dcsimg
 
  en  
names in breadcrumbs
Image of Pterostichus (Hypherpes) herculaneus Mannerheim 1843
Creatures » … » Animal » … » Arthropods » … » Hexapods » Insects » Winged Insects » … » Endopterygotes » Beetles » … » Ground Beetles » …

Pterostichus (Hypherpes) herculaneus Mannerheim 1843

Scientific Names

Preferred Names

Pterostichus (Hypherpes) herculaneus Mannerheim 1843
Species recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus herculaneus
Species recognized by wikipedia NL, Canada Species List, NCBI, and United States Species List
Pterostichus herculaneus Mannerheim 1843
Species recognized by GBIF classification
Pterostichus (Hypherpes) herculaneus Mannerheim 1843
Species recognized by EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus (Hypherpes) herculaneus Mannerheim 1843
Species recognized by Catalogue of Life 26 March 2024, Dynamic Hierarchy Version 2.2.3, and EOL Dynamic Hierarchy Version 2.2.4
Pterostichus (Hypherpes) scenicus Casey
Recognized by NMNH type specimens
Pterostichus herculaneus
Species recognized by wikidata_hierarchy tar gz, wikipedia EN, Barcode of Life Data Systems, iNaturalist data coverage, and GGBN data coverage
Pterostichus herculaneus
Recognized by BHL data coverage
Pterostichus herculaneus Mannerheim 1843
Species recognized by iNaturalist, Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy, TreatmentBank, and GBIF data coverage

Alternative Names

Hypherpes herculaneus (Mannerheim 1843)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus scenicus Casey 1913
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Hypherpes herculaneus (Mannerheim 1843)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus scenicus Casey 1913
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Hypherpes herculaneus (Mannerheim 1843)
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Hypherpes herculaneus (Mannerheim 1843)
Synonym according to Catalogue of Life 26 March 2024
Pterostichus scenicus Casey 1913
Not accepted according to Dynamic Hierarchy Version 2.2.3 and EOL Dynamic Hierarchy Version 2.2.4
Pterostichus scenicus Casey 1913
Synonym according to Catalogue of Life 26 March 2024

Common Names

There are no common names associated with this taxon.

Curated hierarchies for Pterostichus (Hypherpes) herculaneus Mannerheim 1843

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Pterostichus
Pterostichus (Hypherpes) herculaneus Mannerheim 1843 (this page)
Pterostichus (Agastillus) eriwanicus (Tschitscherine 1897)
Pterostichus (Aphaonus) arcanoides (Lorenz 1998)
Pterostichus (Aphaonus) cylindriformis (Reitter 1887)
Pterostichus (Argutor) cursor (Dejean 1828)
Pterostichus (Badistrinus) goschi Jedlicka 1930
Pterostichus (Bothriopterus) ferghanicus Kabak 2001
Pterostichus (Bothriopterus) oblongopunctatus (Fabricius 1787)
Pterostichus (Bothriopterus) oregonus Le Conte 1861
Pterostichus (Bothriopterus) pensylvanicus Le Conte 1873
Pterostichus (Bothriopterus) zyzzovi O. & E. Berlov 1996
Pterostichus (Circinatus) agilis Allegro & Sciaky 2010
Pterostichus (Cryobius) arcticola (Chaudoir 1868)
Pterostichus (Cryobius) asiaticus (Poppius 1906)
Pterostichus (Cryobius) glacialis (Brisout de Barneville 1863)
Pterostichus (Cryobius) korgei Jedlicka 1964
Pterostichus (Cryobius) kultianus Jedlicka 1946
Pterostichus (Cryobius) mandibularoides Ball 1966
Pterostichus (Cryobius) palekhai E. & O. Berlov 1996
Pterostichus (Cryobius) parasimilis Ball 1962
Pterostichus (Cryobius) procerulus (Heyden 1880)
Pterostichus (Cryobius) tatricus Kult 1947
Pterostichus (Eosteropus) abudarini O. & E. Berlov 1997
Pterostichus (Eosteropus) tokui Morita 2007
Pterostichus (Epinialoe) janoi Jedlicka 1952
Pterostichus (Ethira) sharanus (Straneo 1984)
Pterostichus (Eurythoracana) haptoderoides (Tschitscherine 1889)
Pterostichus (Falsargutor) pseudopedius Reitter 1887
Pterostichus (Gastrosticta) obesulus Le Conte 1873
Pterostichus (Gastrosticta) putus Casey 1913
Pterostichus (Haplomaseus) andreae (Tschitscherine 1897)
Pterostichus (Haplomaseus) armenus (Faldermann 1836)
Pterostichus (Haplomaseus) caucasicus (Ménétriés 1832)
Pterostichus (Haptotapinus) vseteckai (Maran 1944)
Pterostichus (Hypherpes) annosus Casey 1913
Pterostichus (Hypherpes) ecarinatus Hatch 1936
Pterostichus (Hypherpes) scutellaris Le Conte 1873
Pterostichus (Hypherpes) setosus Hatch 1951
Pterostichus (Koreonialoe) teretis Park & Kwon 1996
Pterostichus (Lenapterus) agonus G. Horn 1880
Pterostichus (Lenapterus) punctatissimus (Randall 1838)
Pterostichus (Leptoferonia) falli Van Dyke 1926
Pterostichus (Leptoferonia) hatchi Hacker 1968
Pterostichus (Leptoferonia) infernalis Hatch 1936
Pterostichus (Leptoferonia) stapedius Hacker 1968
Pterostichus (Leptoferonia) yosemitensis Hacker 1968
Pterostichus (Lianoe) drescoi Nègre 1957
Pterostichus (Lianoe) tanakai Ishida 1964
Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida 1972
Pterostichus (Melanius) aterrimus (Herbst 1784)
Pterostichus (Metallophilus) rugosus (Gebler 1823)
Pterostichus (Morphohaptoderus) hubeicus Facchini & Sciaky 2003
Pterostichus (Morphohaptoderus) yulongshanensis Sciaky 1997
Pterostichus (Neohaptoderus) jureceki (Jedlicka 1936)
Pterostichus (Neohaptoderus) yunnanensis Jedlicka 1934
Pterostichus (Nialoe) ohbayashii Ishida 1968
Pterostichus (Nialoe) okutamae Tanaka 1963
Pterostichus (Nialoe) opacipennis Jedlicka 1934
Pterostichus (Nialoe) shinbodakensis Sasakawa & H. Ito 2017
Pterostichus (Nialoe) taoi Kasahara 1993
Pterostichus (Nialoe) yakushimanus Nakane & Ishida 1961
Pterostichus (Oreophilus) morio (Duftschmid 1812)
Pterostichus (Oreophilus) spinolae (Dejean 1828)
Pterostichus (Oreoplatysma) abishirensis Belousov 1991
Pterostichus (Oreoplatysma) casaleianus Kirschenhofer 1991
Pterostichus (Oreoplatysma) kvirensis Belousov 1991
Pterostichus (Oreoplatysma) mediocris Dvorak 1995
Pterostichus (Oreoplatysma) orman Jedlicka 1963
Pterostichus (Oreoplatysma) rousianus Kirschenhofer 1987
Pterostichus (Oreoplatysma) satyrus Kurnakov 1962
Pterostichus (Paranialoe) biexcisus Straneo 1955
Pterostichus (Petrophilus) alexandrovi Lafer 1979
Pterostichus (Petrophilus) arsenjevi Lafer 1979
Pterostichus (Petrophilus) pertinax (Tschitscherine 1895)
Pterostichus (Petrophilus) tomensis (Gebler 1847)
Pterostichus (Phonias) diligens (Sturm 1824)
Pterostichus (Phonias) gyrosus (Motschulsky 1866)
Pterostichus (Platysma) sachtlebeni Jedlicka 1962
Pterostichus (Pseudethira) gobettii Straneo 1983
Pterostichus (Pseudethira) immarginatus Straneo 1977
Pterostichus (Pseudethira) matsumurai Habu 1973
Pterostichus (Pseudethira) miehei J. Schmidt 2005
Pterostichus (Pseudethira) weiperti J. Schmidt 2005
Pterostichus (Pseudomaseus) aimaki Jedlicka 1968
Pterostichus (Pseudomaseus) rhaeticus Heer 1837
Pterostichus (Pseudomaseus) tenuis (Casey 1924)
Pterostichus (Pterostichus) auratus Heer 1837
Pterostichus (Pterostichus) devillei Puel 1924
Pterostichus (Pterostichus) esakii Ishida 1959
Pterostichus (Pterostichus) pedemontanus Ganglbauer 1891
Pterostichus (Pterostichus) ruffoi Sciaky 1984
Pterostichus (Rhagadus) modicellus (Tschitscherine 1897)
Pterostichus (Sinosteropus) jani Sciaky & Facchini 2003
Pterostichus (Steropus) catalonicus J. Daniel ex K. & J. Daniel 1906
Pterostichus (Tschitscherinea) mulensis Sciaky 1997
Pterostichus bischoffianus Jedlicka 1936
Pterostichus exochus Andrewes 1930
Pterostichus nishidai Nakane 1989
Pterostichus pentheri Apfelbeck 1918
Pterostichus punctisternus Straneo 1989
Pterostichus umbralis Gistel 1857
1082 additional siblings truncated for brevity.
See the resource file for a full list.

Catalogue of Life 26 March 2024

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Caraboidea
Carabidae Latreille 1802
Pterostichinae Bonelli 1810
Pterostichini Bonelli 1810
Pterostichina Bonelli 1810
Pterostichus Bonelli 1810
Pterostichus subgen. Hypherpes Chaudoir 1838
Pterostichus (Hypherpes) herculaneus Mannerheim 1843 (this page)
Pterostichus (Hypherpes) adoxus (Say 1823)
Pterostichus (Hypherpes) algidus Le Conte 1853
Pterostichus (Hypherpes) amethystinus Mannerheim 1843
Pterostichus (Hypherpes) annosus Casey 1913
Pterostichus (Hypherpes) arcanus Casey 1913
Pterostichus (Hypherpes) baldwini (Casey 1924)
Pterostichus (Hypherpes) barbarinus Casey 1913
Pterostichus (Hypherpes) brachylobus Kavanaugh & La Bonte 2006
Pterostichus (Hypherpes) californicus (Dejean 1828)
Pterostichus (Hypherpes) canallatus Casey 1913
Pterostichus (Hypherpes) castaneus (Dejean 1828)
Pterostichus (Hypherpes) castanipes (Ménétriés 1843)
Pterostichus (Hypherpes) congestus (Ménétriés 1843)
Pterostichus (Hypherpes) craterensis (Hatch 1949)
Pterostichus (Hypherpes) crenicollis Le Conte 1873
Pterostichus (Hypherpes) ecarinatus Hatch 1936
Pterostichus (Hypherpes) esuriens Casey 1913
Pterostichus (Hypherpes) gliscans Casey 1913
Pterostichus (Hypherpes) gracilior Le Conte 1873
Pterostichus (Hypherpes) hornii Le Conte 1873
Pterostichus (Hypherpes) illustris Le Conte 1851
Pterostichus (Hypherpes) inermis Fall 1901
Pterostichus (Hypherpes) isabellae Le Conte 1851
Pterostichus (Hypherpes) jacobinus Casey 1913
Pterostichus (Hypherpes) laborans Casey 1913
Pterostichus (Hypherpes) lacertus Casey 1913
Pterostichus (Hypherpes) lama (Ménétriés 1843)
Pterostichus (Hypherpes) lassulus (Casey 1920)
Pterostichus (Hypherpes) lattini La Bonte 2006
Pterostichus (Hypherpes) luscus (Casey 1918)
Pterostichus (Hypherpes) menetriesii Le Conte 1873
Pterostichus (Hypherpes) mercedianus (Casey 1918)
Pterostichus (Hypherpes) miscellus Casey 1913
Pterostichus (Hypherpes) morionides (Chaudoir 1868)
Pterostichus (Hypherpes) neobrunneus Lindroth 1966
Pterostichus (Hypherpes) nigrocaeruleus Van Dyke 1926
Pterostichus (Hypherpes) obsidianus Casey 1913
Pterostichus (Hypherpes) occultus Casey 1913
Pterostichus (Hypherpes) ordinarius Casey 1913
Pterostichus (Hypherpes) ovalipennis Casey 1913
Pterostichus (Hypherpes) panticulatus Casey 1913
Pterostichus (Hypherpes) pergracilis (Casey 1920)
Pterostichus (Hypherpes) planctus Le Conte 1853
Pterostichus (Hypherpes) protensiformis (Casey 1924)
Pterostichus (Hypherpes) protractus Le Conte 1860
Pterostichus (Hypherpes) restrictus (Casey 1918)
Pterostichus (Hypherpes) scutellaris Le Conte 1873
Pterostichus (Hypherpes) sejungendus (Chaudoir 1868)
Pterostichus (Hypherpes) serripes (Le Conte 1875)
Pterostichus (Hypherpes) setosus Hatch 1951
Pterostichus (Hypherpes) sierranus Casey 1913
Pterostichus (Hypherpes) sponsor Casey 1913
Pterostichus (Hypherpes) spraguei Le Conte 1873
Pterostichus (Hypherpes) suffusus Casey 1913
Pterostichus (Hypherpes) tarsalis Le Conte 1873
Pterostichus (Hypherpes) tristis (Dejean 1828)
Pterostichus (Hypherpes) tuberculofemoratus Hatch 1936
Pterostichus (Hypherpes) vandykei Schaeffer 1910
Pterostichus (Hypherpes) vicinus Mannerheim 1843
Pterostichus (Hypherpes) ybousqueti O. Berlov 1999

Dynamic Hierarchy Version 2.2.3

Pterostichus (Hypherpes) herculaneus Mannerheim 1843 (this page)
Pterostichus (Agastillus) eriwanicus (Tschitscherine 1897)
Pterostichus (Aphaonus) arcanoides (Lorenz 1998)
Pterostichus (Aphaonus) cylindriformis (Reitter 1887)
Pterostichus (Argutor) cursor (Dejean 1828)
Pterostichus (Badistrinus) goschi Jedlicka 1930
Pterostichus (Bothriopterus) ferghanicus Kabak 2001
Pterostichus (Bothriopterus) oblongopunctatus (Fabricius 1787)
Pterostichus (Bothriopterus) oregonus Le Conte 1861
Pterostichus (Bothriopterus) pensylvanicus Le Conte 1873
Pterostichus (Bothriopterus) zyzzovi O. & E. Berlov 1996
Pterostichus (Circinatus) agilis Allegro & Sciaky 2010
Pterostichus (Cryobius) arcticola (Chaudoir 1868)
Pterostichus (Cryobius) asiaticus (Poppius 1906)
Pterostichus (Cryobius) glacialis (Brisout de Barneville 1863)
Pterostichus (Cryobius) korgei Jedlicka 1964
Pterostichus (Cryobius) kultianus Jedlicka 1946
Pterostichus (Cryobius) mandibularoides Ball 1966
Pterostichus (Cryobius) palekhai E. & O. Berlov 1996
Pterostichus (Cryobius) parasimilis Ball 1962
Pterostichus (Cryobius) procerulus (Heyden 1880)
Pterostichus (Cryobius) tatricus Kult 1947
Pterostichus (Eosteropus) abudarini O. & E. Berlov 1997
Pterostichus (Eosteropus) tokui Morita 2007
Pterostichus (Epinialoe) janoi Jedlicka 1952
Pterostichus (Ethira) sharanus (Straneo 1984)
Pterostichus (Eurythoracana) haptoderoides (Tschitscherine 1889)
Pterostichus (Falsargutor) pseudopedius Reitter 1887
Pterostichus (Gastrosticta) obesulus Le Conte 1873
Pterostichus (Gastrosticta) putus Casey 1913
Pterostichus (Haplomaseus) andreae (Tschitscherine 1897)
Pterostichus (Haplomaseus) armenus (Faldermann 1836)
Pterostichus (Haplomaseus) caucasicus (Ménétriés 1832)
Pterostichus (Haptotapinus) vseteckai (Maran 1944)
Pterostichus (Hypherpes) annosus Casey 1913
Pterostichus (Hypherpes) ecarinatus Hatch 1936
Pterostichus (Hypherpes) scutellaris Le Conte 1873
Pterostichus (Hypherpes) setosus Hatch 1951
Pterostichus (Koreonialoe) teretis Park & Kwon 1996
Pterostichus (Lenapterus) agonus G. Horn 1880
Pterostichus (Lenapterus) punctatissimus (Randall 1838)
Pterostichus (Leptoferonia) falli Van Dyke 1926
Pterostichus (Leptoferonia) hatchi Hacker 1968
Pterostichus (Leptoferonia) infernalis Hatch 1936
Pterostichus (Leptoferonia) stapedius Hacker 1968
Pterostichus (Leptoferonia) yosemitensis Hacker 1968
Pterostichus (Lianoe) drescoi Nègre 1957
Pterostichus (Lianoe) tanakai Ishida 1964
Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida 1972
Pterostichus (Melanius) aterrimus (Herbst 1784)
Pterostichus (Metallophilus) rugosus (Gebler 1823)
Pterostichus (Morphohaptoderus) hubeicus Facchini & Sciaky 2003
Pterostichus (Morphohaptoderus) yulongshanensis Sciaky 1997
Pterostichus (Neohaptoderus) jureceki (Jedlicka 1936)
Pterostichus (Neohaptoderus) yunnanensis Jedlicka 1934
Pterostichus (Nialoe) ohbayashii Ishida 1968
Pterostichus (Nialoe) okutamae Tanaka 1963
Pterostichus (Nialoe) opacipennis Jedlicka 1934
Pterostichus (Nialoe) shinbodakensis Sasakawa & H. Ito 2017
Pterostichus (Nialoe) taoi Kasahara 1993
Pterostichus (Nialoe) yakushimanus Nakane & Ishida 1961
Pterostichus (Oreophilus) morio (Duftschmid 1812)
Pterostichus (Oreophilus) spinolae (Dejean 1828)
Pterostichus (Oreoplatysma) abishirensis Belousov 1991
Pterostichus (Oreoplatysma) casaleianus Kirschenhofer 1991
Pterostichus (Oreoplatysma) kvirensis Belousov 1991
Pterostichus (Oreoplatysma) mediocris Dvorak 1995
Pterostichus (Oreoplatysma) orman Jedlicka 1963
Pterostichus (Oreoplatysma) rousianus Kirschenhofer 1987
Pterostichus (Oreoplatysma) satyrus Kurnakov 1962
Pterostichus (Paranialoe) biexcisus Straneo 1955
Pterostichus (Petrophilus) alexandrovi Lafer 1979
Pterostichus (Petrophilus) arsenjevi Lafer 1979
Pterostichus (Petrophilus) pertinax (Tschitscherine 1895)
Pterostichus (Petrophilus) tomensis (Gebler 1847)
Pterostichus (Phonias) diligens (Sturm 1824)
Pterostichus (Phonias) gyrosus (Motschulsky 1866)
Pterostichus (Platysma) sachtlebeni Jedlicka 1962
Pterostichus (Pseudethira) gobettii Straneo 1983
Pterostichus (Pseudethira) immarginatus Straneo 1977
Pterostichus (Pseudethira) matsumurai Habu 1973
Pterostichus (Pseudethira) miehei J. Schmidt 2005
Pterostichus (Pseudethira) weiperti J. Schmidt 2005
Pterostichus (Pseudomaseus) aimaki Jedlicka 1968
Pterostichus (Pseudomaseus) rhaeticus Heer 1837
Pterostichus (Pseudomaseus) tenuis (Casey 1924)
Pterostichus (Pterostichus) auratus Heer 1837
Pterostichus (Pterostichus) devillei Puel 1924
Pterostichus (Pterostichus) esakii Ishida 1959
Pterostichus (Pterostichus) pedemontanus Ganglbauer 1891
Pterostichus (Pterostichus) ruffoi Sciaky 1984
Pterostichus (Rhagadus) modicellus (Tschitscherine 1897)
Pterostichus (Sinosteropus) jani Sciaky & Facchini 2003
Pterostichus (Steropus) catalonicus J. Daniel ex K. & J. Daniel 1906
Pterostichus (Tschitscherinea) mulensis Sciaky 1997
Pterostichus bischoffianus Jedlicka 1936
Pterostichus exochus Andrewes 1930
Pterostichus nishidai Nakane 1989
Pterostichus pentheri Apfelbeck 1918
Pterostichus punctisternus Straneo 1989
Pterostichus umbralis Gistel 1857
1082 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Pterostichus
Pterostichus (Hypherpes) herculaneus Mannerheim 1843 (this page)
Pterostichus (Agastillus) eriwanicus (Tschitscherine 1897)
Pterostichus (Aphaonus) arcanoides (Lorenz 1998)
Pterostichus (Aphaonus) cylindriformis (Reitter 1887)
Pterostichus (Argutor) cursor (Dejean 1828)
Pterostichus (Badistrinus) goschi Jedlicka 1930
Pterostichus (Bothriopterus) ferghanicus Kabak 2001
Pterostichus (Bothriopterus) oblongopunctatus (Fabricius 1787)
Pterostichus (Bothriopterus) oregonus Le Conte 1861
Pterostichus (Bothriopterus) pensylvanicus Le Conte 1873
Pterostichus (Bothriopterus) zyzzovi O. & E. Berlov 1996
Pterostichus (Circinatus) agilis Allegro & Sciaky 2010
Pterostichus (Cryobius) arcticola (Chaudoir 1868)
Pterostichus (Cryobius) asiaticus (Poppius 1906)
Pterostichus (Cryobius) glacialis (Brisout de Barneville 1863)
Pterostichus (Cryobius) korgei Jedlicka 1964
Pterostichus (Cryobius) kultianus Jedlicka 1946
Pterostichus (Cryobius) mandibularoides Ball 1966
Pterostichus (Cryobius) palekhai E. & O. Berlov 1996
Pterostichus (Cryobius) parasimilis Ball 1962
Pterostichus (Cryobius) procerulus (Heyden 1880)
Pterostichus (Cryobius) tatricus Kult 1947
Pterostichus (Eosteropus) abudarini O. & E. Berlov 1997
Pterostichus (Eosteropus) tokui Morita 2007
Pterostichus (Epinialoe) janoi Jedlicka 1952
Pterostichus (Ethira) sharanus (Straneo 1984)
Pterostichus (Eurythoracana) haptoderoides (Tschitscherine 1889)
Pterostichus (Falsargutor) pseudopedius Reitter 1887
Pterostichus (Gastrosticta) obesulus Le Conte 1873
Pterostichus (Gastrosticta) putus Casey 1913
Pterostichus (Haplomaseus) andreae (Tschitscherine 1897)
Pterostichus (Haplomaseus) armenus (Faldermann 1836)
Pterostichus (Haplomaseus) caucasicus (Ménétriés 1832)
Pterostichus (Haptotapinus) vseteckai (Maran 1944)
Pterostichus (Hypherpes) annosus Casey 1913
Pterostichus (Hypherpes) ecarinatus Hatch 1936
Pterostichus (Hypherpes) scutellaris Le Conte 1873
Pterostichus (Hypherpes) setosus Hatch 1951
Pterostichus (Koreonialoe) teretis Park & Kwon 1996
Pterostichus (Lenapterus) agonus G. Horn 1880
Pterostichus (Lenapterus) punctatissimus (Randall 1838)
Pterostichus (Leptoferonia) falli Van Dyke 1926
Pterostichus (Leptoferonia) hatchi Hacker 1968
Pterostichus (Leptoferonia) infernalis Hatch 1936
Pterostichus (Leptoferonia) stapedius Hacker 1968
Pterostichus (Leptoferonia) yosemitensis Hacker 1968
Pterostichus (Lianoe) drescoi Nègre 1957
Pterostichus (Lianoe) tanakai Ishida 1964
Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida 1972
Pterostichus (Melanius) aterrimus (Herbst 1784)
Pterostichus (Metallophilus) rugosus (Gebler 1823)
Pterostichus (Morphohaptoderus) hubeicus Facchini & Sciaky 2003
Pterostichus (Morphohaptoderus) yulongshanensis Sciaky 1997
Pterostichus (Neohaptoderus) jureceki (Jedlicka 1936)
Pterostichus (Nialoe) ohbayashii Ishida 1968
Pterostichus (Nialoe) okutamae Tanaka 1963
Pterostichus (Nialoe) opacipennis Jedlicka 1934
Pterostichus (Nialoe) shinbodakensis Sasakawa & H. Ito 2017
Pterostichus (Nialoe) taoi Kasahara 1993
Pterostichus (Nialoe) yakushimanus Nakane & Ishida 1961
Pterostichus (Oreophilus) morio (Duftschmid 1812)
Pterostichus (Oreophilus) spinolae (Dejean 1828)
Pterostichus (Oreoplatysma) abishirensis Belousov 1991
Pterostichus (Oreoplatysma) casaleianus Kirschenhofer 1991
Pterostichus (Oreoplatysma) kvirensis Belousov 1991
Pterostichus (Oreoplatysma) mediocris Dvorak 1995
Pterostichus (Oreoplatysma) orman Jedlicka 1963
Pterostichus (Oreoplatysma) rousianus Kirschenhofer 1987
Pterostichus (Oreoplatysma) satyrus Kurnakov 1962
Pterostichus (Paranialoe) biexcisus Straneo 1955
Pterostichus (Petrophilus) alexandrovi Lafer 1979
Pterostichus (Petrophilus) arsenjevi Lafer 1979
Pterostichus (Petrophilus) pertinax (Tschitscherine 1895)
Pterostichus (Petrophilus) tomensis (Gebler 1847)
Pterostichus (Phonias) diligens (Sturm 1824)
Pterostichus (Phonias) gyrosus (Motschulsky 1866)
Pterostichus (Platysma) sachtlebeni Jedlicka 1962
Pterostichus (Pseudethira) gobettii Straneo 1983
Pterostichus (Pseudethira) immarginatus Straneo 1977
Pterostichus (Pseudethira) matsumurai Habu 1973
Pterostichus (Pseudethira) miehei J. Schmidt 2005
Pterostichus (Pseudethira) weiperti J. Schmidt 2005
Pterostichus (Pseudomaseus) aimaki Jedlicka 1968
Pterostichus (Pseudomaseus) rhaeticus Heer 1837
Pterostichus (Pseudomaseus) tenuis (Casey 1924)
Pterostichus (Pterostichus) auratus Heer 1837
Pterostichus (Pterostichus) devillei Puel 1924
Pterostichus (Pterostichus) esakii Ishida 1959
Pterostichus (Pterostichus) pedemontanus Ganglbauer 1891
Pterostichus (Pterostichus) ruffoi Sciaky 1984
Pterostichus (Rhagadus) modicellus (Tschitscherine 1897)
Pterostichus (Sinosteropus) jani Sciaky & Facchini 2003
Pterostichus (Steropus) catalonicus J. Daniel ex K. & J. Daniel 1906
Pterostichus (Tschitscherinea) mulensis Sciaky 1997
Pterostichus bischoffianus Jedlicka 1936
Pterostichus exochus Andrewes 1930
Pterostichus nishidai Nakane 1989
Pterostichus pentheri Apfelbeck 1918
Pterostichus punctisternus Straneo 1989
Pterostichus umbralis Gistel 1857
Pterostichus vetustus Heer 1862
1100 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy Version 2.2.4

Pterostichus (Hypherpes) herculaneus Mannerheim 1843 (this page)
Pterostichus (Agastillus) eriwanicus (Tschitscherine 1897)
Pterostichus (Aphaonus) arcanoides (Lorenz 1998)
Pterostichus (Aphaonus) cylindriformis (Reitter 1887)
Pterostichus (Argutor) cursor (Dejean 1828)
Pterostichus (Badistrinus) goschi Jedlicka 1930
Pterostichus (Bothriopterus) ferghanicus Kabak 2001
Pterostichus (Bothriopterus) oblongopunctatus (Fabricius 1787)
Pterostichus (Bothriopterus) oregonus Le Conte 1861
Pterostichus (Bothriopterus) pensylvanicus Le Conte 1873
Pterostichus (Bothriopterus) zyzzovi O. & E. Berlov 1996
Pterostichus (Circinatus) agilis Allegro & Sciaky 2010
Pterostichus (Cryobius) arcticola (Chaudoir 1868)
Pterostichus (Cryobius) asiaticus (Poppius 1906)
Pterostichus (Cryobius) glacialis (Brisout de Barneville 1863)
Pterostichus (Cryobius) korgei Jedlicka 1964
Pterostichus (Cryobius) kultianus Jedlicka 1946
Pterostichus (Cryobius) mandibularoides Ball 1966
Pterostichus (Cryobius) palekhai E. & O. Berlov 1996
Pterostichus (Cryobius) parasimilis Ball 1962
Pterostichus (Cryobius) procerulus (Heyden 1880)
Pterostichus (Cryobius) tatricus Kult 1947
Pterostichus (Eosteropus) abudarini O. & E. Berlov 1997
Pterostichus (Eosteropus) tokui Morita 2007
Pterostichus (Epinialoe) janoi Jedlicka 1952
Pterostichus (Ethira) sharanus (Straneo 1984)
Pterostichus (Eurythoracana) haptoderoides (Tschitscherine 1889)
Pterostichus (Falsargutor) pseudopedius Reitter 1887
Pterostichus (Gastrosticta) obesulus Le Conte 1873
Pterostichus (Gastrosticta) putus Casey 1913
Pterostichus (Haplomaseus) andreae (Tschitscherine 1897)
Pterostichus (Haplomaseus) armenus (Faldermann 1836)
Pterostichus (Haplomaseus) caucasicus (Ménétriés 1832)
Pterostichus (Haptotapinus) vseteckai (Maran 1944)
Pterostichus (Hypherpes) annosus Casey 1913
Pterostichus (Hypherpes) ecarinatus Hatch 1936
Pterostichus (Hypherpes) scutellaris Le Conte 1873
Pterostichus (Hypherpes) setosus Hatch 1951
Pterostichus (Koreonialoe) teretis Park & Kwon 1996
Pterostichus (Lenapterus) agonus G. Horn 1880
Pterostichus (Lenapterus) punctatissimus (Randall 1838)
Pterostichus (Leptoferonia) falli Van Dyke 1926
Pterostichus (Leptoferonia) hatchi Hacker 1968
Pterostichus (Leptoferonia) infernalis Hatch 1936
Pterostichus (Leptoferonia) stapedius Hacker 1968
Pterostichus (Leptoferonia) yosemitensis Hacker 1968
Pterostichus (Lianoe) drescoi Nègre 1957
Pterostichus (Lianoe) tanakai Ishida 1964
Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida 1972
Pterostichus (Melanius) aterrimus (Herbst 1784)
Pterostichus (Metallophilus) rugosus (Gebler 1823)
Pterostichus (Morphohaptoderus) hubeicus Facchini & Sciaky 2003
Pterostichus (Morphohaptoderus) yulongshanensis Sciaky 1997
Pterostichus (Neohaptoderus) jureceki (Jedlicka 1936)
Pterostichus (Neohaptoderus) yunnanensis Jedlicka 1934
Pterostichus (Nialoe) ohbayashii Ishida 1968
Pterostichus (Nialoe) okutamae Tanaka 1963
Pterostichus (Nialoe) opacipennis Jedlicka 1934
Pterostichus (Nialoe) shinbodakensis Sasakawa & H. Ito 2017
Pterostichus (Nialoe) taoi Kasahara 1993
Pterostichus (Nialoe) yakushimanus Nakane & Ishida 1961
Pterostichus (Oreophilus) morio (Duftschmid 1812)
Pterostichus (Oreophilus) spinolae (Dejean 1828)
Pterostichus (Oreoplatysma) abishirensis Belousov 1991
Pterostichus (Oreoplatysma) casaleianus Kirschenhofer 1991
Pterostichus (Oreoplatysma) kvirensis Belousov 1991
Pterostichus (Oreoplatysma) mediocris Dvorak 1995
Pterostichus (Oreoplatysma) orman Jedlicka 1963
Pterostichus (Oreoplatysma) rousianus Kirschenhofer 1987
Pterostichus (Oreoplatysma) satyrus Kurnakov 1962
Pterostichus (Paranialoe) biexcisus Straneo 1955
Pterostichus (Petrophilus) alexandrovi Lafer 1979
Pterostichus (Petrophilus) arsenjevi Lafer 1979
Pterostichus (Petrophilus) pertinax (Tschitscherine 1895)
Pterostichus (Petrophilus) tomensis (Gebler 1847)
Pterostichus (Phonias) diligens (Sturm 1824)
Pterostichus (Phonias) gyrosus (Motschulsky 1866)
Pterostichus (Platysma) sachtlebeni Jedlicka 1962
Pterostichus (Pseudethira) gobettii Straneo 1983
Pterostichus (Pseudethira) immarginatus Straneo 1977
Pterostichus (Pseudethira) matsumurai Habu 1973
Pterostichus (Pseudethira) miehei J. Schmidt 2005
Pterostichus (Pseudethira) weiperti J. Schmidt 2005
Pterostichus (Pseudomaseus) aimaki Jedlicka 1968
Pterostichus (Pseudomaseus) rhaeticus Heer 1837
Pterostichus (Pseudomaseus) tenuis (Casey 1924)
Pterostichus (Pterostichus) auratus Heer 1837
Pterostichus (Pterostichus) devillei Puel 1924
Pterostichus (Pterostichus) esakii Ishida 1959
Pterostichus (Pterostichus) pedemontanus Ganglbauer 1891
Pterostichus (Pterostichus) ruffoi Sciaky 1984
Pterostichus (Rhagadus) modicellus (Tschitscherine 1897)
Pterostichus (Sinosteropus) jani Sciaky & Facchini 2003
Pterostichus (Steropus) catalonicus J. Daniel in K. & J. Daniel 1906
Pterostichus (Tschitscherinea) mulensis Sciaky 1997
Pterostichus bischoffianus Jedlicka 1936
Pterostichus exochus Andrewes 1930
Pterostichus nishidai Nakane 1989
Pterostichus pentheri Apfelbeck 1918
Pterostichus punctisternus Straneo 1989
Pterostichus umbralis Gistel 1857
1082 additional siblings truncated for brevity.
See the resource file for a full list.

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Pterostichus Bonelli 1810
Pterostichus herculaneus Mannerheim 1843 (this page)
Pterostichus adulterinus Wrase & J. Schmidt 2006
Pterostichus alexeji Zamotajlov & Kryzhanovskij 1992
Pterostichus altaiensis (Poppius 1906)
Pterostichus amagisanus Tanaka & Ishida 1972
Pterostichus amanoi Nakane 1968
Pterostichus amoenus (Dejean 1828)
Pterostichus andreae (Tschitscherine 1897)
Pterostichus angoarnigi Morvan 1994
Pterostichus apenninus (Dejean 1831)
Pterostichus arcticola (Chaudoir 1868)
Pterostichus arrowi Jedlicka 1936
Pterostichus asymmetricus Bates 1883
Pterostichus aubryi Coiffait 1952
Pterostichus auratus Heer 1837
Pterostichus balthasari Jedlicka 1937
Pterostichus barbarinus Casey 1913
Pterostichus bifoveolatus Park, Kwon & Lafer 1996
Pterostichus bisetosus (Straneo 1938)
Pterostichus bituberculatus (Tschitscherine 1899)
Pterostichus brunneipennis Straneo 1955
Pterostichus capitolinus Kurnakov 1962
Pterostichus cervenkai Sciaky 1994
Pterostichus crassiusculus (Chaudoir 1868)
Pterostichus craterensis (Hatch 1949)
Pterostichus cylindricus (Herbst 1784)
Pterostichus daisenicus Ishida 1958
Pterostichus dentifer Allegro & Sciaky 2010
Pterostichus dhorpatanicus Straneo 1977
Pterostichus dolens (Tschitscherine 1900)
Pterostichus dostali Kirschenhofer 1981
Pterostichus ebeninus (Dejean 1828)
Pterostichus empetricola (Dejean 1828)
Pterostichus faldermanni Schatzmayr 1929
Pterostichus fenestratus Kasahara & Ohtani 1989
Pterostichus firmus (Lutshnik 1933)
Pterostichus foveolatus (Duftschmid 1812)
Pterostichus galae Farkac & Plutenko 1996
Pterostichus gerstlensis Ball 1962
Pterostichus gravis Jedlicka 1939
Pterostichus hagenbachii Sturm 1824
Pterostichus harmandi (Tschitscherine 1900)
Pterostichus hirobane Habu & Baba 1958
Pterostichus iripennis (Chaudoir 1868)
Pterostichus jurecekianus Maran 1940
Pterostichus kambaiti (Andrewes 1947)
Pterostichus katashinensis Habu 1958
Pterostichus kitakamisanus Sasakawa 2005
Pterostichus lachrymosus (Newman 1838)
Pterostichus lacunosus (Chaudoir 1844)
Pterostichus laferi O. & E. Berlov 1996
Pterostichus lanei Van Dyke 1926
Pterostichus liodactylus (Tschitscherine 1898)
Pterostichus longipes (Poppius 1906)
Pterostichus macer (Marsham 1802)
Pterostichus macrogenys Bates 1883
Pterostichus macrothorax (Poppius 1906)
Pterostichus masidai Ishida 1959
Pterostichus microbus Sciaky & Facchini 2003
Pterostichus multiseta (Straneo 1984)
Pterostichus nigellus A. Morawitz 1862
Pterostichus nigrocaeruleus Van Dyke 1926
Pterostichus nordqvisti (J. Sahlberg 1887)
Pterostichus ohsawacavus Sasakawa 2005
Pterostichus olafi J. Schmidt 2005
Pterostichus pfizenmayeri Poppius 1906
Pterostichus planctus Le Conte 1853
Pterostichus protensiformis (Casey 1924)
Pterostichus pseudopercus (Reitter 1889)
Pterostichus pseudoplatyderus Sciaky 1994
Pterostichus pseudosinensis Sciaky & Facchini 2003
Pterostichus putus Casey 1913
Pterostichus rolex Morvan 1995
Pterostichus rousianus Kirschenhofer 1987
Pterostichus sacheri I. Frivaldszky von Frivald 1865
Pterostichus scalptus Sciaky & Wrase 1997
Pterostichus schmidtii (Chaudoir 1838)
Pterostichus schneideri Sciaky & Facchini 2003
Pterostichus scuticollis (Fairmaire 1889)
Pterostichus serripes (Le Conte 1875)
Pterostichus seunglaki Park & Kwon 1996
Pterostichus shaovuensis Jedlicka 1956
Pterostichus skoupyi Hovorka & Skoupy 2007
Pterostichus stantonensis Ball 1966
Pterostichus stygicus (Say 1823)
Pterostichus styx (Andrewes 1937)
Pterostichus subiasi (Ortuño & Zaballos 1992)
Pterostichus swaneticus (Reitter 1883)
Pterostichus tamarae Wrase & Kirschenhofer 1991
Pterostichus tatricus Kult 1947
Pterostichus tenuimarginatus (Chaudoir 1868)
Pterostichus tiliaceoradix Ball 1962
Pterostichus triangularis Sciaky & Facchini 2003
Pterostichus trinarius (Casey 1918)
Pterostichus triseriatus (Gebler 1847)
Pterostichus tuberculofemoratus Hatch 1936
Pterostichus validiceps Reitter 1887
Pterostichus variabilis (Ménétriés 1832)
Pterostichus vicinus Mannerheim 1843
Pterostichus weigeli J. Schmidt 2005
Pterostichus zyzzovi O. & E. Berlov 1996
1090 additional siblings truncated for brevity.
See the resource file for a full list.

Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Coleoptera Linnaeus 1758
Adephaga Schellenberg 1806
Carabidae Latreille 1802
Harpalinae Bonelli 1810
Pterostichini Bonelli 1810
Pterostichus Bonelli 1810
Pterostichus subgen. Hypherpes Chaudoir 1838
Pterostichus herculaneus Mannerheim 1843 (this page)
Pterostichus adoxus (Say 1823)
Pterostichus algidus Le Conte 1853
Pterostichus amethystinus Mannerheim 1843
Pterostichus annosus Casey 1913
Pterostichus arcanus Casey 1913
Pterostichus baldwini (Casey 1924)
Pterostichus barbarinus Casey 1913
Pterostichus brachylobus Kavanaugh & La Bonte 2006
Pterostichus californicus (Dejean 1828)
Pterostichus canallatus Casey 1913
Pterostichus castaneus (Dejean 1828)
Pterostichus castanipes (Ménétriés 1843)
Pterostichus congestus (Ménétriés 1843)
Pterostichus craterensis (Hatch 1949)
Pterostichus crenicollis Le Conte 1873
Pterostichus ecarinatus Hatch 1936
Pterostichus esuriens Casey 1913
Pterostichus gliscans Casey 1913
Pterostichus gracilior Le Conte 1873
Pterostichus hornii Le Conte 1873
Pterostichus illustris Le Conte 1851
Pterostichus inermis Fall 1901
Pterostichus isabellae Le Conte 1851
Pterostichus jacobinus Casey 1913
Pterostichus laborans Casey 1913
Pterostichus lacertus Casey 1913
Pterostichus lama (Ménétriés 1843)
Pterostichus lassulus (Casey 1920)
Pterostichus lattini La Bonte 2006
Pterostichus luscus (Casey 1918)
Pterostichus menetriesii Le Conte 1873
Pterostichus mercedianus (Casey 1918)
Pterostichus miscellus Casey 1913
Pterostichus morionides (Chaudoir 1868)
Pterostichus neobrunneus Lindroth 1966
Pterostichus nigrocaeruleus Van Dyke 1926
Pterostichus obsidianus Casey 1913
Pterostichus occultus Casey 1913
Pterostichus ordinarius Casey 1913
Pterostichus ovalipennis Casey 1913
Pterostichus panticulatus Casey 1913
Pterostichus pergracilis (Casey 1920)
Pterostichus planctus Le Conte 1853
Pterostichus protensiformis (Casey 1924)
Pterostichus protractus Le Conte 1860
Pterostichus restrictus (Casey 1918)
Pterostichus scutellaris Le Conte 1873
Pterostichus serripes (Le Conte 1875)
Pterostichus setosus Hatch 1951
Pterostichus sierranus Casey 1913
Pterostichus sponsor Casey 1913
Pterostichus spraguei Le Conte 1873
Pterostichus suffusus Casey 1913
Pterostichus tarsalis Le Conte 1873
Pterostichus tristis (Dejean 1828)
Pterostichus tuberculofemoratus Hatch 1936
Pterostichus vandykei Schaeffer 1910
Pterostichus vicinus Mannerheim 1843
Pterostichus ybousqueti Berlov 1999

NCBI

root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Adephaga
Caraboidea
Carabidae
Pterostichinae
Pterostichus
'Hypherpes subgenus complex'
Hypherpes
Pterostichus herculaneus (this page)
Pterostichus adoxus
Pterostichus algidus
Pterostichus amethystinus
Pterostichus brachylobus
Pterostichus californicus
Pterostichus canallatus
Pterostichus castaneus
Pterostichus congestus
Pterostichus crenicollis
Pterostichus diabolus
Pterostichus ecarinatus
Pterostichus inermis
Pterostichus intectus
Pterostichus isabellae
Pterostichus jacobinus
Pterostichus lama
Pterostichus lattini
Pterostichus menetriesii
Pterostichus miscellus
Pterostichus morionides
Pterostichus neobrunneus
Pterostichus nigrocaeruleus
Pterostichus protractus
Pterostichus serripes
Pterostichus tarsalis
Pterostichus tristis
Pterostichus tuberculofemoratus

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Adephaga
Caraboidea
Carabidae
Harpalinae
Pterostichini
Pterostichus
Pterostichus herculaneus (this page)
Pterostichus aanistschenkoi
Pterostichus aapsorum
Pterostichus abaciformis
Pterostichus abagonensis
Pterostichus abasarukini
Pterostichus abasinus
Pterostichus abaxoides
Pterostichus abensis
Pterostichus abishirensis
Pterostichus abnormis
Pterostichus abudarini
Pterostichus acuspina
Pterostichus acutidens
Pterostichus acutipes
Pterostichus adatarasanus
Pterostichus ademidovae
Pterostichus adoxus
Pterostichus adstrictus
Pterostichus adulterinus
Pterostichus aemiliae
Pterostichus aeneocupreus
Pterostichus aenescens
Pterostichus aeneus
Pterostichus aereipennis
Pterostichus agilis
Pterostichus agonus
Pterostichus aibgensis
Pterostichus aimaki
Pterostichus ainus
Pterostichus akitai
Pterostichus akkusianus
Pterostichus akozyrevi
Pterostichus aksekianus
Pterostichus alacer
Pterostichus alamedae
Pterostichus alexandrovi
Pterostichus alexeji
Pterostichus algidus
Pterostichus altaianus
Pterostichus altaicus
Pterostichus altaiensis
Pterostichus amagisanus
Pterostichus amanoi
Pterostichus amaroides
Pterostichus ambigenus
Pterostichus ambiguus
Pterostichus amethystinus
Pterostichus amoenus
Pterostichus amurensis
Pterostichus analis
Pterostichus anatolicus
Pterostichus andreae
Pterostichus andreinii
Pterostichus andrewesi
Pterostichus angoarnigi
Pterostichus angustus
Pterostichus annosus
Pterostichus anomostriatus
Pterostichus apenninus
Pterostichus apfelbecki
Pterostichus apiculatiphallus
Pterostichus aralarensis
Pterostichus araraticus
Pterostichus arator
Pterostichus aratus
Pterostichus arcanoides
Pterostichus arcanus
Pterostichus archangaicus
Pterostichus arcticola
Pterostichus arcuaticarinatus
Pterostichus armenus
Pterostichus arnoldii
Pterostichus arribasi
Pterostichus arrowianus
Pterostichus arvernus
Pterostichus asahinus
Pterostichus asiaticus
Pterostichus asturicus
Pterostichus astutus
Pterostichus asymmetricus
Pterostichus aterrimus
Pterostichus atratus
Pterostichus atrox
Pterostichus aubryi
Pterostichus audax
Pterostichus auratus
Pterostichus auriga
Pterostichus avaricus
Pterostichus baenningeri
Pterostichus blandus
Pterostichus calcaratus
Pterostichus diligens
Pterostichus minor
Pterostichus nigrita
Pterostichus oblongopunctatus
Pterostichus obscuripennis
Pterostichus parvulus
Pterostichus strenuus
Pterostichus subgen. Morphohaptoderus
Pterostichus vernalis
1308 additional siblings truncated for brevity.
See the resource file for a full list.
EOL is hosted by: