Nipple Bract Arrowhead

Herbs, perennial, to 112 cm; rhizomes not coarse; stolons absent; corms absent. Leaves submersed and emergent; petiole triangular, 11.5--35 cm; blade elliptic to ovate, 25--56 cm. Inflorescences racemes or panicles, of 4--10 whorls, emersed, 8--25 ´ 5--36 cm; peduncles to 76 cm; bracts connate ¼ total length, lanceolate, 4--8 mm, papillose; fruiting pedicels ascending to spreading, cylindric, 1--4.5 cm. Flowers to 3 cm diam.; sepals appressed to spreading; filaments slightly dilated, shorter than anthers, glabrous; pistillate pedicellate, without ring of sterile stamens. Fruiting heads 0.7--1 cm diam.; achenes obovoid, abaxially keeled, 1.2--1.5 ´ 0.7--1.1 mm, beaked; faces not tuberculate, wings 1, ± entire, glands 0--1; beak lateral, erect, 0.1--0.3 mm.

Ark., La., Okla., Tex.

Flowering spring--fall.

Swamps, marshes, bogs, ditches, borders of lakes; 0--300m.

Sagittaria papillosa Buch. Abh. Nat. Ver
Bremen 2 : 44. 1869.
Sagittaria lancifolia papulosa Micheli, in DC. Monog. Phan. 3 : 74. 1881.
Plants mostly emersed, 3-10 dm. tall ; leaves erect or. nearly so, the blades linear or linear-lanceolate, 10-15 cm. long, acute, as long as the petioles or shorter ; scapes usually longer than the leaves, mostly branched ; whorls of the inflorescence three to ten or more, the lower ones remote ; pedicels of pistillate flowers 1-1.5 cm. long ; bracts ovate, 3-4 mm. long, densely granular-papillose, united to about the middle ; sepals ovate, roughened like the bracts, becoming 5-6 mm. long ; corollas 2-3 cm. broad ; filaments not dilated, glabrous ; fruit-heads subglobose, 6-10 mm. in diameter ; achenes cuneate, about 1.5 mm. long, the beak lateral and slightly above the middle of the achene-body.
Type locality : Not given [Texas] . Distribution : Louisiana and Texas.

Sagittaria papillosa, the nipplebract arrowhead, is a plant species native to the south-central United States (Texas, Oklahoma, Louisiana, Arkansas and Mississippi).

Sagittaria papillosa grows in wet places such as marshes and the banks of lakes and slow-moving streams. It is a perennial herb up to 120 cm tall. Petioles are triangular in cross-section, the leaf blade very narrowly elliptical to ovate, not lobed. The species is distinguished from others in the genus by having bumps (papillae) resembling nipples on the flower bracts.