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Image of Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823)
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Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823)

Scientific Names

Preferred Names

Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823)
Species recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus montanellus
Species recognized by United States Species List
Pterostichus pinguedineus
Species recognized by wikipedia NL, Canada Species List, North Pacific Species List, NCBI, Arctic Arthropods, and United States Species List
Pterostichus pinguedineus (Eschscholtz 1823)
Species recognized by GBIF classification
Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823)
Species recognized by EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius beringi Casey
Recognized by NMNH type specimens
Cryobius washingtoni Casey
Recognized by NMNH type specimens
Poecilus pinguedineus
Species recognized by wikidata_hierarchy tar gz
Pterostichus (Cryobius) pinguedineus
Species recognized by Paleobiology Database
Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823)
Species recognized by Catalogue of Life 7 February 2023 and Dynamic Hierarchy Version 2.2 - Test
Pterostichus (Cryobius) splendidus Park 2013
Species recognized by Catalogue of Life 7 February 2023
Pterostichus mandibularis
Species recognized by wikidata_hierarchy tar gz
Pterostichus pinguedineus
Species recognized by wikidata_hierarchy tar gz, wikipedia EN, Barcode of Life Data Systems, and Global Biotic Interactions
Pterostichus pinguedineus
Recognized by Alaskan Arthropods
Pterostichus pinguedineus (Eschscholtz 1823)
Species recognized by Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy and TreatmentBank

Alternative Names

Argutor mandibularis Kirby 1837
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius alaskensis Poppius 1906
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius beringensis (Poppius 1908)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius beringi Casey 1918
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius fortestriatus Poppius 1906
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius holmbergi Poppius 1906
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius incognitus Poppius 1906
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius montanellus (Poppius 1907)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius pinguedineus (Eschscholtz 1823)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius repandus Poppius 1906
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius subnitidulus Poppius 1906
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Cryobius washingtoni Casey 1920
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Feronia despecta J. Sahlberg 1885
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Feronia diplogma Chaudoir 1868
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Feronia frigida Dejean 1828
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Feronia laeviuscula J. Sahlberg 1885
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Feronia splendida J. Sahlberg 1885
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Feronia stuxbergi Mäklin 1878
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Feronia subsinuosa Chaudoir 1868
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Feronia sulcipennis J. Sahlberg 1885
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Poecilus pinguedineus Eschscholtz 1823
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus alaskensis (Poppius 1906)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus anadyricus Csiki 1930
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus beringensis Poppius 1908
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus beringi (Casey 1918)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus despectus (J. Sahlberg 1885)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus diplogmus (Chaudoir 1868)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus fortestriatus (Poppius 1906)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus frigidus (Dejean 1828)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus holmbergi (Poppius 1906)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus incognitus (Poppius 1906)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus laeviusculus (J. Sahlberg 1885)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus mandibularis (Kirby 1837)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus montanellus Poppius 1907
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus repandus (Poppius 1906)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus splendidus (J. Sahlberg 1885)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus stuxbergi (Mäklin 1878)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus subnitidulus (Poppius 1906)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus subsinuosus (Chaudoir 1868)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus sulcipennis (J. Sahlberg 1885)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pterostichus washingtoni (Casey 1920)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Argutor mandibularis Kirby 1837
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius alaskensis Poppius 1906
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius beringensis (Poppius 1908)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius beringi Casey 1918
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius fortestriatus Poppius 1906
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius holmbergi Poppius 1906
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius incognitus Poppius 1906
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius montanellus (Poppius 1907)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius pinguedineus (Eschscholtz 1823)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius repandus Poppius 1906
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius subnitidulus Poppius 1906
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Cryobius washingtoni Casey 1920
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Feronia despecta J. Sahlberg 1885
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Feronia diplogma Chaudoir 1868
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Feronia frigida Dejean 1828
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Feronia laeviuscula J. Sahlberg 1885
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Feronia splendida J. Sahlberg 1885
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Feronia stuxbergi Mäklin 1878
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Feronia subsinuosa Chaudoir 1868
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Feronia sulcipennis J. Sahlberg 1885
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Poecilus pinguedineus Eschscholtz 1823
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus alaskensis (Poppius 1906)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus anadyricus Csiki 1930
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus beringensis Poppius 1908
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus beringi (Casey 1918)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus despectus (J. Sahlberg 1885)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus diplogmus (Chaudoir 1868)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus fortestriatus (Poppius 1906)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus frigidus (Dejean 1828)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus holmbergi (Poppius 1906)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus incognitus (Poppius 1906)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus laeviusculus (J. Sahlberg 1885)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus mandibularis (Kirby 1837)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus montanellus Poppius 1907
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus repandus (Poppius 1906)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus splendidus (J. Sahlberg 1885)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus stuxbergi (Mäklin 1878)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus subnitidulus (Poppius 1906)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus subsinuosus (Chaudoir 1868)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus sulcipennis (J. Sahlberg 1885)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pterostichus washingtoni (Casey 1920)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Argutor mandibularis Kirby 1837
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Argutor mandibularis Kirby 1837
Synonym according to Catalogue of Life 7 February 2023
Cryobius alaskensis Poppius 1906
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius alaskensis Poppius 1906
Synonym according to Catalogue of Life 7 February 2023
Cryobius beringensis (Poppius 1908)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius beringensis (Poppius 1908)
Synonym according to Catalogue of Life 7 February 2023
Cryobius beringi Casey 1918
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius beringi Casey 1918
Synonym according to Catalogue of Life 7 February 2023
Cryobius fortestriatus Poppius 1906
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius fortestriatus Poppius 1906
Synonym according to Catalogue of Life 7 February 2023
Cryobius holmbergi Poppius 1906
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius holmbergi Poppius 1906
Synonym according to Catalogue of Life 7 February 2023
Cryobius incognitus Poppius 1906
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius incognitus Poppius 1906
Synonym according to Catalogue of Life 7 February 2023
Cryobius montanellus (Poppius 1907)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius montanellus (Poppius 1907)
Synonym according to Catalogue of Life 7 February 2023
Cryobius pinguedineus (Eschscholtz 1823)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius pinguedineus (Eschscholtz 1823)
Synonym according to Catalogue of Life 7 February 2023
Cryobius repandus Poppius 1906
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius repandus Poppius 1906
Synonym according to Catalogue of Life 7 February 2023
Cryobius subnitidulus Poppius 1906
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius subnitidulus Poppius 1906
Synonym according to Catalogue of Life 7 February 2023
Cryobius washingtoni Casey 1920
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Cryobius washingtoni Casey 1920
Synonym according to Catalogue of Life 7 February 2023
Feronia despecta J. Sahlberg 1885
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Feronia despecta J. Sahlberg 1885
Synonym according to Catalogue of Life 7 February 2023
Feronia diplogma Chaudoir 1868
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Feronia diplogma Chaudoir 1868
Synonym according to Catalogue of Life 7 February 2023
Feronia frigida Dejean 1828
Ambiguous synonym according to Catalogue of Life 7 February 2023
Feronia frigida Dejean 1828
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Feronia laeviuscula J. Sahlberg 1885
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Feronia laeviuscula J. Sahlberg 1885
Synonym according to Catalogue of Life 7 February 2023
Feronia splendida J. Sahlberg 1885
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Feronia splendida J. Sahlberg 1885
Synonym according to Catalogue of Life 7 February 2023
Feronia stuxbergi Mäklin 1878
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Feronia stuxbergi Mäklin 1878
Synonym according to Catalogue of Life 7 February 2023
Feronia subsinuosa Chaudoir 1868
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Feronia subsinuosa Chaudoir 1868
Synonym according to Catalogue of Life 7 February 2023
Feronia sulcipennis J. Sahlberg 1885
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Feronia sulcipennis J. Sahlberg 1885
Synonym according to Catalogue of Life 7 February 2023
Poecilus pinguedineus
Recombined according to Paleobiology Database
Poecilus pinguedineus Eschscholtz 1823
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Poecilus pinguedineus Eschscholtz 1823
Synonym according to Catalogue of Life 7 February 2023
Pterostichus alaskensis (Poppius 1906)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus alaskensis (Poppius 1906)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus anadyricus Csiki 1930
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus anadyricus Csiki 1930
Synonym according to Catalogue of Life 7 February 2023
Pterostichus beringensis Poppius 1908
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus beringensis Poppius 1908
Synonym according to Catalogue of Life 7 February 2023
Pterostichus beringi (Casey 1918)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus beringi (Casey 1918)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus despectus (J. Sahlberg 1885)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus despectus (J. Sahlberg 1885)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus diplogmus (Chaudoir 1868)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus diplogmus (Chaudoir 1868)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus fortestriatus (Poppius 1906)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus fortestriatus (Poppius 1906)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus frigidus (Dejean 1828)
Ambiguous synonym according to Catalogue of Life 7 February 2023
Pterostichus frigidus (Dejean 1828)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus holmbergi (Poppius 1906)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus holmbergi (Poppius 1906)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus incognitus (Poppius 1906)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus incognitus (Poppius 1906)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus laeviusculus (J. Sahlberg 1885)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus laeviusculus (J. Sahlberg 1885) junior homonym
Ambiguous synonym according to Catalogue of Life 7 February 2023
Pterostichus mandibularis (Kirby 1837)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus mandibularis (Kirby 1837)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus mandibularis Kirby 1837
Invalid according to Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy
Pterostichus montanellus Poppius 1907
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus montanellus Poppius 1907
Synonym according to Catalogue of Life 7 February 2023
Pterostichus repandus (Poppius 1906)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus repandus (Poppius 1906)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus splendidus (J. Sahlberg 1885)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus splendidus (J. Sahlberg 1885)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus stuxbergi (Mäklin 1878)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus stuxbergi (Mäklin 1878)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus subnitidulus (Poppius 1906)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus subnitidulus (Poppius 1906)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus subsinuosus (Chaudoir 1868)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus subsinuosus (Chaudoir 1868)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus sulcipennis (J. Sahlberg 1885)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus sulcipennis (J. Sahlberg 1885)
Synonym according to Catalogue of Life 7 February 2023
Pterostichus washingtoni (Casey 1920)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pterostichus washingtoni (Casey 1920)
Synonym according to Catalogue of Life 7 February 2023

Common Names

There are no common names associated with this taxon.

Curated hierarchies for Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823)

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Pterostichus
Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823) (this page)
Pterostichus (Agastillus) eriwanicus (Tschitscherine 1897)
Pterostichus (Aphaonus) arcanoides (Lorenz 1998)
Pterostichus (Aphaonus) cylindriformis (Reitter 1887)
Pterostichus (Argutor) cursor (Dejean 1828)
Pterostichus (Badistrinus) goschi Jedlicka 1930
Pterostichus (Bothriopterus) ferghanicus Kabak 2001
Pterostichus (Bothriopterus) oblongopunctatus (Fabricius 1787)
Pterostichus (Bothriopterus) oregonus Le Conte 1861
Pterostichus (Bothriopterus) pensylvanicus Le Conte 1873
Pterostichus (Bothriopterus) zyzzovi O. & E. Berlov 1996
Pterostichus (Circinatus) agilis Allegro & Sciaky 2010
Pterostichus (Cryobius) arcticola (Chaudoir 1868)
Pterostichus (Cryobius) asiaticus (Poppius 1906)
Pterostichus (Cryobius) glacialis (Brisout de Barneville 1863)
Pterostichus (Cryobius) korgei Jedlicka 1964
Pterostichus (Cryobius) kultianus Jedlicka 1946
Pterostichus (Cryobius) mandibularoides Ball 1966
Pterostichus (Cryobius) palekhai E. & O. Berlov 1996
Pterostichus (Cryobius) parasimilis Ball 1962
Pterostichus (Cryobius) procerulus (Heyden 1880)
Pterostichus (Cryobius) tatricus Kult 1947
Pterostichus (Eosteropus) abudarini O. & E. Berlov 1997
Pterostichus (Eosteropus) tokui Morita 2007
Pterostichus (Epinialoe) janoi Jedlicka 1952
Pterostichus (Ethira) sharanus (Straneo 1984)
Pterostichus (Eurythoracana) haptoderoides (Tschitscherine 1889)
Pterostichus (Falsargutor) pseudopedius Reitter 1887
Pterostichus (Gastrosticta) obesulus Le Conte 1873
Pterostichus (Gastrosticta) putus Casey 1913
Pterostichus (Haplomaseus) andreae (Tschitscherine 1897)
Pterostichus (Haplomaseus) armenus (Faldermann 1836)
Pterostichus (Haplomaseus) caucasicus (Ménétriés 1832)
Pterostichus (Haptotapinus) vseteckai (Maran 1944)
Pterostichus (Hypherpes) annosus Casey 1913
Pterostichus (Hypherpes) ecarinatus Hatch 1936
Pterostichus (Hypherpes) scutellaris Le Conte 1873
Pterostichus (Hypherpes) setosus Hatch 1951
Pterostichus (Koreonialoe) teretis Park & Kwon 1996
Pterostichus (Lenapterus) agonus G. Horn 1880
Pterostichus (Lenapterus) punctatissimus (Randall 1838)
Pterostichus (Leptoferonia) falli Van Dyke 1926
Pterostichus (Leptoferonia) hatchi Hacker 1968
Pterostichus (Leptoferonia) infernalis Hatch 1936
Pterostichus (Leptoferonia) stapedius Hacker 1968
Pterostichus (Leptoferonia) yosemitensis Hacker 1968
Pterostichus (Lianoe) drescoi Nègre 1957
Pterostichus (Lianoe) tanakai Ishida 1964
Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida 1972
Pterostichus (Melanius) aterrimus (Herbst 1784)
Pterostichus (Metallophilus) rugosus (Gebler 1823)
Pterostichus (Morphohaptoderus) hubeicus Facchini & Sciaky 2003
Pterostichus (Morphohaptoderus) yulongshanensis Sciaky 1997
Pterostichus (Neohaptoderus) jureceki (Jedlicka 1936)
Pterostichus (Neohaptoderus) yunnanensis Jedlicka 1934
Pterostichus (Nialoe) ohbayashii Ishida 1968
Pterostichus (Nialoe) okutamae Tanaka 1963
Pterostichus (Nialoe) opacipennis Jedlicka 1934
Pterostichus (Nialoe) shinbodakensis Sasakawa & H. Ito 2017
Pterostichus (Nialoe) taoi Kasahara 1993
Pterostichus (Nialoe) yakushimanus Nakane & Ishida 1961
Pterostichus (Oreophilus) morio (Duftschmid 1812)
Pterostichus (Oreophilus) spinolae (Dejean 1828)
Pterostichus (Oreoplatysma) abishirensis Belousov 1991
Pterostichus (Oreoplatysma) casaleianus Kirschenhofer 1991
Pterostichus (Oreoplatysma) kvirensis Belousov 1991
Pterostichus (Oreoplatysma) mediocris Dvorak 1995
Pterostichus (Oreoplatysma) orman Jedlicka 1963
Pterostichus (Oreoplatysma) rousianus Kirschenhofer 1987
Pterostichus (Oreoplatysma) satyrus Kurnakov 1962
Pterostichus (Paranialoe) biexcisus Straneo 1955
Pterostichus (Petrophilus) alexandrovi Lafer 1979
Pterostichus (Petrophilus) arsenjevi Lafer 1979
Pterostichus (Petrophilus) pertinax (Tschitscherine 1895)
Pterostichus (Petrophilus) tomensis (Gebler 1847)
Pterostichus (Phonias) diligens (Sturm 1824)
Pterostichus (Phonias) gyrosus (Motschulsky 1866)
Pterostichus (Platysma) sachtlebeni Jedlicka 1962
Pterostichus (Pseudethira) gobettii Straneo 1983
Pterostichus (Pseudethira) immarginatus Straneo 1977
Pterostichus (Pseudethira) matsumurai Habu 1973
Pterostichus (Pseudethira) miehei J. Schmidt 2005
Pterostichus (Pseudethira) weiperti J. Schmidt 2005
Pterostichus (Pseudomaseus) aimaki Jedlicka 1968
Pterostichus (Pseudomaseus) rhaeticus Heer 1837
Pterostichus (Pseudomaseus) tenuis (Casey 1924)
Pterostichus (Pterostichus) auratus Heer 1837
Pterostichus (Pterostichus) devillei Puel 1924
Pterostichus (Pterostichus) esakii Ishida 1959
Pterostichus (Pterostichus) pedemontanus Ganglbauer 1891
Pterostichus (Pterostichus) ruffoi Sciaky 1984
Pterostichus (Rhagadus) modicellus (Tschitscherine 1897)
Pterostichus (Sinosteropus) jani Sciaky & Facchini 2003
Pterostichus (Steropus) catalonicus J. Daniel ex K. & J. Daniel 1906
Pterostichus (Tschitscherinea) mulensis Sciaky 1997
Pterostichus bischoffianus Jedlicka 1936
Pterostichus exochus Andrewes 1930
Pterostichus nishidai Nakane 1989
Pterostichus pentheri Apfelbeck 1918
Pterostichus punctisternus Straneo 1989
Pterostichus umbralis Gistel 1857
1082 additional siblings truncated for brevity.
See the resource file for a full list.

Catalogue of Life 7 February 2023

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Caraboidea
Carabidae Latreille 1802
Pterostichinae Bonelli 1810
Pterostichini Bonelli 1810
Pterostichina Bonelli 1810
Pterostichus Bonelli 1810
Pterostichus subgen. Cryobius Chaudoir 1838
Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823) (this page)
Pterostichus (Cryobius) abaxoides (Dejean 1828)
Pterostichus (Cryobius) acrogonus (Chaudoir 1843)
Pterostichus (Cryobius) akkusianus Kirschenhofer 1981
Pterostichus (Cryobius) altaiensis (Poppius 1906)
Pterostichus (Cryobius) amaroides (Dejean 1828)
Pterostichus (Cryobius) amblypterus (Chaudoir 1868)
Pterostichus (Cryobius) amoenus (Dejean 1828)
Pterostichus (Cryobius) amurensis (Poppius 1906)
Pterostichus (Cryobius) anatolicus Jedlicka 1963
Pterostichus (Cryobius) apenninus (Dejean 1831)
Pterostichus (Cryobius) aralarensis (Mateu 1945)
Pterostichus (Cryobius) arcticola (Chaudoir 1868)
Pterostichus (Cryobius) auriga Ball 1962
Pterostichus (Cryobius) barryorum Ball 1962
Pterostichus (Cryobius) biocryus Ball 1962
Pterostichus (Cryobius) blandulus L. Miller 1859
Pterostichus (Cryobius) blumenthali Heinz 1965
Pterostichus (Cryobius) brevicornis (Kirby 1837)
Pterostichus (Cryobius) breviusculus (R. F. Sahlberg 1844)
Pterostichus (Cryobius) bryanti (Van Dyke 1951)
Pterostichus (Cryobius) bryantoides Ball 1962
Pterostichus (Cryobius) cacumensis Ball 1966
Pterostichus (Cryobius) cantabricus (L. Schaufuss 1862)
Pterostichus (Cryobius) caribou Ball 1962
Pterostichus (Cryobius) carradei (Gautier des Cottes 1866)
Pterostichus (Cryobius) champenoisi (Croissandeau 1893)
Pterostichus (Cryobius) colasi (Jeannel 1937)
Pterostichus (Cryobius) coreicus Jedlicka 1962
Pterostichus (Cryobius) ehlersi (Heyden 1881)
Pterostichus (Cryobius) empetricola (Dejean 1828)
Pterostichus (Cryobius) espanoli (J. & E. Vives 1977)
Pterostichus (Cryobius) euxinus Straneo 1935
Pterostichus (Cryobius) exceptus (J. Sahlberg 1885)
Pterostichus (Cryobius) firmus (Lutshnik 1933)
Pterostichus (Cryobius) fulvescens (Motschulsky 1844)
Pterostichus (Cryobius) gerstlensis Ball 1962
Pterostichus (Cryobius) glacialis (Brisout de Barneville 1863)
Pterostichus (Cryobius) glukhomanka Sundukov 2013
Pterostichus (Cryobius) herzi (Poppius 1906)
Pterostichus (Cryobius) homalonotus (Tschitscherine 1894)
Pterostichus (Cryobius) horvatovichi Kirschenhofer 1991
Pterostichus (Cryobius) infimus (Chaudoir 1868)
Pterostichus (Cryobius) inopinatus (Lutshnik 1933)
Pterostichus (Cryobius) iripennis (Chaudoir 1868)
Pterostichus (Cryobius) ishiharai Ishida & Shibata 1961
Pterostichus (Cryobius) jacobsoni (Poppius 1906)
Pterostichus (Cryobius) jaechi Kirschenhofer 1997
Pterostichus (Cryobius) josephi Csiki 1930
Pterostichus (Cryobius) kadjaranze (Jedlicka 1947)
Pterostichus (Cryobius) kaninensis (Poppius 1906)
Pterostichus (Cryobius) kaynashensis Schweiger 1967
Pterostichus (Cryobius) kolymensis Erjiomin 1998
Pterostichus (Cryobius) korgei Jedlicka 1964
Pterostichus (Cryobius) kotzebuei Ball 1962
Pterostichus (Cryobius) kurosai Tanaka 1958
Pterostichus (Cryobius) kurosawai Tanaka 1958
Pterostichus (Cryobius) lamuticus (Poppius 1906)
Pterostichus (Cryobius) latiusculus (Chaudoir 1868)
Pterostichus (Cryobius) longipes (Poppius 1906)
Pterostichus (Cryobius) lucidus (Motschulsky 1844)
Pterostichus (Cryobius) macedonicus Apfelbeck 1918
Pterostichus (Cryobius) macrothorax (Poppius 1906)
Pterostichus (Cryobius) maeklini (Poppius 1906)
Pterostichus (Cryobius) mandibularoides Ball 1966
Pterostichus (Cryobius) marani Jedlicka 1967
Pterostichus (Cryobius) middendorffi (J. Sahlberg 1875)
Pterostichus (Cryobius) negligens (Sturm 1824)
Pterostichus (Cryobius) nigripalpis (Poppius 1906)
Pterostichus (Cryobius) nivalis (R. F. Sahlberg 1844)
Pterostichus (Cryobius) nordqvisti (J. Sahlberg 1885)
Pterostichus (Cryobius) ochoticus (R. F. Sahlberg 1844)
Pterostichus (Cryobius) palekhai E. & O. Berlov 1996
Pterostichus (Cryobius) parasimilis Ball 1962
Pterostichus (Cryobius) parviceps (Poppius 1906)
Pterostichus (Cryobius) pithyusicus (Lutshnik 1933)
Pterostichus (Cryobius) planus (J. Sahlberg 1885)
Pterostichus (Cryobius) poppiusianus (Jakobson 1907)
Pterostichus (Cryobius) procerulus (Heyden 1880)
Pterostichus (Cryobius) properans (Chaudoir 1868)
Pterostichus (Cryobius) pseudostuxbergi (Poppius 1906)
Pterostichus (Cryobius) pumilio (Dejean 1828)
Pterostichus (Cryobius) punctiger (J. Sahlberg 1880)
Pterostichus (Cryobius) quadraticollis (Chaudoir 1846)
Pterostichus (Cryobius) riparius (Dejean 1828)
Pterostichus (Cryobius) rufonitens (Fairmaire 1866)
Pterostichus (Cryobius) safonovae Budarin 1995
Pterostichus (Cryobius) scitus (Mäklin 1878)
Pterostichus (Cryobius) shilenkovi Erjiomin & Kabak 1991
Pterostichus (Cryobius) similis Mannerheim 1852
Pterostichus (Cryobius) soperi Ball 1966
Pterostichus (Cryobius) spornyi Sundukov 2013
Pterostichus (Cryobius) stantonensis Ball 1966
Pterostichus (Cryobius) subgibbus (Motschulsky 1860)
Pterostichus (Cryobius) subiasi (Ortuño & Zaballos 1992)
Pterostichus (Cryobius) surgens Le Conte 1878
Pterostichus (Cryobius) tareumiut Ball 1962
Pterostichus (Cryobius) tatricus Kult 1947
Pterostichus (Cryobius) tichomirovi Erjiomin 1990
Pterostichus (Cryobius) tiliaceoradix Ball 1962
Pterostichus (Cryobius) tokmakovae Sundukov 2013
30 additional siblings truncated for brevity.
See the resource file for a full list.

Catalogue of Life 7 February 2023

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Caraboidea
Carabidae Latreille 1802
Pterostichinae Bonelli 1810
Pterostichini Bonelli 1810
Pterostichina Bonelli 1810
Pterostichus Bonelli 1810
Pterostichus subgen. Cryobius Chaudoir 1838
Pterostichus (Cryobius) splendidus Park 2013 (this page)
Pterostichus (Cryobius) abaxoides (Dejean 1828)
Pterostichus (Cryobius) acrogonus (Chaudoir 1843)
Pterostichus (Cryobius) akkusianus Kirschenhofer 1981
Pterostichus (Cryobius) altaiensis (Poppius 1906)
Pterostichus (Cryobius) amaroides (Dejean 1828)
Pterostichus (Cryobius) amblypterus (Chaudoir 1868)
Pterostichus (Cryobius) amoenus (Dejean 1828)
Pterostichus (Cryobius) amurensis (Poppius 1906)
Pterostichus (Cryobius) anatolicus Jedlicka 1963
Pterostichus (Cryobius) apenninus (Dejean 1831)
Pterostichus (Cryobius) aralarensis (Mateu 1945)
Pterostichus (Cryobius) arcticola (Chaudoir 1868)
Pterostichus (Cryobius) auriga Ball 1962
Pterostichus (Cryobius) barryorum Ball 1962
Pterostichus (Cryobius) biocryus Ball 1962
Pterostichus (Cryobius) blandulus L. Miller 1859
Pterostichus (Cryobius) blumenthali Heinz 1965
Pterostichus (Cryobius) brevicornis (Kirby 1837)
Pterostichus (Cryobius) breviusculus (R. F. Sahlberg 1844)
Pterostichus (Cryobius) bryanti (Van Dyke 1951)
Pterostichus (Cryobius) bryantoides Ball 1962
Pterostichus (Cryobius) cacumensis Ball 1966
Pterostichus (Cryobius) cantabricus (L. Schaufuss 1862)
Pterostichus (Cryobius) caribou Ball 1962
Pterostichus (Cryobius) carradei (Gautier des Cottes 1866)
Pterostichus (Cryobius) champenoisi (Croissandeau 1893)
Pterostichus (Cryobius) colasi (Jeannel 1937)
Pterostichus (Cryobius) coreicus Jedlicka 1962
Pterostichus (Cryobius) ehlersi (Heyden 1881)
Pterostichus (Cryobius) empetricola (Dejean 1828)
Pterostichus (Cryobius) espanoli (J. & E. Vives 1977)
Pterostichus (Cryobius) euxinus Straneo 1935
Pterostichus (Cryobius) exceptus (J. Sahlberg 1885)
Pterostichus (Cryobius) firmus (Lutshnik 1933)
Pterostichus (Cryobius) fulvescens (Motschulsky 1844)
Pterostichus (Cryobius) gerstlensis Ball 1962
Pterostichus (Cryobius) glacialis (Brisout de Barneville 1863)
Pterostichus (Cryobius) glukhomanka Sundukov 2013
Pterostichus (Cryobius) herzi (Poppius 1906)
Pterostichus (Cryobius) homalonotus (Tschitscherine 1894)
Pterostichus (Cryobius) horvatovichi Kirschenhofer 1991
Pterostichus (Cryobius) infimus (Chaudoir 1868)
Pterostichus (Cryobius) inopinatus (Lutshnik 1933)
Pterostichus (Cryobius) iripennis (Chaudoir 1868)
Pterostichus (Cryobius) ishiharai Ishida & Shibata 1961
Pterostichus (Cryobius) jacobsoni (Poppius 1906)
Pterostichus (Cryobius) jaechi Kirschenhofer 1997
Pterostichus (Cryobius) josephi Csiki 1930
Pterostichus (Cryobius) kadjaranze (Jedlicka 1947)
Pterostichus (Cryobius) kaninensis (Poppius 1906)
Pterostichus (Cryobius) kaynashensis Schweiger 1967
Pterostichus (Cryobius) kolymensis Erjiomin 1998
Pterostichus (Cryobius) korgei Jedlicka 1964
Pterostichus (Cryobius) kotzebuei Ball 1962
Pterostichus (Cryobius) kurosai Tanaka 1958
Pterostichus (Cryobius) kurosawai Tanaka 1958
Pterostichus (Cryobius) lamuticus (Poppius 1906)
Pterostichus (Cryobius) latiusculus (Chaudoir 1868)
Pterostichus (Cryobius) longipes (Poppius 1906)
Pterostichus (Cryobius) lucidus (Motschulsky 1844)
Pterostichus (Cryobius) macedonicus Apfelbeck 1918
Pterostichus (Cryobius) macrothorax (Poppius 1906)
Pterostichus (Cryobius) maeklini (Poppius 1906)
Pterostichus (Cryobius) mandibularoides Ball 1966
Pterostichus (Cryobius) marani Jedlicka 1967
Pterostichus (Cryobius) middendorffi (J. Sahlberg 1875)
Pterostichus (Cryobius) negligens (Sturm 1824)
Pterostichus (Cryobius) nigripalpis (Poppius 1906)
Pterostichus (Cryobius) nivalis (R. F. Sahlberg 1844)
Pterostichus (Cryobius) nordqvisti (J. Sahlberg 1885)
Pterostichus (Cryobius) ochoticus (R. F. Sahlberg 1844)
Pterostichus (Cryobius) palekhai E. & O. Berlov 1996
Pterostichus (Cryobius) parasimilis Ball 1962
Pterostichus (Cryobius) parviceps (Poppius 1906)
Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823)
Pterostichus (Cryobius) pithyusicus (Lutshnik 1933)
Pterostichus (Cryobius) planus (J. Sahlberg 1885)
Pterostichus (Cryobius) poppiusianus (Jakobson 1907)
Pterostichus (Cryobius) procerulus (Heyden 1880)
Pterostichus (Cryobius) properans (Chaudoir 1868)
Pterostichus (Cryobius) pseudostuxbergi (Poppius 1906)
Pterostichus (Cryobius) pumilio (Dejean 1828)
Pterostichus (Cryobius) punctiger (J. Sahlberg 1880)
Pterostichus (Cryobius) quadraticollis (Chaudoir 1846)
Pterostichus (Cryobius) riparius (Dejean 1828)
Pterostichus (Cryobius) rufonitens (Fairmaire 1866)
Pterostichus (Cryobius) safonovae Budarin 1995
Pterostichus (Cryobius) scitus (Mäklin 1878)
Pterostichus (Cryobius) shilenkovi Erjiomin & Kabak 1991
Pterostichus (Cryobius) similis Mannerheim 1852
Pterostichus (Cryobius) soperi Ball 1966
Pterostichus (Cryobius) spornyi Sundukov 2013
Pterostichus (Cryobius) stantonensis Ball 1966
Pterostichus (Cryobius) subgibbus (Motschulsky 1860)
Pterostichus (Cryobius) subiasi (Ortuño & Zaballos 1992)
Pterostichus (Cryobius) surgens Le Conte 1878
Pterostichus (Cryobius) tareumiut Ball 1962
Pterostichus (Cryobius) tatricus Kult 1947
Pterostichus (Cryobius) tichomirovi Erjiomin 1990
Pterostichus (Cryobius) tiliaceoradix Ball 1962
30 additional siblings truncated for brevity.
See the resource file for a full list.

Dynamic Hierarchy Version 2.2 - Test

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Pterostichus
Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823) (this page)
Pterostichus (Agastillus) eriwanicus (Tschitscherine 1897)
Pterostichus (Aphaonus) arcanoides (Lorenz 1998)
Pterostichus (Aphaonus) cylindriformis (Reitter 1887)
Pterostichus (Argutor) cursor (Dejean 1828)
Pterostichus (Badistrinus) goschi Jedlicka 1930
Pterostichus (Bothriopterus) ferghanicus Kabak 2001
Pterostichus (Bothriopterus) oblongopunctatus (Fabricius 1787)
Pterostichus (Bothriopterus) oregonus Le Conte 1861
Pterostichus (Bothriopterus) pensylvanicus Le Conte 1873
Pterostichus (Bothriopterus) zyzzovi O. & E. Berlov 1996
Pterostichus (Circinatus) agilis Allegro & Sciaky 2010
Pterostichus (Cryobius) arcticola (Chaudoir 1868)
Pterostichus (Cryobius) asiaticus (Poppius 1906)
Pterostichus (Cryobius) glacialis (Brisout de Barneville 1863)
Pterostichus (Cryobius) korgei Jedlicka 1964
Pterostichus (Cryobius) kultianus Jedlicka 1946
Pterostichus (Cryobius) mandibularoides Ball 1966
Pterostichus (Cryobius) palekhai E. & O. Berlov 1996
Pterostichus (Cryobius) parasimilis Ball 1962
Pterostichus (Cryobius) procerulus (Heyden 1880)
Pterostichus (Cryobius) tatricus Kult 1947
Pterostichus (Eosteropus) abudarini O. & E. Berlov 1997
Pterostichus (Eosteropus) tokui Morita 2007
Pterostichus (Epinialoe) janoi Jedlicka 1952
Pterostichus (Ethira) sharanus (Straneo 1984)
Pterostichus (Eurythoracana) haptoderoides (Tschitscherine 1889)
Pterostichus (Falsargutor) pseudopedius Reitter 1887
Pterostichus (Gastrosticta) obesulus Le Conte 1873
Pterostichus (Gastrosticta) putus Casey 1913
Pterostichus (Haplomaseus) andreae (Tschitscherine 1897)
Pterostichus (Haplomaseus) armenus (Faldermann 1836)
Pterostichus (Haplomaseus) caucasicus (Ménétriés 1832)
Pterostichus (Haptotapinus) vseteckai (Maran 1944)
Pterostichus (Hypherpes) annosus Casey 1913
Pterostichus (Hypherpes) ecarinatus Hatch 1936
Pterostichus (Hypherpes) scutellaris Le Conte 1873
Pterostichus (Hypherpes) setosus Hatch 1951
Pterostichus (Koreonialoe) teretis Park & Kwon 1996
Pterostichus (Lenapterus) agonus G. Horn 1880
Pterostichus (Lenapterus) punctatissimus (Randall 1838)
Pterostichus (Leptoferonia) falli Van Dyke 1926
Pterostichus (Leptoferonia) hatchi Hacker 1968
Pterostichus (Leptoferonia) infernalis Hatch 1936
Pterostichus (Leptoferonia) stapedius Hacker 1968
Pterostichus (Leptoferonia) yosemitensis Hacker 1968
Pterostichus (Lianoe) drescoi Nègre 1957
Pterostichus (Lianoe) tanakai Ishida 1964
Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida 1972
Pterostichus (Melanius) aterrimus (Herbst 1784)
Pterostichus (Metallophilus) rugosus (Gebler 1823)
Pterostichus (Morphohaptoderus) hubeicus Facchini & Sciaky 2003
Pterostichus (Morphohaptoderus) yulongshanensis Sciaky 1997
Pterostichus (Neohaptoderus) jureceki (Jedlicka 1936)
Pterostichus (Neohaptoderus) yunnanensis Jedlicka 1934
Pterostichus (Nialoe) ohbayashii Ishida 1968
Pterostichus (Nialoe) okutamae Tanaka 1963
Pterostichus (Nialoe) opacipennis Jedlicka 1934
Pterostichus (Nialoe) shinbodakensis Sasakawa & H. Ito 2017
Pterostichus (Nialoe) taoi Kasahara 1993
Pterostichus (Nialoe) yakushimanus Nakane & Ishida 1961
Pterostichus (Oreophilus) morio (Duftschmid 1812)
Pterostichus (Oreophilus) spinolae (Dejean 1828)
Pterostichus (Oreoplatysma) abishirensis Belousov 1991
Pterostichus (Oreoplatysma) casaleianus Kirschenhofer 1991
Pterostichus (Oreoplatysma) kvirensis Belousov 1991
Pterostichus (Oreoplatysma) mediocris Dvorak 1995
Pterostichus (Oreoplatysma) orman Jedlicka 1963
Pterostichus (Oreoplatysma) rousianus Kirschenhofer 1987
Pterostichus (Oreoplatysma) satyrus Kurnakov 1962
Pterostichus (Paranialoe) biexcisus Straneo 1955
Pterostichus (Petrophilus) alexandrovi Lafer 1979
Pterostichus (Petrophilus) arsenjevi Lafer 1979
Pterostichus (Petrophilus) pertinax (Tschitscherine 1895)
Pterostichus (Petrophilus) tomensis (Gebler 1847)
Pterostichus (Phonias) diligens (Sturm 1824)
Pterostichus (Phonias) gyrosus (Motschulsky 1866)
Pterostichus (Platysma) sachtlebeni Jedlicka 1962
Pterostichus (Pseudethira) gobettii Straneo 1983
Pterostichus (Pseudethira) immarginatus Straneo 1977
Pterostichus (Pseudethira) matsumurai Habu 1973
Pterostichus (Pseudethira) miehei J. Schmidt 2005
Pterostichus (Pseudethira) weiperti J. Schmidt 2005
Pterostichus (Pseudomaseus) aimaki Jedlicka 1968
Pterostichus (Pseudomaseus) rhaeticus Heer 1837
Pterostichus (Pseudomaseus) tenuis (Casey 1924)
Pterostichus (Pterostichus) auratus Heer 1837
Pterostichus (Pterostichus) devillei Puel 1924
Pterostichus (Pterostichus) esakii Ishida 1959
Pterostichus (Pterostichus) pedemontanus Ganglbauer 1891
Pterostichus (Pterostichus) ruffoi Sciaky 1984
Pterostichus (Rhagadus) modicellus (Tschitscherine 1897)
Pterostichus (Sinosteropus) jani Sciaky & Facchini 2003
Pterostichus (Steropus) catalonicus J. Daniel ex K. & J. Daniel 1906
Pterostichus (Tschitscherinea) mulensis Sciaky 1997
Pterostichus bischoffianus Jedlicka 1936
Pterostichus exochus Andrewes 1930
Pterostichus nishidai Nakane 1989
Pterostichus pentheri Apfelbeck 1918
Pterostichus punctisternus Straneo 1989
Pterostichus umbralis Gistel 1857
1082 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Pterostichus
Pterostichus (Cryobius) pinguedineus (Eschscholtz 1823) (this page)
Pterostichus (Agastillus) eriwanicus (Tschitscherine 1897)
Pterostichus (Aphaonus) arcanoides (Lorenz 1998)
Pterostichus (Aphaonus) cylindriformis (Reitter 1887)
Pterostichus (Argutor) cursor (Dejean 1828)
Pterostichus (Badistrinus) goschi Jedlicka 1930
Pterostichus (Bothriopterus) ferghanicus Kabak 2001
Pterostichus (Bothriopterus) oblongopunctatus (Fabricius 1787)
Pterostichus (Bothriopterus) oregonus Le Conte 1861
Pterostichus (Bothriopterus) pensylvanicus Le Conte 1873
Pterostichus (Bothriopterus) zyzzovi O. & E. Berlov 1996
Pterostichus (Circinatus) agilis Allegro & Sciaky 2010
Pterostichus (Cryobius) arcticola (Chaudoir 1868)
Pterostichus (Cryobius) asiaticus (Poppius 1906)
Pterostichus (Cryobius) glacialis (Brisout de Barneville 1863)
Pterostichus (Cryobius) korgei Jedlicka 1964
Pterostichus (Cryobius) kultianus Jedlicka 1946
Pterostichus (Cryobius) mandibularoides Ball 1966
Pterostichus (Cryobius) palekhai E. & O. Berlov 1996
Pterostichus (Cryobius) parasimilis Ball 1962
Pterostichus (Cryobius) procerulus (Heyden 1880)
Pterostichus (Cryobius) tatricus Kult 1947
Pterostichus (Eosteropus) abudarini O. & E. Berlov 1997
Pterostichus (Eosteropus) tokui Morita 2007
Pterostichus (Epinialoe) janoi Jedlicka 1952
Pterostichus (Ethira) sharanus (Straneo 1984)
Pterostichus (Eurythoracana) haptoderoides (Tschitscherine 1889)
Pterostichus (Falsargutor) pseudopedius Reitter 1887
Pterostichus (Gastrosticta) obesulus Le Conte 1873
Pterostichus (Gastrosticta) putus Casey 1913
Pterostichus (Haplomaseus) andreae (Tschitscherine 1897)
Pterostichus (Haplomaseus) armenus (Faldermann 1836)
Pterostichus (Haplomaseus) caucasicus (Ménétriés 1832)
Pterostichus (Haptotapinus) vseteckai (Maran 1944)
Pterostichus (Hypherpes) annosus Casey 1913
Pterostichus (Hypherpes) ecarinatus Hatch 1936
Pterostichus (Hypherpes) scutellaris Le Conte 1873
Pterostichus (Hypherpes) setosus Hatch 1951
Pterostichus (Koreonialoe) teretis Park & Kwon 1996
Pterostichus (Lenapterus) agonus G. Horn 1880
Pterostichus (Lenapterus) punctatissimus (Randall 1838)
Pterostichus (Leptoferonia) falli Van Dyke 1926
Pterostichus (Leptoferonia) hatchi Hacker 1968
Pterostichus (Leptoferonia) infernalis Hatch 1936
Pterostichus (Leptoferonia) stapedius Hacker 1968
Pterostichus (Leptoferonia) yosemitensis Hacker 1968
Pterostichus (Lianoe) drescoi Nègre 1957
Pterostichus (Lianoe) tanakai Ishida 1964
Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida 1972
Pterostichus (Melanius) aterrimus (Herbst 1784)
Pterostichus (Metallophilus) rugosus (Gebler 1823)
Pterostichus (Morphohaptoderus) hubeicus Facchini & Sciaky 2003
Pterostichus (Morphohaptoderus) yulongshanensis Sciaky 1997
Pterostichus (Neohaptoderus) jureceki (Jedlicka 1936)
Pterostichus (Nialoe) ohbayashii Ishida 1968
Pterostichus (Nialoe) okutamae Tanaka 1963
Pterostichus (Nialoe) opacipennis Jedlicka 1934
Pterostichus (Nialoe) shinbodakensis Sasakawa & H. Ito 2017
Pterostichus (Nialoe) taoi Kasahara 1993
Pterostichus (Nialoe) yakushimanus Nakane & Ishida 1961
Pterostichus (Oreophilus) morio (Duftschmid 1812)
Pterostichus (Oreophilus) spinolae (Dejean 1828)
Pterostichus (Oreoplatysma) abishirensis Belousov 1991
Pterostichus (Oreoplatysma) casaleianus Kirschenhofer 1991
Pterostichus (Oreoplatysma) kvirensis Belousov 1991
Pterostichus (Oreoplatysma) mediocris Dvorak 1995
Pterostichus (Oreoplatysma) orman Jedlicka 1963
Pterostichus (Oreoplatysma) rousianus Kirschenhofer 1987
Pterostichus (Oreoplatysma) satyrus Kurnakov 1962
Pterostichus (Paranialoe) biexcisus Straneo 1955
Pterostichus (Petrophilus) alexandrovi Lafer 1979
Pterostichus (Petrophilus) arsenjevi Lafer 1979
Pterostichus (Petrophilus) pertinax (Tschitscherine 1895)
Pterostichus (Petrophilus) tomensis (Gebler 1847)
Pterostichus (Phonias) diligens (Sturm 1824)
Pterostichus (Phonias) gyrosus (Motschulsky 1866)
Pterostichus (Platysma) sachtlebeni Jedlicka 1962
Pterostichus (Pseudethira) gobettii Straneo 1983
Pterostichus (Pseudethira) immarginatus Straneo 1977
Pterostichus (Pseudethira) matsumurai Habu 1973
Pterostichus (Pseudethira) miehei J. Schmidt 2005
Pterostichus (Pseudethira) weiperti J. Schmidt 2005
Pterostichus (Pseudomaseus) aimaki Jedlicka 1968
Pterostichus (Pseudomaseus) rhaeticus Heer 1837
Pterostichus (Pseudomaseus) tenuis (Casey 1924)
Pterostichus (Pterostichus) auratus Heer 1837
Pterostichus (Pterostichus) devillei Puel 1924
Pterostichus (Pterostichus) esakii Ishida 1959
Pterostichus (Pterostichus) pedemontanus Ganglbauer 1891
Pterostichus (Pterostichus) ruffoi Sciaky 1984
Pterostichus (Rhagadus) modicellus (Tschitscherine 1897)
Pterostichus (Sinosteropus) jani Sciaky & Facchini 2003
Pterostichus (Steropus) catalonicus J. Daniel ex K. & J. Daniel 1906
Pterostichus (Tschitscherinea) mulensis Sciaky 1997
Pterostichus bischoffianus Jedlicka 1936
Pterostichus exochus Andrewes 1930
Pterostichus nishidai Nakane 1989
Pterostichus pentheri Apfelbeck 1918
Pterostichus punctisternus Straneo 1989
Pterostichus umbralis Gistel 1857
Pterostichus vetustus Heer 1862
1100 additional siblings truncated for brevity.
See the resource file for a full list.

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Pterostichus Bonelli 1810
Pterostichus pinguedineus (Eschscholtz 1823) (this page)
Pterostichus adulterinus Wrase & J. Schmidt 2006
Pterostichus alexeji Zamotajlov & Kryzhanovskij 1992
Pterostichus altaiensis (Poppius 1906)
Pterostichus amagisanus Tanaka & Ishida 1972
Pterostichus amanoi Nakane 1968
Pterostichus amoenus (Dejean 1828)
Pterostichus andreae (Tschitscherine 1897)
Pterostichus angoarnigi Morvan 1994
Pterostichus apenninus (Dejean 1831)
Pterostichus arcticola (Chaudoir 1868)
Pterostichus arrowi Jedlicka 1936
Pterostichus asymmetricus Bates 1883
Pterostichus aubryi Coiffait 1952
Pterostichus auratus Heer 1837
Pterostichus balthasari Jedlicka 1937
Pterostichus barbarinus Casey 1913
Pterostichus bifoveolatus Park, Kwon & Lafer 1996
Pterostichus bisetosus (Straneo 1938)
Pterostichus bituberculatus (Tschitscherine 1899)
Pterostichus brunneipennis Straneo 1955
Pterostichus capitolinus Kurnakov 1962
Pterostichus cervenkai Sciaky 1994
Pterostichus crassiusculus (Chaudoir 1868)
Pterostichus craterensis (Hatch 1949)
Pterostichus cylindricus (Herbst 1784)
Pterostichus daisenicus Ishida 1958
Pterostichus dentifer Allegro & Sciaky 2010
Pterostichus dhorpatanicus Straneo 1977
Pterostichus dolens (Tschitscherine 1900)
Pterostichus dostali Kirschenhofer 1981
Pterostichus ebeninus (Dejean 1828)
Pterostichus empetricola (Dejean 1828)
Pterostichus faldermanni Schatzmayr 1929
Pterostichus fenestratus Kasahara & Ohtani 1989
Pterostichus firmus (Lutshnik 1933)
Pterostichus foveolatus (Duftschmid 1812)
Pterostichus galae Farkac & Plutenko 1996
Pterostichus gerstlensis Ball 1962
Pterostichus gravis Jedlicka 1939
Pterostichus hagenbachii Sturm 1824
Pterostichus harmandi (Tschitscherine 1900)
Pterostichus hirobane Habu & Baba 1958
Pterostichus iripennis (Chaudoir 1868)
Pterostichus jurecekianus Maran 1940
Pterostichus kambaiti (Andrewes 1947)
Pterostichus katashinensis Habu 1958
Pterostichus kitakamisanus Sasakawa 2005
Pterostichus lachrymosus (Newman 1838)
Pterostichus lacunosus (Chaudoir 1844)
Pterostichus laferi O. & E. Berlov 1996
Pterostichus lanei Van Dyke 1926
Pterostichus liodactylus (Tschitscherine 1898)
Pterostichus longipes (Poppius 1906)
Pterostichus macer (Marsham 1802)
Pterostichus macrogenys Bates 1883
Pterostichus macrothorax (Poppius 1906)
Pterostichus masidai Ishida 1959
Pterostichus microbus Sciaky & Facchini 2003
Pterostichus multiseta (Straneo 1984)
Pterostichus nigellus A. Morawitz 1862
Pterostichus nigrocaeruleus Van Dyke 1926
Pterostichus nordqvisti (J. Sahlberg 1887)
Pterostichus ohsawacavus Sasakawa 2005
Pterostichus olafi J. Schmidt 2005
Pterostichus pfizenmayeri Poppius 1906
Pterostichus planctus Le Conte 1853
Pterostichus protensiformis (Casey 1924)
Pterostichus pseudopercus (Reitter 1889)
Pterostichus pseudoplatyderus Sciaky 1994
Pterostichus pseudosinensis Sciaky & Facchini 2003
Pterostichus putus Casey 1913
Pterostichus rolex Morvan 1995
Pterostichus rousianus Kirschenhofer 1987
Pterostichus sacheri I. Frivaldszky von Frivald 1865
Pterostichus scalptus Sciaky & Wrase 1997
Pterostichus schmidtii (Chaudoir 1838)
Pterostichus schneideri Sciaky & Facchini 2003
Pterostichus scuticollis (Fairmaire 1889)
Pterostichus serripes (Le Conte 1875)
Pterostichus seunglaki Park & Kwon 1996
Pterostichus shaovuensis Jedlicka 1956
Pterostichus skoupyi Hovorka & Skoupy 2007
Pterostichus stantonensis Ball 1966
Pterostichus stygicus (Say 1823)
Pterostichus styx (Andrewes 1937)
Pterostichus subiasi (Ortuño & Zaballos 1992)
Pterostichus swaneticus (Reitter 1883)
Pterostichus tamarae Wrase & Kirschenhofer 1991
Pterostichus tatricus Kult 1947
Pterostichus tenuimarginatus (Chaudoir 1868)
Pterostichus tiliaceoradix Ball 1962
Pterostichus triangularis Sciaky & Facchini 2003
Pterostichus trinarius (Casey 1918)
Pterostichus triseriatus (Gebler 1847)
Pterostichus tuberculofemoratus Hatch 1936
Pterostichus validiceps Reitter 1887
Pterostichus variabilis (Ménétriés 1832)
Pterostichus vicinus Mannerheim 1843
Pterostichus weigeli J. Schmidt 2005
Pterostichus zyzzovi O. & E. Berlov 1996
1090 additional siblings truncated for brevity.
See the resource file for a full list.

Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Coleoptera Linnaeus 1758
Adephaga Schellenberg 1806
Carabidae Latreille 1802
Harpalinae Bonelli 1810
Pterostichini Bonelli 1810
Pterostichus Bonelli 1810
Pterostichus subgen. Criobius Chaudoir 1838
Pterostichus pinguedineus (Eschscholtz 1823) (this page)
Pterostichus arcticola (Chaudoir 1868)
Pterostichus auriga Ball 1962
Pterostichus barryorum Ball 1962
Pterostichus brevicornis (Kirby 1837)
Pterostichus bryanti (Van Dyke 1951)
Pterostichus caribou Ball 1962
Pterostichus chipewyan Ball 1962
Pterostichus empetricola (Dejean 1828)
Pterostichus gerstlensis Ball 1962
Pterostichus hudsonicus Le Conte 1863
Pterostichus kotzebuei Ball 1962
Pterostichus mandibularoides Ball 1966
Pterostichus nivalis (R. Sahlberg 1844)
Pterostichus parasimilis Ball 1962
Pterostichus planus (J. Sahlberg 1885)
Pterostichus riparius (Dejean 1828)
Pterostichus similis Mannerheim 1852
Pterostichus soperi Ball 1966
Pterostichus surgens Le Conte 1878
Pterostichus tareumiut Ball 1962
Pterostichus ventricosus (Eschscholtz 1823)
Pterostichus woodi Ball & Currie 1997

NCBI

root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Adephaga
Caraboidea
Carabidae
Pterostichinae
Pterostichus
Cryobius
Pterostichus pinguedineus (this page)
Pterostichus barryorum
Pterostichus brevicornis
Pterostichus caribou
Pterostichus empetricola
Pterostichus kurosawai
Pterostichus negligens
Pterostichus nemoralis
Pterostichus nivalis
Pterostichus pumilio
Pterostichus riparius
Pterostichus subsinuatus
Pterostichus surgens
Pterostichus ventricosus

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Adephaga
Caraboidea
Carabidae
Harpalinae
Pterostichini
Poecilus
Poecilus
Poecilus pinguedineus (this page)
Poecilus tyrrhenicus
Poecilus ventricosus

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Adephaga
Caraboidea
Carabidae
Harpalinae
Pterostichini
Pterostichus
Pterostichus pinguedineus (this page)
Pterostichus aanistschenkoi
Pterostichus aapsorum
Pterostichus abaciformis
Pterostichus abagonensis
Pterostichus abasarukini
Pterostichus abasinus
Pterostichus abaxoides
Pterostichus abensis
Pterostichus abishirensis
Pterostichus abnormis
Pterostichus abudarini
Pterostichus acuspina
Pterostichus acutidens
Pterostichus acutipes
Pterostichus adatarasanus
Pterostichus ademidovae
Pterostichus adoxus
Pterostichus adstrictus
Pterostichus adulterinus
Pterostichus aemiliae
Pterostichus aeneocupreus
Pterostichus aenescens
Pterostichus aeneus
Pterostichus aereipennis
Pterostichus agilis
Pterostichus agonus
Pterostichus aibgensis
Pterostichus aimaki
Pterostichus ainus
Pterostichus akitai
Pterostichus akkusianus
Pterostichus akozyrevi
Pterostichus aksekianus
Pterostichus alacer
Pterostichus alamedae
Pterostichus alexandrovi
Pterostichus alexeji
Pterostichus algidus
Pterostichus altaianus
Pterostichus altaicus
Pterostichus altaiensis
Pterostichus amagisanus
Pterostichus amanoi
Pterostichus amaroides
Pterostichus ambigenus
Pterostichus ambiguus
Pterostichus amethystinus
Pterostichus amoenus
Pterostichus amurensis
Pterostichus analis
Pterostichus anatolicus
Pterostichus andreae
Pterostichus andreinii
Pterostichus andrewesi
Pterostichus angoarnigi
Pterostichus angustus
Pterostichus annosus
Pterostichus anomostriatus
Pterostichus apenninus
Pterostichus apfelbecki
Pterostichus apiculatiphallus
Pterostichus aralarensis
Pterostichus araraticus
Pterostichus arator
Pterostichus aratus
Pterostichus arcanoides
Pterostichus arcanus
Pterostichus archangaicus
Pterostichus arcticola
Pterostichus arcuaticarinatus
Pterostichus armenus
Pterostichus arnoldii
Pterostichus arribasi
Pterostichus arrowianus
Pterostichus arvernus
Pterostichus asahinus
Pterostichus asiaticus
Pterostichus asturicus
Pterostichus astutus
Pterostichus asymmetricus
Pterostichus aterrimus
Pterostichus atratus
Pterostichus atrox
Pterostichus aubryi
Pterostichus audax
Pterostichus auratus
Pterostichus auriga
Pterostichus avaricus
Pterostichus baenningeri
Pterostichus blandus
Pterostichus calcaratus
Pterostichus diligens
Pterostichus minor
Pterostichus nigrita
Pterostichus oblongopunctatus
Pterostichus obscuripennis
Pterostichus parvulus
Pterostichus strenuus
Pterostichus subgen. Morphohaptoderus
Pterostichus vernalis
1308 additional siblings truncated for brevity.
See the resource file for a full list.

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Adephaga
Caraboidea
Carabidae
Harpalinae
Pterostichini
Pterostichus
Pterostichus mandibularis (this page)
Pterostichus aanistschenkoi
Pterostichus aapsorum
Pterostichus abaciformis
Pterostichus abagonensis
Pterostichus abasarukini
Pterostichus abasinus
Pterostichus abaxoides
Pterostichus abensis
Pterostichus abishirensis
Pterostichus abnormis
Pterostichus abudarini
Pterostichus acuspina
Pterostichus acutidens
Pterostichus acutipes
Pterostichus adatarasanus
Pterostichus ademidovae
Pterostichus adoxus
Pterostichus adstrictus
Pterostichus adulterinus
Pterostichus aemiliae
Pterostichus aeneocupreus
Pterostichus aenescens
Pterostichus aeneus
Pterostichus aereipennis
Pterostichus agilis
Pterostichus agonus
Pterostichus aibgensis
Pterostichus aimaki
Pterostichus ainus
Pterostichus akitai
Pterostichus akkusianus
Pterostichus akozyrevi
Pterostichus aksekianus
Pterostichus alacer
Pterostichus alamedae
Pterostichus alexandrovi
Pterostichus alexeji
Pterostichus algidus
Pterostichus altaianus
Pterostichus altaicus
Pterostichus altaiensis
Pterostichus amagisanus
Pterostichus amanoi
Pterostichus amaroides
Pterostichus ambigenus
Pterostichus ambiguus
Pterostichus amethystinus
Pterostichus amoenus
Pterostichus amurensis
Pterostichus analis
Pterostichus anatolicus
Pterostichus andreae
Pterostichus andreinii
Pterostichus andrewesi
Pterostichus angoarnigi
Pterostichus angustus
Pterostichus annosus
Pterostichus anomostriatus
Pterostichus apenninus
Pterostichus apfelbecki
Pterostichus apiculatiphallus
Pterostichus aralarensis
Pterostichus araraticus
Pterostichus arator
Pterostichus aratus
Pterostichus arcanoides
Pterostichus arcanus
Pterostichus archangaicus
Pterostichus arcticola
Pterostichus arcuaticarinatus
Pterostichus armenus
Pterostichus arnoldii
Pterostichus arribasi
Pterostichus arrowianus
Pterostichus arvernus
Pterostichus asahinus
Pterostichus asiaticus
Pterostichus asturicus
Pterostichus astutus
Pterostichus asymmetricus
Pterostichus aterrimus
Pterostichus atratus
Pterostichus atrox
Pterostichus aubryi
Pterostichus audax
Pterostichus auratus
Pterostichus auriga
Pterostichus avaricus
Pterostichus baenningeri
Pterostichus blandus
Pterostichus calcaratus
Pterostichus diligens
Pterostichus minor
Pterostichus nigrita
Pterostichus oblongopunctatus
Pterostichus obscuripennis
Pterostichus parvulus
Pterostichus strenuus
Pterostichus subgen. Morphohaptoderus
Pterostichus vernalis
1308 additional siblings truncated for brevity.
See the resource file for a full list.
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