From Cupello, M. & Vaz-de-Mello (2013) Taxonomic revision of the South American dung beetle genus Gromphas Brull, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa, 3722 (4), 439-482.
From Cupello, M. & Vaz-de-Mello (2013) Taxonomic revision of the South American dung beetle genus Gromphas Brull, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa, 3722 (4), 439-482.
From Cupello, M. & Vaz-de-Mello (2013) Taxonomic revision of the South American dung beetle genus Gromphas Brull, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa, 3722 (4), 439-482.
From Cupello, M. & Vaz-de-Mello (2013) Taxonomic revision of the South American dung beetle genus Gromphas Brull, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa, 3722 (4), 439-482.
From Cupello, M. & Vaz-de-Mello (2013) Taxonomic revision of the South American dung beetle genus Gromphas Brull, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa, 3722 (4), 439-482.
From Cupello, M. & Vaz-de-Mello (2013) Taxonomic revision of the South American dung beetle genus Gromphas Brull, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa, 3722 (4), 439-482.