Aegialia crassa Le Conte 1857

Aegialia (Aegialia) crassa LeConte

Aegialia crassa LeConte, 1860:42—Horn, 1871:294—Brown, 1931:48—Hatch, 1971:442: Stebnicka, 1977:463—Gordon and Cartwright, 1977:43

Aegialia crassa insularis Brown, 1931:49—Stebnicka, 1977:463—Gordon and Cartwright, 1977:43

Aegialia crassa var insularis—Hatch, 1971:442

DESCRIPTION—Length 40 to 50 mm, greatest width 190 to 260 mm Form oval, convex, very broad posteriorly (Figure 15) Color black except anterior leg, each femur reddish black Head densely granulate, impunctate; clypeal apex feebly emarginate medially, gena distinctly produced Pronotum slightly alutaceous, punctations coarse basomedially, becoming fine, indistinct anteriorly, laterally, or fine basally with apical, lateral areas impunctate; base without marginal line Elytron feebly alutaceous, interval weakly convex with barely perceptible, fine punctations, stria weakly impressed, feebly punctate, punctations separated by about a diameter Metasternum smooth with 2 to 4 coarse punctations medially, heavily alutaceous laterally Wings reduced, probably not functional Middle tibia short, robust, surface denticles present in apical , 2 incomplete transverse carinae present, apical spurs slender, outer spur as long as first 3 tarsal segments, inner spur as long as first 4 tarsal segments Hind femur with apical flange barely evident; hindtibia robust, wide, with numerous surface denticles and 2 rows of denticles on inner margin from base to apex, one incomplete carina in apical , apical spurs strongly spatulate, both spurs slightly shorter than first 3 tarsal segments combined Male genitalia as in Figure 49; female genitalia plates as figured by Stebnicka (1977, fig 88)

LECTOTYPE—Designated herein It is unclear from the original description how many type specimens LeConte had Therefore we designate a female from the LeConte collection labeled “[gold disc]/TYPE 3724 [red paper]/J LeConte Collection” as the lectotype There are 5 other specimens in the series, 2 of which bear gold discs The latter 2 specimens may also be type material

TYPE LOCALITY—San Francisco, California (lectotype here designated)

TYPE DEPOSITORY—MCZ

REMARKS—Aegialia crassa is a common species on some of the coastal dune systems of the Pacific Coast and is readily distinguished from those species lacking a basal pronotal line by the reduced pronotal punctation and extremely short, robust, middle and hindtibae

DISTRIBUTION—Southeastern British Columbia to Baja California

Aegialia (Aegialia) magnifica Gordon and Cartwright

Aegialia (Aegialia) magnifica Gordon and Cartwright, 1977:43

DESCRIPTION—Length 440 to 590 mm, greatest width 248 to 325 mm Form oval, convex, very broad posteriorly (Figure 16) Color pale red except mouthparts and ventral surface yellowish red Head smooth, shiny, finely alutaceous, fine punctations separated by a diameter or less, becoming coarser, denser at anterolateral angle; clypeal apex feebly emarginate, gena slightly produced Pronotum smooth, shiny, fine punctations separated by less than to 3 times a diameter; base with marginal line Elytron smooth, shiny, interval nearly flat, finely punctate, punctations separated by twice a diameter, stria feebly impressed, finely punctate, punctations separated by twice a diameter Metasternum polished, impunctate medially, coarsely punctate and alutaceous laterally Middle tibia with 2 transverse carinae, lacking surface denticles, apical spurs slender, outer spur as long as first 3 tarsal segments, inner spur as long as first 4 tarsal segments; hindfemur densely, coarsely punctate, apical flange produced, inner angle obsolete; hindtibia robust, wide, a row of denticles present on inner margin from base to apex, with incomplete transverse carina in basal third, complete carina in apical (Figure 71), apical spurs somewhat spatulate, inner spur as long as first 2 tarsal segments, outer spur as long as first 3 tarsal segments (Figure 72) Male genitalia as in Figure 25; female genital plate as in Figure 28

TYPE LOCALITY—Nevada, Nye Co, “Big Dune,” Amargosa Desert

TYPE DEPOSITORY—USNM

REMARKS—This large, red, smooth surfaced species is the most distinctive member of the North American aegialiine fauna and is thus far known only from 2 inland dunes in Nevada

DISTRIBUTION—Nevada: type locality and Lava Dune, NW Lathrop Wells

Aegialia (Aegialia) crescenta Gordon and Cartwright

DESCRIPTION—Length 375 to 50 mm, greatest width 205 to 270 mm Form convex, oval (Figure 17) Color dark reddish brown except ventral surface, legs, and mouthparts pale yellowish brown Head shiny, sometimes feebly alutaceous, with band of coarse, nearly contiguous punctations from basal portion of clypeus to vertex, apical portion of clypeus less coarsely punctate; clypeal apex feebly emarginate medially Pronotum smooth, shiny, disc usually with indistinct, feebly impressed furrow extending from base nearly to anterior margin, midline usually narrowly impunctate, rest of pronotum with coarse, sparse punctations separated by less than to 3 times a diameter; base with marginal line Elytron smooth, shiny, interval flat, punctations on interval in single, slightly irregular, median row, separated by less than to 3 times a diameter, stria distinctly impressed, coarsely punctate, punctations separated by a diameter Metasternum densely, finely punctate medially, intermixed fine, coarse punctations laterally Functional wings absent Middle tibia with 2 transverse carinae, lacking surface denticles, apical spurs as long as first 3 tarsal segments; hindfemur sparsely, coarsely punctate, apical flange slightly produced, inner angle obsolete; hindtibia robust, wide, surface denticles in basal third, with incomplete transverse carina in basal third, complete carina in apical (Figure 73), apical spurs somewhat spatulate, inner spur as long as first 2 tarsal segments, outer spur as long as first 2½ tarsal segments (Figure 74) Male genitalia as in Figure 26 Female genitalia plate as in Figure 29

TYPE LOCALITY—Nevada, Nye Co, Crescent Dune

TYPE DEPOSITORY—USNM

REMARKS—This is another inland dune species from Nevada, which is closely related to A hardyi but not similar to any other described species See “Remarks” under A hardyi

DISTRIBUTION—Nevada: type locality New Record: Nevada: Clark Co, Game Range Dunes

Aegialia (Aegialia) hardyi Gordon and Cartwright

Aegialia (Aegialia) hardyi Gordon and Cartwright, 1977:47

DESCRIPTION—Length 333 to 552 mm, greatest width 175 to 180 mm (Figure 18) Description as for A crescenta except differences noted as follows: head with vertex punctate; pronotal punctations coarse, dense, separated by a diameter or less, punctations present on midline, basal marginal line not visible in dorsal view; hindfemur with scattered, fine punctations; hindtibia relatively slender, hindtibia, spurs as in Figures 75, 76 Male genitalia as in Figure 27; female genital plate as in Figure 30

TYPE LOCALITY—Nevada, Churchill Co, Sand Mountain Dunes

TYPE DEPOSITORY—USNM

REMARKS—The similarity between A hardyi and A crescenta is striking; however, the differences listed above seem to be constant The slender hindtibia and finely punctate hindfemur of A hardyi are significantly different from the robust, wide hindtibia and coarsely punctate femur of A crescenta

DISTRIBUTION—Nevada: type locality

Aegialia (Aegialia) concinna Gordon and Cartwright

Aegialia (Aegialia) concinna Gordon and Cartwright, 1977:48

DESCRIPTION—Length 325 to 40 mm, greatest width 170 to 20 mm Form convex, elongate oval (Figure 19) Color pale brownish yellow to reddish brown except ventral surface always paler than dorsum Head mostly granulate with some close, coarse punctations, vertex smooth; clypeus barely perceptibly emarginate medially Pronotum smooth, shiny, coarsely punctate, punctations separated by a diameter or less; base with marginal line Elytron smooth, shiny, interval convex with fine punctations arranged in single, irregular row, stria deeply impressed, coarsely punctate, punctations separated by twice a diameter Metasternum smooth, shiny medially, alutaceous with coarse punctations laterally Functional wings absent Middle tibia strongly enlarged apically with 2 transverse carinae, lacking surface denticles except in basal third, apical spurs slender, outer spur as long as first 2 tarsal segments, inner spur as long as first 3 tarsal segments; hindfemur with apical flange slightly produced, inner angle obsolete; hindtibia slender, short, row of surface denticles present in basal third near inner margin, with carina in basal third nearly obsolete, carina in apical complete, lacking surface denticles, apical spurs slender, somewhat spatulate, inner spur as long as first 2 tarsal segments, outer spur as long as first 3 tarsal segments Male genitalia as in Figure 50; female genital plate as in Figure 51

TYPE LOCALITY—California, Fresno Co, 18 mi (29 km) SW Mendota

TYPE DEPOSITORY—USNM

REMARKS—This small, pale species is quite unlike any of the other inland dune species to which it is most similar, and even more unlike the remainder of the Aegialia, sensu stricto, fauna When it was described by Gordon and Cartwright in 1977, only 4 specimens, all females, were available Since then, many more specimens have been collected

DISTRIBUTION—California: type locality New Records: California: Contra Costa Co, Antioch; Fresno Co, Monocline Ridge; San Benito Co, Panoche Rd