2010
Copris (Copris) scorpio Zelenka 1993
Copris (Copris) serius Nguyen-phung 1987
Copris (Copris) sexdentatus Thunberg 1818
Copris (Copris) siamensis Gillet 1910
Copris (Copris) siangensis Biswas 1980
Copris (Copris) sierrensis Matthews 1961
Copris (Copris) simonettai Cambefort 1992
Copris (Copris) simulator Nguyen-phung & Cambefort 1986
Copris (Copris) singularis Nguyen-phung & Cambefort 1986
Copris (Copris) sinicus Hope 1842
Copris (Copris) sinon (Olivier 1789)
Copris (Copris) sodalis Walker 1858
Copris (Copris) sonensis Bui, Dumack & Bonkowski 2018
Copris (Copris) sorex Balthasar 1942
Copris (Copris) sphaeropterus Harold 1877
Copris (Copris) spinator Harold 1881
Copris (Copris) subdolus Balthasar 1958
Copris (Copris) subpunctatus Gillet 1910
Copris (Copris) subsidens Péringuey 1901
Copris (Copris) subterraneus Heer 1862
Copris (Copris) sumatrensis Gillet 1921
Copris (Copris) susanae Darling & Génier 2018
Copris (Copris) szechouanicus Balthasar 1958
Copris (Copris) tetraodon Gillet 1910
Copris (Copris) tricorniprothoracicus Ochi, Kon & Kawahara 2020
Copris (Copris) tridentatus Solis & Kohlmann 2003
Copris (Copris) tripartitus Waterhouse 1875
Copris (Copris) truncatus Felsche 1901
Copris (Copris) typhoeus Gerstaecker 1884
Copris (Copris) urus Boheman 1857
Copris (Copris) usambaricus Gillet 1908
Copris (Copris) vankhaii Nguyen-phung 1988
Copris (Copris) victorini Boheman 1857
Copris (Copris) vietnamicus Kabakov 1994
Copris (Copris) vietnamimontanus Ochi, Kon & Pham 2018
Copris (Copris) vilhenai Ferreira 1962
Copris (Copris) vrydaghi Ferreira 1962
Copris (Copris) warneri Mccleve & Kohlmann 2005
Copris (Copris) wiesei Kolbe 1914
Copris (Copris) youngai Balthasar 1967
Copris (Copris) zhangi Ochi, Kon & Bai 2009
Copris subgen. Paracopris Balthasar 1939
Copris subgen. Sinocopris Ochi, Kon & Bai 2009

Dynamic Hierarchy Version 2.2 - Test

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Copris (this page)
Copris (Paracopris) andrewesi Waterhouse 1891
Copris (Paracopris) anomiopseoides Boucomont 1924
Copris (Paracopris) cariniceps Felsche 1910
Copris (Paracopris) compressipennis Gillet 1910
Copris (Paracopris) cribratus Gillet 1927
Copris (Paracopris) davisoni Waterhouse 1891
Copris (Paracopris) excisus Waterhouse 1891
Copris (Paracopris) furciceps Felsche 1910
Copris (Paracopris) imitans Felsche 1910
Copris (Paracopris) kasagii Ochi & Kon 1996
Copris (Paracopris) pedarioides Lansberge 1886
Copris (Paracopris) punctulatus Wiedemann 1823
Copris (Paracopris) punjabensis Gillet 1921
Copris (Paracopris) ramosiceps Gillet 1921
Copris (Paracopris) signatus Walker 1858
Copris (Paracopris) surdus Arrow 1931
Copris (Sinocopris) barclayi Ochi, Kon & Bai 2009
Copris (Sinocopris) basipunctatus Balthasar 1942
Copris (Sinocopris) cheni Ochi, Kon & Bai 2007
Copris (Sinocopris) kiuchii Masumoto 1989
Copris (Sinocopris) kusakabei Ochi, Kon & Kawahara 2005
Copris (Sinocopris) laevigatus Gillet 1927
Copris (Sinocopris) obenbergeri Balthasar 1933
Copris (Sinocopris) ochus (Motschulsky 1860)
Copris (Sinocopris) parapecuarius Ochi, Kon & Kawahara 2008
Copris (Sinocopris) pecuarius Lewis 1884
Copris (Sinocopris) pseudochus Ochi & Kon 2004
Copris (Sinocopris) quasilaevigatus Ochi, Kon & Bai 2007
Copris (Sinocopris) sabinus Gillet 1910
Copris (Sinocopris) sacontala Redtenbacher 1848
Copris (Sinocopris) tsukamotoi Ochi, Kon & Kawahara 2008
Copris (Sinocopris) uenoi Hanboonsong, Masumoto & Ochi 2003
Copris acutidens Motschulsky 1860
Copris afgoi Cambefort 1992
Copris agnus Sharp 1875
Copris algol Nguyen-phung 1989
Copris amabilis Kolbe 1914
Copris amazonicus Darling & Génier 2018
Copris amyntor Klug 1855
Copris anceus (Olivier 1789)
Copris angusticornis Arrow 1933
Copris angustus Nguyen-phung & Cambefort 1986
Copris antares Ferreira 1958
Copris arcturus Gillet 1907
Copris ares Zelenka 1993
Copris arimotoi Ochi & Araya 1992
Copris arizonensis Schaeffer 1906
Copris armatus Harold 1869
Copris armeniacus Faldermann 1835
Copris armiger Gillet 1910
Copris arrowi Felsche 1910
Copris aspericollis Gillet 1910
Copris atropolitus Gillet 1910
Copris basilewskyi Ferreira 1962
Copris bellator Chevrolat 1844
Copris bengalensis Gillet 1911
Copris bidens Kolbe 1893
Copris bootes Klug 1855
Copris boucardi Harold 1869
Copris bovinus Gillet 1908
Copris brachypterus Nomura 1964
Copris brevicornis Darling & Génier 2018
Copris brittoni Ferreira 1962
Copris caelatus (Fabricius 1794)
Copris caliginosus Kohlmann, Cano & Delgado 2003
Copris cambeforti Nguyen-phung 1988
Copris cambodiensis Ochi, Kon & Kawahara 2008
Copris camerunus Felsche 1904
Copris caobangensis Bui, Dumack & Bonkowski 2018
Copris capensis Waterhouse 1891
Copris carinicus Gillet 1910
Copris carmelita Fabricius 1801
Copris cassius Péringuey 1901
Copris celebensis Lansberge 1886
Copris chimalapensis Mora-Aguilar & Delgado 2015
Copris colmanti Gillet 1908
Copris complexus Nguyen-phung 1989
Copris confucius Harold 1877
Copris coriarius Gillet 1907
Copris cornifrons Boheman 1860
Copris corniger Sahlberg 1823
Copris corpulentus Gillet 1910
Copris costaricensis Gahan 1894
Copris crassus Deschodt & Davis 2015
Copris cribricollis Gillet 1910
Copris curlettii Moretto 2011
Copris davidi Darling & Génier 2018
Copris davisi Nguyen-phung & Cambefort 1986
Copris decellei Cambefort 1992
Copris delicatus Arrow 1931
Copris denticulatus Nguyen-phung 1988
Copris diversus Waterhouse 1891
Copris draco Arrow 1906
Copris dracunculus Ferreira 1959
Copris dudleyi Cambefort 1992
Copris eburneus Cambefort 1992
Copris elpheneroides Felsche 1910
Copris elphenor Klug 1855
Copris erratus Lansberge 1886
Copris evanidus Klug 1855
Copris fallaciosus Gillet 1907
Copris fallax Felsche 1910
Copris felschei Reitter 1892
Copris fidius (Olivier 1789)
Copris frankenbergerianus Balthasar 1958
Copris freyi Ferreira 1961
Copris fricator (Fabricius 1787)
Copris fukiensis Balthasar 1952
Copris garambae Cambefort 1992
Copris gazellarum Gillet 1918
Copris gibbulus Lansberge 1886
Copris gilleti Kolbe 1907
Copris gladiator Arrow 1933
Copris gopheri Hubbard 1894
Copris gracilis Waterhouse 1891
Copris graueri Kolbe 1914
Copris halffteri Matthews 1959
Copris harrisi Waterhouse 1891
Copris hartli Frey 1975
Copris hatayamai Ochi & Araya 1992
Copris hispanus (Linnaeus 1764)
Copris howdeni Matthews & Halffter 1959
Copris humilis Gillet 1908
Copris hybridus Gillet 1910
Copris igualensis Warner 1990
Copris illotus Balthasar 1942
Copris inaequabilis Zhang 1997
Copris incertus Say 1835
Copris inemarginatus Blatchley 1918
Copris inhalatus Quedenfeldt 1884
Copris insidiosus Péringuey 1901
Copris integer Reiche 1847
Copris interioris Kolbe 1897
Copris iris Sharp 1875
Copris jacchoides Nguyen-phung & Cambefort 1987
Copris jacchus (Fabricius 1775)
Copris jahi Nguyen-phung & Cambefort 1986
Copris johkii Ochi & Kon 2014
Copris juanmorai Mora-Aguilar & Delgado 2015
Copris jucundus Gillet 1932
Copris kachinensis Ochi, Kon & Kawahara 2008
Copris kasanka Josso & Prévost 2016
Copris katangae Cambefort & Nguyen-phung 1996
Copris keralensis Gill 1986
Copris klapperichi Balthasar 1942
Copris klugi Harold 1869
Copris laeviceps Harold 1869
Copris laiiformis Nguyen-phung & Cambefort 1987
Copris laioides Boucomont 1932
Copris laius Harold 1868
Copris lannathai Hanboonsong, Masumoto & Ochi 2003
Copris lecontei Matthews 1961
Copris leruei Josso & Prévost 2017
Copris lugubris Boheman 1858
Copris lunaris (Linnaeus 1758)
Copris macacus Lansberge 1886
Copris macclevei Warner 1990
Copris macer Péringuey 1901
Copris maesi Ratcliffe 1998
Copris magicus Harold 1881
Copris manni Ochi, Kon & Bai 2009
Copris marcus Gillet 1921
Copris martinae Nguyen-phung 1988
Copris martinezi Matthews & Halffter 1968
Copris matthewsi Delgado & Kohlmann 2001
Copris medogensis Zhang 1988
Copris megaceratoides Waterhouse 1891
Copris megasoma Matthews & Halffter 1959
Copris mesacanthus Harold 1878
Copris mexicanus Matthews & Halffter 1959
Copris minutus (Drury 1773)
Copris moechus Le Conte 1854
Copris moffartsi Gillet 1907
Copris mongkhoni Hanboonsong, Masumoto & Ochi 2003
Copris montivagus Gillet 1908
Copris moroni Darling & Génier 2018
Copris morphaeus Gillet 1932
Copris mourgliai Zelenka 1993
Copris neglectus Moxey 1963
Copris neotridens Génier & Prévost 2017
Copris nepos Gillet 1908
Copris nevinsoni Waterhouse 1891
Copris novaki Zelenka 1992
Copris nubilosus Kohlmann, Cano & Delgado 2003
Copris numa Lansberge 1886
Copris obesus Boheman 1857
Copris orion Klug 1835
Copris orphanus Guérin-Méneville 1847
Copris paraszechouanicus Ochi, Kon & Pham 2018
Copris pauliani Nguyen-phung & Cambefort 1986
Copris phungae Cambefort 1992
Copris phylax Gillet 1908
Copris poggii Ochi & Kon 2005
Copris potanini Semenov 1891
Copris prevosti Josso 2011
Copris pseudobootes Ferreira 1962
Copris pseudomoffartsi Moretto 2011
Copris pseudosinicus Balthasar 1958
Copris pueli Mollandin De Boissy 1905
Copris punctatus Gillet 1910
Copris puncticollis Boheman 1857
Copris punctipennis Boucomont 1914
Copris rebouchei Harold 1869
Copris remotus Le Conte 1866
Copris renwarti Nguyen-phung & Cambefort 1986
Copris repertus Walker 1858
Copris ribbei Lansberge 1886
Copris riekoae Ochi & Kon 2003
Copris ritsemae Harold 1875
Copris rugosus Gillet 1908
Copris ruricola Balthasar 1933
Copris sallei Harold 1869
Copris sarpedon Harold 1868
Copris saundersi Harold 1869
Copris schoolmeestersi Ochi & Kon 2010
Copris scorpio Zelenka 1993
Copris serius Nguyen-phung 1987
Copris sexdentatus Thunberg 1818
Copris siamensis Gillet 1910
Copris siangensis Biswas 1980
Copris sierrensis Matthews 1961
Copris simonettai Cambefort 1992
Copris simulator Nguyen-phung & Cambefort 1986
Copris singularis Nguyen-phung & Cambefort 1986
Copris sinicus Hope 1842
Copris sinon (Olivier 1789)
Copris sodalis Walker 1858
Copris sonensis Bui, Dumack & Bonkowski 2018
Copris sorex Balthasar 1942
Copris sphaeropterus Harold 1877
Copris spinator Harold 1881
Copris subdolus Balthasar 1958
Copris subpunctatus Gillet 1910
Copris subsidens Péringuey 1901
Copris sumatrensis Gillet 1921
Copris susanae Darling & Génier 2018
Copris szechouanicus Balthasar 1958
Copris tetraodon Gillet 1910
Copris tridentatus Solis & Kohlmann 2003
Copris tripartitus Waterhouse 1875
Copris truncatus Felsche 1901
Copris typhoeus Gerstaecker 1884
Copris umbilicatus Abeille De Perrin 1901
Copris urus Boheman 1857
Copris usambaricus Gillet 1908
Copris vankhaii Nguyen-phung 1988
Copris victorini Boheman 1857
Copris vietnamicus Kabakov 1994
Copris vietnamimontanus Ochi, Kon & Pham 2018
Copris vilhenai Ferreira 1962
Copris vrydaghi Ferreira 1962
Copris warneri Mccleve & Kohlmann 2005
Copris wiesei Kolbe 1914
Copris youngai Balthasar 1967
Copris zhangi Ochi, Kon & Bai 2009
Acanthonitis
Aliuscanthoniola
Allonitis
Anoctus
Anomiopsoides
Anonychonitis
Anoplodrepanus
Apotolamprus
Attavicinus
Ausmontins
Bdelyrus
Besourenga
Byrrhidium
Cambefortantus
Cambefortius
Canthon
Canthonidia
Catharsiocopris
Catharsius
Chalcocopris
Cheironitis
Clypeodrepanus
Copridaspidus
Coprophanaeus
Delopleurus
Deltochilum
Deltorhinum
Dendropaemon
Digitonthophagus
Diorygopyx
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drogo
Dwesasilvasedis
Epidrepanus
Escarabaeus
Eudinopus
Euoniticellus
Eusaproecius
Frankenbergerius
Gilletellus
Glyphoderus
Gymnopleurus
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holocephalus
Homocopris
Homocorpis
Hypocanthidium
Isocopris
Jossonthophagus
Latodrepanus
Liatongus
Martinezidium
Megathopa
Melanocanthon
Mentophilus
Microcopris
Mimonthophagus
Mnematidium
Monoplistes
Morettius
Neosaproecius
Ochicanthon
Oniticellus
Onitis
Onthophagus
Oxysternon
Pachylomera
Paracanthon
Paracryptocanthon
Paragymnopleurus
Paronthobium
Pedaria
Phaedotrogus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Scaptocnemis
Scarabaeus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Silvinha
Streblopus
Sulcophanaeus
Temnoplectron
Theotimius
Trichillidium
Trichillum
Tropidonitis
178 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Copris (this page)
Copris (Paracopris) andrewesi Waterhouse 1891
Copris (Paracopris) anomiopseoides Boucomont 1924
Copris (Paracopris) cariniceps Felsche 1910
Copris (Paracopris) compressipennis Gillet 1910
Copris (Paracopris) cribratus Gillet 1927
Copris (Paracopris) davisoni Waterhouse 1891
Copris (Paracopris) excisus Waterhouse 1891
Copris (Paracopris) furciceps Felsche 1910
Copris (Paracopris) imitans Felsche 1910
Copris (Paracopris) kasagii Ochi & Kon 1996
Copris (Paracopris) pedarioides Lansberge 1886
Copris (Paracopris) punctulatus Wiedemann 1823
Copris (Paracopris) punjabensis Gillet 1921
Copris (Paracopris) ramosiceps Gillet 1921
Copris (Paracopris) signatus Walker 1858
Copris (Paracopris) surdus Arrow 1931
Copris (Sinocopris) barclayi Ochi, Kon & Bai 2009
Copris (Sinocopris) basipunctatus Balthasar 1942
Copris (Sinocopris) cheni Ochi, Kon & Bai 2007
Copris (Sinocopris) kiuchii Masumoto 1989
Copris (Sinocopris) kusakabei Ochi, Kon & Kawahara 2005
Copris (Sinocopris) laevigatus Gillet 1927
Copris (Sinocopris) obenbergeri Balthasar 1933
Copris (Sinocopris) ochus (Motschulsky 1860)
Copris (Sinocopris) parapecuarius Ochi, Kon & Kawahara 2008
Copris (Sinocopris) pecuarius Lewis 1884
Copris (Sinocopris) pseudochus Ochi & Kon 2004
Copris (Sinocopris) quasilaevigatus Ochi, Kon & Bai 2007
Copris (Sinocopris) sabinus Gillet 1910
Copris (Sinocopris) sacontala Redtenbacher 1848
Copris (Sinocopris) tsukamotoi Ochi, Kon & Kawahara 2008
Copris (Sinocopris) uenoi Hanboonsong, Masumoto & Ochi 2003
Copris (Sinocopris) yangi Ochi, Kon & Bai 2007
Copris acutidens Motschulsky 1860
Copris afgoi Cambefort 1992
Copris agnus Sharp 1875
Copris algol Nguyen-phung 1989
Copris amabilis Kolbe 1914
Copris amazonicus Darling & Génier 2018
Copris amyntor Klug 1855
Copris anceus (Olivier 1789)
Copris angusticornis Arrow 1933
Copris angustus Nguyen-phung & Cambefort 1986
Copris antares Ferreira 1958
Copris arcturus Gillet 1907
Copris ares Zelenka 1993
Copris arimotoi Ochi & Araya 1992
Copris arizonensis Schaeffer 1906
Copris armatus Harold 1869
Copris armeniacus Faldermann 1835
Copris armiger Gillet 1910
Copris arrowi Felsche 1910
Copris aspericollis Gillet 1910
Copris atropolitus Gillet 1910
Copris basilewskyi Ferreira 1962
Copris bellator Chevrolat 1844
Copris bengalensis Gillet 1911
Copris bidens Kolbe 1893
Copris bootes Klug 1855
Copris boucardi Harold 1869
Copris bovinus Gillet 1908
Copris brachypterus Nomura 1964
Copris brevicornis Darling & Génier 2018
Copris brittoni Ferreira 1962
Copris caelatus (Fabricius 1794)
Copris caliginosus Kohlmann, Cano & Delgado 2003
Copris cambeforti Nguyen-phung 1988
Copris cambodiensis Ochi, Kon & Kawahara 2008
Copris camerunus Felsche 1904
Copris caobangensis Bui, Dumack & Bonkowski 2018
Copris capensis Waterhouse 1891
Copris carinicus Gillet 1910
Copris carmelita Fabricius 1801
Copris cassius Péringuey 1901
Copris celebensis Lansberge 1886
Copris chimalapensis Mora-Aguilar & Delgado 2015
Copris colmanti Gillet 1908
Copris complexus Nguyen-phung 1989
Copris confucius Harold 1877
Copris coriarius Gillet 1907
Copris cornifrons Boheman 1860
Copris corniger Sahlberg 1823
Copris corpulentus Gillet 1910
Copris costaricensis Gahan 1894
Copris crassus Deschodt & Davis 2015
Copris cribricollis Gillet 1910
Copris curlettii Moretto 2011
Copris davidi Darling & Génier 2018
Copris davisi Nguyen-phung & Cambefort 1986
Copris decellei Cambefort 1992
Copris delicatus Arrow 1931
Copris denticulatus Nguyen-phung 1988
Copris diversus Waterhouse 1891
Copris draco Arrow 1906
Copris dracunculus Ferreira 1959
Copris druidum Heer 1862
Copris dudleyi Cambefort 1992
Copris eburneus Cambefort 1992
Copris elpheneroides Felsche 1910
Copris elphenor Klug 1855
Copris erratus Lansberge 1886
Copris evanidus Klug 1855
Copris fallaciosus Gillet 1907
Copris fallax Felsche 1910
Copris felschei Reitter 1892
Copris fidius (Olivier 1789)
Copris frankenbergerianus Balthasar 1958
Copris freyi Ferreira 1961
Copris fricator (Fabricius 1787)
Copris fukiensis Balthasar 1952
Copris garambae Cambefort 1992
Copris gazellarum Gillet 1918
Copris gibbulus Lansberge 1886
Copris gilleti Kolbe 1907
Copris gladiator Arrow 1933
Copris gopheri Hubbard 1894
Copris gracilis Waterhouse 1891
Copris graueri Kolbe 1914
Copris halffteri Matthews 1959
Copris harrisi Waterhouse 1891
Copris hartli Frey 1975
Copris hatayamai Ochi & Araya 1992
Copris hispanus (Linnaeus 1764)
Copris howdeni Matthews & Halffter 1959
Copris humilis Gillet 1908
Copris hybridus Gillet 1910
Copris igualensis Warner 1990
Copris illotus Balthasar 1942
Copris inaequabilis Zhang 1997
Copris incertus Say 1835
Copris inemarginatus Blatchley 1918
Copris inhalatus Quedenfeldt 1884
Copris insidiosus Péringuey 1901
Copris integer Reiche 1847
Copris interioris Kolbe 1897
Copris iris Sharp 1875
Copris jacchoides Nguyen-phung & Cambefort 1987
Copris jacchus (Fabricius 1775)
Copris jahi Nguyen-phung & Cambefort 1986
Copris johkii Ochi & Kon 2014
Copris juanmorai Mora-Aguilar & Delgado 2015
Copris jucundus Gillet 1932
Copris kachinensis Ochi, Kon & Kawahara 2008
Copris kartlinus Kabakov 1988
Copris kasanka Josso & Prévost 2016
Copris katangae Cambefort & Nguyen-phung 1996
Copris keralensis Gill 1986
Copris klapperichi Balthasar 1942
Copris klugi Harold 1869
Copris laeviceps Harold 1869
Copris laiiformis Nguyen-phung & Cambefort 1987
Copris laioides Boucomont 1932
Copris laius Harold 1868
Copris lannathai Hanboonsong, Masumoto & Ochi 2003
Copris leakeyorum Paulian 1976
Copris lecontei Matthews 1961
Copris leruei Josso & Prévost 2017
Copris lugubris Boheman 1858
Copris lunaris (Linnaeus 1758)
Copris macacus Lansberge 1886
Copris macclevei Warner 1990
Copris macer Péringuey 1901
Copris maesi Ratcliffe 1998
Copris magicus Harold 1881
Copris manni Ochi, Kon & Bai 2009
Copris marcus Gillet 1921
Copris martinae Nguyen-phung 1988
Copris martinezi Matthews & Halffter 1968
Copris matthewsi Delgado & Kohlmann 2001
Copris medogensis Zhang 1988
Copris megaceratoides Waterhouse 1891
Copris megasoma Matthews & Halffter 1959
Copris mesacanthus Harold 1878
Copris mexicanus Matthews & Halffter 1959
Copris minutus (Drury 1773)
Copris misellus Péringuey 1901
Copris moechus Le Conte 1854
Copris moffartsi Gillet 1907
Copris mongkhoni Hanboonsong, Masumoto & Ochi 2003
Copris montivagus Gillet 1908
Copris moroni Darling & Génier 2018
Copris morphaeus Gillet 1932
Copris mourgliai Zelenka 1993
Copris neglectus Moxey 1963
Copris neotridens Génier & Prévost 2017
Copris nepos Gillet 1908
Copris nevinsoni Waterhouse 1891
Copris novaki Zelenka 1992
Copris nubilosus Kohlmann, Cano & Delgado 2003
Copris numa Lansberge 1886
Copris obesus Boheman 1857
Copris orion Klug 1835
Copris orphanus Guérin-Méneville 1847
Copris paraszechouanicus Ochi, Kon & Pham 2018
Copris pauliani Nguyen-phung & Cambefort 1986
Copris phungae Cambefort 1992
Copris phylax Gillet 1908
Copris poggii Ochi & Kon 2005
Copris potanini Semenov 1891
Copris prevosti Josso 2011
Copris pristinus Pierce 1946
Copris pseudobootes Ferreira 1962
Copris pseudomoffartsi Moretto 2011
Copris pseudosinicus Balthasar 1958
Copris pueli Mollandin De Boissy 1905
Copris punctatus Gillet 1910
Copris puncticollis Boheman 1857
Copris punctipennis Boucomont 1914
Copris rebouchei Harold 1869
Copris remotus Le Conte 1866
Copris renwarti Nguyen-phung & Cambefort 1986
Copris repertus Walker 1858
Copris ribbei Lansberge 1886
Copris riekoae Ochi & Kon 2003
Copris ritsemae Harold 1875
Copris rugosus Gillet 1908
Copris ruricola Balthasar 1933
Copris sallei Harold 1869
Copris sarpedon Harold 1868
Copris saundersi Harold 1869
Copris schoolmeestersi Ochi & Kon 2010
Copris scorpio Zelenka 1993
Copris serius Nguyen-phung 1987
Copris sexdentatus Thunberg 1818
Copris siamensis Gillet 1910
Copris siangensis Biswas 1980
Copris sierrensis Matthews 1961
Copris simonettai Cambefort 1992
Copris simulator Nguyen-phung & Cambefort 1986
Copris singularis Nguyen-phung & Cambefort 1986
Copris sinicus Hope 1842
Copris sinon (Olivier 1789)
Copris sodalis Walker 1858
Copris sonensis Bui, Dumack & Bonkowski 2018
Copris sorex Balthasar 1942
Copris sphaeropterus Harold 1877
Copris spinator Harold 1881
Copris subdolus Balthasar 1958
Copris subpunctatus Gillet 1910
Copris subsidens Péringuey 1901
Copris subterraneus Heer 1862
Copris sumatrensis Gillet 1921
Copris susanae Darling & Génier 2018
Copris szechouanicus Balthasar 1958
Copris tetraodon Gillet 1910
Copris tridentatus Solis & Kohlmann 2003
Copris tripartitus Waterhouse 1875
Copris truncatus Felsche 1901
Copris typhoeus Gerstaecker 1884
Copris umbilicatus Abeille de Perrin 1901
Copris urus Boheman 1857
Copris usambaricus Gillet 1908
Copris vankhaii Nguyen-phung 1988
Copris victorini Boheman 1857
Copris vietnamicus Kabakov 1994
Copris vietnamimontanus Ochi, Kon & Pham 2018
Copris vilhenai Ferreira 1962
Copris vrydaghi Ferreira 1962
Copris warneri Mccleve & Kohlmann 2005
Copris wiesei Kolbe 1914
Copris youngai Balthasar 1967
Copris zhangi Ochi, Kon & Bai 2009
Acanthonitis
Aliuscanthoniola
Allonitis
Anoctus
Anomiopsoides
Anonychonitis
Anoplodrepanus
Apotolamprus
Ateuchites
Attavicinus
Ausmontins
Bdelyrus
Besourenga
Byrrhidium
Cambefortantus
Cambefortius
Canthon
Canthonidia
Catharsiocopris
Catharsius
Chalcocopris
Cheironitis
Clypeodrepanus
Copridaspidus
Coprophanaeus
Delopleurus
Deltochilum
Deltorhinum
Dendropaemon
Digitonthophagus
Diorygopyx
Dorbignyolus
Drepanocerus
Drepanoplatynus
Drogo
Dwesasilvasedis
Epidrepanus
Escarabaeus
Eudinopus
Euoniticellus
Eusaproecius
Frankenbergerius
Gilletellus
Glyphoderus
Gymnopleurus
Helictopleurus
Heliocopris
Heteroclitopus
Heteronitis
Holcorobeus
Holocephalus
Homocopris
Homocorpis
Hypocanthidium
Isocopris
Jossonthophagus
Latodrepanus
Liatongus
Martinezidium
Megathopa
Melanocanthon
Microcopris
Mimonthophagus
Mnematidium
Monoplistes
Morettius
Neosaproecius
Ochicanthon
Oniticellus
Onitis
Onthophagus
Oxysternon
Pachylomera
Paracanthon
Paracryptocanthon
Paronthobium
Pedaria
Phaedotrogus
Pinacopodius
Pinacotarsus
Platyonitis
Pleuronitis
Pseudocopris
Pseudonthobium
Pseudopedaria
Pseudosaproecius
Scaptocnemis
Scarabaeus
Scatonomus
Scatrichus
Sceliages
Scybalocanthon
Silvinha
Streblopus
Sulcophanaeus
Temnoplectron
Theotimius
Trichillidium
Trichillum
Tropidonitis
189 additional siblings truncated for brevity.
See the resource file for a full list.

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Copris Geoffroy 1762 (this page)
Copris acutidens Motschulsky 1860
Copris afgoi Cambefort 1992
Copris agnus Sharp 1875
Copris algol Nguyen-phung 1989
Copris amabilis Kolbe 1914
Copris amazonicus Darling & Génier 2018
Copris amyntor Klug 1855
Copris anceus (Olivier 1789)
Copris andrewesi Waterhouse 1891
Copris angusticornis Arrow 1933
Copris angustus Nguyen-phung & Cambefort 1986
Copris anomiopseoides Boucomont 1924
Copris antares Ferreira 1958
Copris arcturus Gillet 1907
Copris ares Zelenka 1993
Copris arimotoi Ochi & Araya 1992
Copris arizonensis Schaeffer 1906
Copris armatus Harold 1869
Copris armeniacus Faldermann 1835
Copris armiger Gillet 1910
Copris arrowi Felsche 1910
Copris aspericollis Gillet 1910
Copris atropolitus Gillet 1910
Copris bacchus Thunberg 1787
Copris barclayi Ochi, Kon & Bai 2009
Copris basilewskyi Ferreira 1962
Copris basipunctatus Balthasar 1942
Copris bellator Chevrolat 1844
Copris bengalensis Gillet 1911
Copris bidens Kolbe 1893
Copris binodis
Copris bootes Klug 1855
Copris boucardi Harold 1869
Copris bovinus Gillet 1908
Copris brachypterus Nomura 1964
Copris brevicornis Darling & Génier 2018
Copris brittoni Ferreira 1962
Copris caelatus (Fabricius 1794)
Copris caliginosus Kohlmann, Cano & Delgado 2003
Copris cambeforti Nguyen-phung 1988
Copris cambodiensis Ochi, Kon & Kawahara 2008
Copris camerunus Felsche 1904
Copris caobangensis Bui, Dumack & Bonkowski 2018
Copris capensis Waterhouse 1891
Copris cariniceps Felsche 1910
Copris carinicus Gillet 1910
Copris carmelita Fabricius 1801
Copris cassius Péringuey 1901
Copris celebensis Lansberge 1886
Copris cheni Ochi, Kon & Bai 2007
Copris chimalapensis Mora-Aguilar & Delgado 2015
Copris ciliatus
Copris colmanti Gillet 1908
Copris complexus Nguyen-phung 1989
Copris compressipennis Gillet 1910
Copris confucius Harold 1877
Copris coriarius Gillet 1907
Copris cornifrons Boheman 1860
Copris corniger Sahlberg 1823
Copris corpulentus Gillet 1910
Copris costaricensis Gahan 1894
Copris costariensis Matthews
Copris crassus Deschodt & Davis 2015
Copris cribratus Gillet 1927
Copris cribricollis Gillet 1910
Copris curlettii Moretto 2011
Copris davidi Darling & Génier 2018
Copris davisi Nguyen-phung & Cambefort 1986
Copris davisoni Waterhouse 1891
Copris decellei Cambefort 1992
Copris delicatus Arrow 1931
Copris denticulatus Nguyen-phung 1988
Copris diversus Waterhouse 1891
Copris draco Arrow 1906
Copris dracunculus Ferreira 1959
Copris druidum Heer 1862
Copris dudleyi Cambefort 1992
Copris eburneus Cambefort 1992
Copris elpheneroides Felsche 1910
Copris elphenor Klug 1855
Copris erratus Lansberge 1886
Copris evanidus Klug 1855
Copris excisus Waterhouse 1891
Copris fallaciosus Gillet 1907
Copris fallax Felsche 1910
Copris felschei Reitter 1892
Copris fidius (Olivier 1789)
Copris frankenbergerianus Balthasar 1958
Copris freyi Ferreira 1961
Copris fricator (Fabricius 1787)
Copris fukiensis Balthasar 1952
Copris furciceps Felsche 1910
Copris garambae Cambefort 1992
Copris gazellarum Gillet 1918
Copris gibbulus Lansberge 1886
Copris gilleti Kolbe 1907
Copris gladiator Arrow 1933
Copris gopheri Hubbard 1894
Copris gracilis Waterhouse 1891
Copris graueri Kolbe 1914
Copris halffteri Matthews 1959
Copris harrisi Waterhouse 1891
Copris hartli Frey 1975
Copris hatayamai Ochi & Araya 1992
Copris hispanus (Linnaeus 1764)
Copris howdeni Matthews & Halffter 1959
Copris humilis Gillet 1908
Copris hybridus Gillet 1910
Copris igualensis Warner 1990
Copris illotus Balthasar 1942
Copris imitans Felsche 1910
Copris inaequabilis Zhang 1997
Copris incertus Say 1835
Copris inemarginatus Blatchley 1918
Copris inhalatus Quedenfeldt 1884
Copris insidiosus Péringuey 1901
Copris integer Reiche 1847
Copris interioris Kolbe 1897
Copris iris Sharp 1875
Copris jacchoides Nguyen-phung & Cambefort 1987
Copris jacchus (Fabricius 1775)
Copris jahi Nguyen-phung & Cambefort 1986
Copris johkii Ochi & Kon 2014
Copris juanmorai Mora-Aguilar & Delgado 2015
Copris jucundus Gillet 1932
Copris kachinensis Ochi, Kon & Kawahara 2008
Copris kartlinus Kabakov 1988
Copris kasagii Ochi & Kon 1996
Copris kasanka Josso & Prévost 2016
Copris katangae Cambefort & Nguyen-phung 1996
Copris keralensis Gill 1986
Copris kiuchii Masumoto 1989
Copris klapperichi Balthasar 1942
Copris klugi Harold 1869
Copris kusakabei Ochi, Kon & Kawahara 2005
Copris laeviceps Harold 1869
Copris laevigatus Gillet 1927
Copris laiiformis Nguyen-phung & Cambefort 1987
Copris laioides Boucomont 1932
Copris laius Harold 1868
Copris lannathai Hanboonsong, Masumoto & Ochi 2003
Copris leakeyorum Paulian 1976
Copris lecontei Matthews 1961
Copris leruei Josso & Prévost 2017
Copris lugubris Boheman 1858
Copris lunaris (Linnaeus 1758)
Copris macacus Lansberge 1886
Copris macclevei Warner 1990
Copris macer Péringuey 1901
Copris maesi Ratcliffe 1998
Copris magicus Harold 1881
Copris manni Ochi, Kon & Bai 2009
Copris marcus Gillet 1921
Copris martinae Nguyen-phung 1988
Copris martinezi Matthews & Halffter 1968
Copris matthewsi Delgado & Kohlmann 2001
Copris medogensis Zhang 1988
Copris megaceratoides Waterhouse 1891
Copris megasoma Matthews & Halffter 1959
Copris mesacanthus Harold 1878
Copris mexicanus Matthews & Halffter 1959
Copris minutus (Drury 1773)
Copris misellus Péringuey 1901
Copris moechus Le Conte 1854
Copris moffartsi Gillet 1907
Copris mongkhoni Hanboonsong, Masumoto & Ochi 2003
Copris montivagus Gillet 1908
Copris moroni Darling & Génier 2018
Copris morphaeus Gillet 1932
Copris mourgliai Zelenka 1993
Copris neglectus Moxey 1963
Copris neotridens Génier & Prévost 2017
Copris nepos Gillet 1908
Copris nevinsoni Waterhouse 1891
Copris novaki Zelenka 1992
Copris nubilosus Kohlmann, Cano & Delgado 2003
Copris numa Lansberge 1886
Copris obenbergeri Balthasar 1933
Copris obesus Boheman 1857
Copris ochus (Motschulsky 1860)
Copris orion Klug 1835
Copris orphanus Guérin-Méneville 1847
Copris parapecuarius Ochi, Kon & Kawahara 2008
Copris paraszechouanicus Ochi, Kon & Pham 2018
Copris pauliani Nguyen-phung & Cambefort 1986
Copris pecuarius Lewis 1884
Copris pedarioides Lansberge 1886
Copris phungae Cambefort 1992
Copris phylax Gillet 1908
Copris poggii Ochi & Kon 2005
Copris potanini Semenov 1891
Copris prevosti Josso 2011
Copris pristinus Pierce 1946
Copris pseudobootes Ferreira 1962
Copris pseudochus Ochi & Kon 2004
Copris pseudomoffartsi Moretto 2011
Copris pseudosinicus Balthasar 1958
Copris pueli Mollandin De Boissy 1905
Copris punctatus Gillet 1910
Copris puncticollis Boheman 1857
Copris punctipennis Boucomont 1914
Copris punctulatus Wiedemann 1823
Copris punjabensis Gillet 1921
Copris quasilaevigatus Ochi, Kon & Bai 2007
Copris ramosiceps Gillet 1921
Copris rebouchei Harold 1869
Copris remotus Le Conte 1866
Copris renwarti Nguyen-phung & Cambefort 1986
Copris repertus Walker 1858
Copris ribbei Lansberge 1886
Copris riekoae Ochi & Kon 2003
Copris ritsemae Harold 1875
Copris rugosus Gillet 1908
Copris ruricola Balthasar 1933
Copris sabinus Gillet 1910
Copris sacontala Redtenbacher 1848
Copris sallei Harold 1869
Copris sarpedon Harold 1868
Copris saundersi Harold 1869
Copris schoolmeestersi Ochi & Kon 2010
Copris scorpio Zelenka 1993
Copris serius Nguyen-phung 1987
Copris sexdentatus Thunberg 1818
Copris siamensis Gillet 1910
Copris siangensis Biswas 1980
Copris sierrensis Matthews 1961
Copris signatus Walker 1858
Copris simonettai Cambefort 1992
Copris simulator Nguyen-phung & Cambefort 1986
Copris singularis Nguyen-phung & Cambefort 1986
Copris sinicus Hope 1842
Copris sinon (Olivier 1789)
Copris sodalis Walker 1858
Copris sonensis Bui, Dumack & Bonkowski 2018
Copris sorex Balthasar 1942
Copris sphaeropterus Harold 1877
Copris spinator Harold 1881
Copris subdolus Balthasar 1958
Copris subpunctatus Gillet 1910
Copris subsidens Péringuey 1901
Copris subterraneus Heer 1862
Copris sumatrensis Gillet 1921
Copris surdus Arrow 1931
Copris susanae Darling & Génier 2018
Copris szechouanicus Balthasar 1958
Copris tetraodon Gillet 1910
Copris tridentatus Solis & Kohlmann 2003
Copris tripartitus Waterhouse 1875
Copris truncatus Felsche 1901
Copris tsukamotoi Ochi, Kon & Kawahara 2008
Copris typhoeus Gerstaecker 1884
Copris uenoi Hanboonsong, Masumoto & Ochi 2003
Copris umbilicatus Abeille De Perrin 1901
Copris urus Boheman 1857
Copris usambaricus Gillet 1908
Copris vankhaii Nguyen-phung 1988
Copris victorini Boheman 1857
Copris vietnamicus Kabakov 1994
Copris vietnamimontanus Ochi, Kon & Pham 2018
Copris vilhenai Ferreira 1962
Copris vrydaghi Ferreira 1962
Copris warneri Mccleve & Kohlmann 2005
Copris wiesei Kolbe 1914
Copris yangi Ochi, Kon & Bai 2007
Copris youngai Balthasar 1967
Copris zhangi Ochi, Kon & Bai 2009
Acanthonitis Janssens 1937
Afrodrepanus Krikken 2009
Agamopus Bates 1887
Amietina Cambefort 1981
Anoctus Sharp 1875
Anonychonitis Janssens 1950
Anoplosiagum Blanchard 1850
Aptenocanthon Matthews 1974
Aptychonitis Janssens 1937
Ateuchus Weber 1801
Attavicinus Philips & Bell 2008
Ausmontins Deschodt & Davis 2018
Baloghonthobium Paulian 1986
Bohepilissus Paulian 1975
Bolbites Harold 1868
Caccobiomorphus Balthasar 1964
Caccobius Thomson 1859
Caeconthobium Paulian 1984
Cambefortius Branco 1989
Canthochilum Chapin 1934
Canthonosoma Mac Leay 1871
Catharsiocopris Balthasar 1967
Cephalodesmius Westwood 1842
Cetonischema Reitter 1899
Cheironitis Lansberge 1875
Chiron Macleay 1819
Clypeodrepanus Krikken 2009
Copridaspidus Boucomont 1920
Coprophanaeus D' Olsoufieff 1924
Coptorhina Hope 1833
Ctilocephala Burmeister 1855
Degallieridium
Deltepilissus Silvério Pereira 1949
Dendropaemon Perty 1830
Diathermus Kraatz 1897
Dicranocara Frolov & Scholtz 2003
Dicronorhina Hope 1837
Dorbignyolus Branco 1989
Ennearabdus Van Lansberge 1874
Eucetonia Schoch 1894
Euglypta Mohnike 1873
Eusaproecius Branco 1989
Feeridium Vaz de Mello 2008
Garreta Janssens 1940
Glaphyrocanthon Martínez 1948
Glyphoderus Westwood 1837
Haematonotus Kraatz 1880
Hammondantus Cambefort 1978
Heterorrhina
Holocanthon Martínez & Pereira 1956
Hybosorites Nikolajev 1996
Lachnosterna Hope 1837
Leotrichillum
Lithoscarabaeus Nikolajev 1992
Lophorrhinides
Macroderes Westwood 1842
Macrophylla Hope 1837
Mecinonota Kraatz 1892
Megalorhina Westwood 1847
Melanocanthon Halffter 1958
Mentophilus Laporte de Castelnau 1840
Metacatharsius Paulian 1939
Micrelaphinis Schoch 1896
Miltotrogus Reitter 1902
Mimeoma Casey 1915
Mimonthophagus Balthasar 1963
Mnematium Macleay 1821
Nebulasilvius Deschodt & Scholtz 2008
Neosaproecius Branco 1990
Neptunides Thomson 1879
Nesosisyphus Vinson 1946
Nesovinsonia Martínez & Pereira 1959
Nunoidium
Onychothecus Boucomont 1912
Oxycetonia Arrow 1910
Oxygrylius Casey 1915
Pachylomera Kirby 1832
Pachysoma Macleay 1821
Paroniticellus Balthasar 1963
Parvuhowdenius Deschodt & Scholtz 2008
Peckolus Scholtz & Howden 1987
Pereiraidium
Phycocus Broun 1883
Pinacopodius Branco 1992
Pleuronitis Van Lansberge 1875
Proteroscarabaeus Grabau 1923
Pseudeuryomia Kraatz 1894
Pseudoapterogyna Martínez De La Escalera 1914
Pycnopanelus Arrow 1931
Rhinacosmus Kraatz 1895
Scarabaeus
Scybalophagus
Silvinha
Sisyphus Latreille 1807
Stenovalgus Kolbe 1892
Sukelus Branco 1992
Thaumastopeus Kraatz 1885
Tiniocellus Péringuey 1902
Trichiomorphus Bourgoin 1919
Xenocanthon Martínez 1952
258 additional siblings truncated for brevity.
See the resource file for a full list.

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Scarabaeidae
Litocopris Waterhouse 1891 (this page)
Litocopris punctiventris Waterhouse 1891
Litocopris ruandanus Balthasar 1961
Litocopris simplex (Harold 1868)
Acanthonitis Janssens 1937
Afrodrepanus Krikken 2009
Agamopus Bates 1887
Amietina Cambefort 1981
Anoctus Sharp 1875
Anonychonitis Janssens 1950
Anoplosiagum Blanchard 1850
Aptenocanthon Matthews 1974
Aptychonitis Janssens 1937
Ateuchus Weber 1801
Attavicinus Philips & Bell 2008
Ausmontins Deschodt & Davis 2018
Baloghonthobium Paulian 1986
Bohepilissus Paulian 1975
Bolbites Harold 1868
Caccobiomorphus Balthasar 1964
Caccobius Thomson 1859
Caeconthobium Paulian 1984
Cambefortius Branco 1989
Canthochilum Chapin 1934
Canthonosoma Mac Leay 1871
Catharsiocopris Balthasar 1967
Cephalodesmius Westwood 1842
Cetonischema Reitter 1899
Cheironitis Lansberge 1875
Chiron Macleay 1819
Clypeodrepanus Krikken 2009
Copridaspidus Boucomont 1920
Coprophanaeus D' Olsoufieff 1924
Coptorhina Hope 1833
Ctilocephala Burmeister 1855
Degallieridium
Deltepilissus Silvério Pereira 1949
Dendropaemon Perty 1830
Diathermus Kraatz 1897
Dicranocara Frolov & Scholtz 2003
Dicronorhina Hope 1837
Dorbignyolus Branco 1989
Ennearabdus Van Lansberge 1874
Eucetonia Schoch 1894
Euglypta Mohnike 1873
Eusaproecius Branco 1989
Feeridium Vaz de Mello 2008
Garreta Janssens 1940
Glaphyrocanthon Martínez 1948
Glyphoderus Westwood 1837
Haematonotus Kraatz 1880
Hammondantus Cambefort 1978
Heterorrhina
Holocanthon Martínez & Pereira 1956
Hybosorites Nikolajev 1996
Lachnosterna Hope 1837
Leotrichillum
Lithoscarabaeus Nikolajev 1992
Lophorrhinides
Macroderes Westwood 1842
Macrophylla Hope 1837
Mecinonota Kraatz 1892
Megalorhina Westwood 1847
Melanocanthon Halffter 1958
Mentophilus Laporte de Castelnau 1840
Metacatharsius Paulian 1939
Micrelaphinis Schoch 1896
Miltotrogus Reitter 1902
Mimeoma Casey 1915
Mimonthophagus Balthasar 1963
Mnematium Macleay 1821
Nebulasilvius Deschodt & Scholtz 2008
Neosaproecius Branco 1990
Neptunides Thomson 1879
Nesosisyphus Vinson 1946
Nesovinsonia Martínez & Pereira 1959
Nunoidium
Onychothecus Boucomont 1912
Oxycetonia Arrow 1910
Oxygrylius Casey 1915
Pachylomera Kirby 1832
Pachysoma Macleay 1821
Paroniticellus Balthasar 1963
Parvuhowdenius Deschodt & Scholtz 2008
Peckolus Scholtz & Howden 1987
Pereiraidium
Phycocus Broun 1883
Pinacopodius Branco 1992
Pleuronitis Van Lansberge 1875
Proteroscarabaeus Grabau 1923
Pseudeuryomia Kraatz 1894
Pseudoapterogyna Martínez De La Escalera 1914
Pycnopanelus Arrow 1931
Rhinacosmus Kraatz 1895
Scarabaeus
Scybalophagus
Silvinha
Sisyphus Latreille 1807
Stenovalgus Kolbe 1892
Sukelus Branco 1992
Thaumastopeus Kraatz 1885
Tiniocellus Péringuey 1902
Trichiomorphus Bourgoin 1919
Xenocanthon Martínez 1952
258 additional siblings truncated for brevity.
See the resource file for a full list.

Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Coleoptera Linnaeus 1758
Polyphaga Emery 1886
Scarabeiformia Crowson 1960
Scarabaeoidea Latreille 1802
Scarabaeidae Latreille 1802
Scarabaeinae Latreille 1802
Coprini Leach 1815
Copris Geoffroy 1762 (this page)
Copris arizonensis Schaeffer 1906
Copris fricator (Fabricius 1787)
Copris gopheri Hubbard 1894
Copris howdeni Matthews & Halffter 1959
Copris incertus Say 1835
Copris inemarginatus Blatchley 1918
Copris klugi Harold 1869
Copris laeviceps Harold 1862
Copris lecontei Matthews 1962
Copris lugubris Boheman 1858
Copris macclevei Warner 1990
Copris martinezi Matthews & Halffter 1968
Copris minutus (Drury 1773)
Copris moechus Le Conte 1854
Copris remotus Le Conte 1866
Canthidium Erichson 1847
Dichotomius Hope 1838
Ontherus Erichson 1847
Uroxys Westwood 1842

NCBI

root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Scarabaeinae
Coprini
Copris (this page)
Copris aeneus
Copris agnus
Copris amyntor
Copris confucius
Copris cornifrons
Copris fidius
Copris hispanicus
Copris jacchoides
Copris laeviceps
Copris lugubris
Copris lunaris
Copris minutus
Copris ochus
Copris ramosiceps
Copris sinicus
Copris sonensis
Copris tripartitus
unclassified Copris
Coptodactyla
Homocopris
Litocopris

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Scarabaeinae
Coprini
Copris
Copris (this page)
Copris crassus
Copris davidis
Copris susanae
Copris algol
Copris amabilis
Copris andrewesi
Copris angustus
Copris anomiopseoides
Copris antares
Copris apicepunctatus
Copris arimotoi
Copris basilewskyi
Copris bihamatus
Copris boucardi
Copris brachypterus
Copris brittoni
Copris cambeforti
Copris cariniceps
Copris cassius
Copris celebensis
Copris colmanti
Copris complexus
Copris compressipennis
Copris confucius
Copris coriarius
Copris cornifrons
Copris corniger
Copris corpulentus
Copris costaricensis
Copris cribratus
Copris cribricollis
Copris cyaneus
Copris davisi
Copris davisoni
Copris decellei
Copris delicatus
Copris denticulatus
Copris diversus
Copris doriae
Copris draco
Copris dracunculus
Copris dudleyi
Copris eburneus
Copris elpheneroides
Copris elphenor
Copris ensifer
Copris excisus
Copris fallaciosus
Copris fujiokai
Copris furciceps
Copris garambae
Copris gazellarum
Copris glabricollis
Copris halffteri
Copris hatayamai
Copris hidakai
Copris hispanus
Copris howdeni
Copris igualensis
Copris imitans
Copris inemarginatus
Copris jacchoides
Copris jahi
Copris jucundus
Copris kasagii
Copris keralensis
Copris laiiformis
Copris laioides
Copris lecontei
Copris macclevei
Copris maesi
Copris martinae
Copris martinezi
Copris mexicanus
Copris morphaeus
Copris novaki
Copris pauliani
Copris pedarioides
Copris platycerus
Copris potanini
Copris pseudobootes
Copris punctiventris
Copris punctulatus
Copris quadridens
Copris reflexus
Copris renwarti
Copris ruandanus
Copris rugosus
Copris sanctus
Copris sarpedon
Copris serius
Copris simulator
Copris singularis
Copris smaragdifer
Copris subpunctatus
Copris uenoi
Copris vankhaii
Copris vietnamicus
Copris vilhenai
Copris viridis
Copris vrydaghi
Copris wiesei
Copris youngai
171 additional siblings truncated for brevity.
See the resource file for a full list.

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Polyphaga
Scarabaeiformia
Scarabaeoidea
Scarabaeidae
Scarabaeinae
Coprini
Copris (this page)
Copris
Copris acutidens
Copris afgoi
Copris agnus
Copris algol
Copris amabilis
Copris amyntor
Copris anceus
Copris andrewesi
Copris angusticornis
Copris angustus
Copris anomiopseoides
Copris antares
Copris apicepunctatus
Copris arcturus
Copris ares
Copris arimotoi
Copris arizonensis
Copris armatus
Copris armeniacus
Copris armiger
Copris arrowi
Copris aspericollis
Copris atropolitus
Copris barclayi
Copris basilewskyi
Copris basipunctatus
Copris bellator
Copris bengalensis
Copris bidens
Copris bihamatus
Copris bootes
Copris boucardi
Copris bovinus
Copris brachypterus
Copris brittoni
Copris caelatus
Copris caliginosus
Copris cambeforti
Copris cambodiensis
Copris camerunus
Copris caobangensis
Copris capensis
Copris cariniceps
Copris carinicus
Copris carmelita
Copris cassius
Copris celebensis
Copris cheni
Copris chimalapensis
Copris colmanti
Copris complexus
Copris compressipennis
Copris confucius
Copris coriarius
Copris cornifrons
Copris corniger
Copris corpulentus
Copris costaricensis
Copris cribratus
Copris cribricollis
Copris curlettii
Copris cyaneus
Copris davisi
Copris davisoni
Copris decellei
Copris delicatus
Copris denticulatus
Copris diversus
Copris doriae
Copris draco
Copris dracunculus
Copris dudleyi
Copris eburneus
Copris elpheneroides
Copris elphenor
Copris ensifer
Copris erratus
Copris evanidus
Copris excisus
Copris fallaciosus
Copris fallax
Copris felschei
Copris fidius
Copris frankenbergerianus
Copris freyi
Copris fricator
Copris fujiokai
Copris fukiensis
Copris furciceps
Copris garambae
Copris gazellarum
Copris gibbulus
Copris gilleti
Copris glabricollis
Copris gladiator
Copris gopheri
Copris gracilis
Copris graueri
Copris halffteri
Copris harrisi
Copris hartli
Copris hatayamai
Copris hidakai
Copris hispanus
Copris hosakai
Copris howdeni
Copris humilis
Copris hybridus
Copris igualensis
Copris illotus
Copris imitans
Copris inaequabilis
Copris incertus
Copris inemarginatus
Copris inhalatus
Copris insidiosus
Copris integer
Copris interioris
Copris iris
Copris jacchoides
Copris jacchus
Copris jahi
Copris johkii
Copris juanmorai
Copris jucundus
Copris kachinensis
Copris kasagii
Copris kasanka
Copris katangae
Copris keralensis
Copris kiuchii
Copris klapperichi
Copris klugi
Copris kusakabei
Copris laeviceps
Copris laevigatus
Copris laiiformis
Copris laioides
Copris laius
Copris lannathai
Copris lecontei
Copris lugubris
Copris lunaris
Copris macacus
Copris macclevei
Copris macer
Copris maesi
Copris magicus
Copris manni
Copris marcus
Copris martinae
Copris martinezi
Copris matthewsi
Copris medogensis
Copris megaceratoides
Copris megasoma
Copris mesacanthus
Copris mexicanus
Copris mindorensis
Copris minutissimus
Copris minutus
Copris misellus
Copris miyakei
Copris moechus
Copris moffartsi
Copris mongkhoni
Copris montivagus
Copris morphaeus
Copris mourgliai
Copris muticus
Copris neglectus
Copris nepos
Copris nevinsoni
Copris novaki
Copris nubilosus
Copris numa
Copris obenbergeri
Copris obesus
Copris ochus
Copris orion
Copris orphanus
Copris parapecuarius
Copris pauliani
Copris pecuarius
Copris pedarioides
Copris phungae
Copris phylax
Copris platycerus
Copris poggii
Copris potanini
Copris prevosti
Copris propinquus
Copris pseudobootes
Copris pseudochus
Copris pseudomoffartsi
Copris pseudosinicus
Copris pueli
Copris punctatus
Copris puncticollis
Copris punctipennis
Copris punctiventris
Copris punctulatus
Copris punjabensis
Copris quadridens
Copris quasilaevigatus
Copris ramosiceps
Copris rebouchei
Copris reflexus
Copris remotus
Copris renwarti
Copris repertus
Copris ribbei
Copris riekoae
Copris ritsemae
Copris ruandanus
Copris rugosus
Copris ruricola
Copris sabinus
Copris sacontala
Copris sallei
Copris sanctus
Copris sarpedon
Copris saundersi
Copris schoolmeestersi
Copris scorpio
Copris serius
Copris sexdentatus
Copris siamensis
Copris siangensis
Copris sierrensis
Copris signatus
Copris simonettai
Copris simplex
Copris simulator
Copris singularis
Copris sinicus
Copris sinon
Copris smaragdifer
Copris sodalis
Copris sorex
Copris subdolus
Copris subpunctatus
Copris subsidens
Copris subterraneus
Copris sumatrensis
Copris surdus
Copris szechouanicus
Copris tetraodon
Copris tridens
Copris tridentatus
Copris tripartitus
Copris truncatus
Copris tsukamotoi
Copris typhoeus
Copris uenoi
Copris umbilicatus
Copris urus
Copris usambaricus
Copris vankhaii
Copris victorini
Copris vietnamicus
Copris vilhenai
Copris viridis
Copris vitalisi
Copris vrydaghi
Copris warneri
Copris wiesei
Copris yangi
Copris youngai
Copris zhangi
Sinocopris
Canthidium
Catharsius
Dichotomius
Heliocopris
Litocopris
Ontherus
Synapsis
EOL is hosted by: