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Acritarchs are organic microfossils, known from approximately 1800 million years ago to the present. The classification is a catch all term used to refer to any organic microfossils that cannot be assigned to other groups. Their diversity reflects major ecological events such as the appearance of predation and the Cambrian explosion.
Acritarchs were originally defined as non-acid soluble (i.e. non-carbonate, non-siliceous) organic-walled microfossils consisting of a central cavity, and whose biological affinities cannot be determined with certainty.[2][3][4] Most commonly they are composed of thermally altered acid insoluble carbon compounds (kerogen).
Acritarchs may include the remains of a wide range of quite different kinds of organisms—ranging from the egg cases of small metazoans to resting cysts of many kinds of chlorophyta (green algae). It is likely that most acritarch species from the Paleozoic represent various stages of the life cycle of algae that were ancestral to the dinoflagellates.[5] The nature of the organisms associated with older acritarchs is generally not well understood, though many are probably related to unicellular marine algae. In theory, when the biological source (taxon) of an acritarch does become known, that particular microfossil is removed from the acritarchs and classified with its proper group.
While the classification of acritarchs into form genera is entirely artificial, it is not without merit, as the form taxa show traits similar to those of genuine taxa—for example an 'explosion' in the Cambrian and a mass extinction at the end of the Permian.
Acritarchs were most likely eukaryotes. While archaea and bacteria (prokaryotes) usually produce simple fossils of a very small size, eukaryotic unicellular fossils are usually larger and more complex, with external morphological projections and ornamentation such as spines and hairs that only eukaryotes can produce; as most acritarchs have external projections (e.g., hair, spines, thick cell membranes, etc.), they are predominantly eukaryotes, although simple eukaryote acritarchs also exist.[6]
The recent application of atomic force microscopy, confocal microscopy, Raman spectroscopy, and other sophisticated analytic techniques to the study of the ultrastructure, life history, and systematic affinities of mineralized, but originally organic-walled microfossils,[7][8][9][10][11] has shown that some acritarchs are actually fossilized microalgae. In the end, it may well be, as Moczydłowska et al. suggested in 2011, that many acritarchs will, in fact, turn out to be algae.[12][13]
Acritarchs are found in sedimentary rocks from the present back into the Archean.[14] They are typically isolated from siliciclastic sedimentary rocks using hydrofluoric acid but are occasionally extracted from carbonate-rich rocks. They are excellent candidates for index fossils used for dating rock formations in the Paleozoic Era and when other fossils are not available. Because most acritarchs are thought to be marine (pre-Triassic), they are also useful for palaeoenvironmental interpretation.
The Archean and earliest Proterozoic microfossils termed "acritarchs" may actually be prokaryotes. The earliest eukaryotic acritarchs known (as of 2020) are from between 1950 and 2150 million years ago.[15]
At about 1 billion years ago the organisms responsible for acritarchs started to increase in abundance, diversity, size, complexity of shape and especially size and number of spines. Their populations crashed during periods of extensive worldwide glaciations that covered the majority of the planet, but they proliferated in the Cambrian explosion and reached their highest diversity in the Paleozoic. The increased spininess 1000 million years ago possibly resulted from the need for defense against predators, especially predators large enough to swallow them or tear them apart. Other groups of small organisms from the Neoproterozoic era also show signs of anti-predator defenses.[16]
Further evidence that acritarchs were subject to herbivory around this time comes from a consideration of taxon longevity. The abundance of planktonic organisms that evolved between 1,700 and 1,400 million years ago was limited by nutrient availability – a situation which limits the origination of new species because the existing organisms are so specialised to their niches, and no other niches are available for occupation. Approximately 1,000 million years ago, species longevity fell sharply, suggesting that predation pressure, probably by protist herbivores, became an important factor. Predation would have kept populations in check, meaning that some nutrients were left unused, and new niches were available for new species to occupy.[17]
Acritarch was coined in 1963 from the Greek ákritos meaning confused (a kritēs, without critic) and archē meaning origin (confer archaic).[18]
This is a list of genera according to Fossilid.info.[19]
Material was copied from this source, which is available under a Creative Commons Attribution 4.0 International License. 
Acritarch from the Weng'an biotaAcritarchs are organic microfossils, known from approximately 1800 million years ago to the present. The classification is a catch all term used to refer to any organic microfossils that cannot be assigned to other groups. Their diversity reflects major ecological events such as the appearance of predation and the Cambrian explosion.
Les acritarches sont des microfossiles à paroi organique, c’est-à-dire des palynomorphes, auxquels il n'est pas possible d'attribuer une affinité biologique avec certitude (on les regroupe dans les Biota incertae sedis).
Le nom acritarche dérive du grec akritos, signifiant « incertain ou confus », et de arche, signifiant « origine », « ce qui est premier ». Le terme a été introduit pour la première fois par W.R. Evitt en 1963[1].
On pourrait caricaturer les acritarches comme étant tous les palynomorphes qui ont été laissés de côté dans les préparations palynologiques (lames qu'étudient au microscope les spécialistes en palynologie) après avoir réalisé toutes les autres identifications. Il est certain que différents organismes sont regroupés sous le terme acritarche, qui est donc un groupe polyphylétique. Certains sont probablement des kystes de dinoflagellés trop mal conservés pour être correctement identifiés, d'autres appartiennent sans aucun doute aux algues vertes et encore d'autres ressemblent fortement à certaines prasinophycées. Les acritarches sont connus depuis le Précambrien (les plus anciens connus sont datés de 1,87 milliard d'années), ils sont abondants au cours du Paléozoïque et puis régressent très fortement et disparaissent presque complètement par la suite.
La grande majorité des acritarches sont trouvés dans des sédiments de rivières et/ou associés à des organismes aquatiques. Ils ont été découverts par Manon[Qui ?] au XVIe. Par ailleurs d'après leur morphologie, leur distribution, et leur composition, la plupart des acritarches sont supposés être du phytoplancton et étaient donc à la base de la chaîne alimentaire de l'écosystème marin durant le Protérozoïque et le Paléozoïque.
Deunff (1954) décrit Veryhachium comme des « microorganismes planctoniques fossiles conservés à l'état de matière organique, le plus souvent de couleur jaune ou brune parfois noire, coque à paroi lisse ou granuleuse. Forme générale dépendant du nombre de processus (de 1 à 8), non globuleuse, sauf lorsqu’il n’y a qu'une seule corne. Processus presque toujours pointus, généralement longs ou très longs, souvent recourbés. Diamètre du corps de la coque 10 à 40 µm, rarement inférieur ou supérieur. Envergure totale pouvant atteindre 100 µm et plus ».
Plusieurs centaines d’espèces ont été décrites depuis 1954. Elles ont été attribuées à différents groupes morphologiques. Servais et al. (2007) distinguent deux groupes dans l’Ordovicien inférieur, Veryhachium trispinosum de forme triangulaire, et Veryhachium lairdii, de forme rectangulaire[2].
Les acritarches sont des microfossiles à paroi organique, c’est-à-dire des palynomorphes, auxquels il n'est pas possible d'attribuer une affinité biologique avec certitude (on les regroupe dans les Biota incertae sedis).
