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Description ( Inglês )

fornecido por Flora of Zimbabwe
Evergreen shrubs. Leaves alternate. Inflorescence a dense, terminal head, surrounded by coloured bracts. Flowers bisexual, zygomorphic; perianth tube open on lower side; anthers sessile. Fruit a leathery follicle. Seeds winged.
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Mark Hyde, Bart Wursten and Petra Ballings
citação bibliográfica
Hyde, M.A., Wursten, B.T. and Ballings, P. (2002-2014). Telopea Flora of Zimbabwe website. Accessed 28 August 2014 at http://www.zimbabweflora.co.zw/cult/genus.php?genus_id=2123
autor
Mark Hyde
autor
Bart Wursten
autor
Petra Ballings
original
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site do parceiro
Flora of Zimbabwe

Telopea ( Alemão )

fornecido por wikipedia DE

Telopea ist eine Pflanzengattung in der Familie der Silberbaumgewächse (Proteaceae) mit fünf Arten.

Beschreibung

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Balgfrüchte mit mehreren beigen Samen von Telopea speciosissima.

Telopea-Arten sind immergrüne Sträucher oder selten kleine Bäume, die Wuchshöhen von 3 bis 5 Meter[1] erreichen und meist mehrere Stämme aufweisen. Die spiralig angeordneten Laubblätter sind an jungen und älteren Pflanzen unterschiedlich geformt. An jungen Pflanzen sind die Laubblätter einfach mit glattem oder gezahntem Blattrand, an älteren Pflanzen sind sie einfach aber mit gewellten bis gelappten Rändern. Die Laubblätter sind 10 bis 20 cm lang und 2 bis 3 cm breit.

Die endständigen Gesamtblütenstände sind dicht, ei- oder kugelförmig, weisen einen Durchmesser von 6 bis 15 cm und einen Ring von rötlichen Tragblättern auf; sie enthalten bis zu 250 Einzelblüten. Die Gesamtblütenstände bestehen aus pseudo-traubigen Teilblütenständen, in denen die Blüten paarweise zusammenstehen. Die zygomorphen Blüten sind rot. Die Bestäubung erfolgt durch Vögel. In der Knospe sind die Blütenhüllblätter stark eingekrümmt. Es ist ein hufeisenförmiges Nektarium vorhanden.

Die Balgfrüchte sind verholzt. Die Samen sind geflügelt.

Systematik und Verbreitung

Das natürliche Vorkommensgebiet befindet sich im Südosten Australiens: in New South Wales, Victoria und Tasmanien.

Innerhalb der Silberbaumgewächse scheinen die nächsten Verwandten die Gattungen Alloxylon, Oreocallis und Embothrium zu sein, die alle rot-blühende Pflanzen am Südpazifik Australiens sind. Zusammen bilden sie die Untertribus der Embothriinae aus der Tribus Embothrieae in der Unterfamilie Grevilleoideae innerhalb der Familie der Proteaceae.

Die Gattung Telopea wurde 1810 von Robert Brown in On the natural order of plants called Proteaceae. in Transactions of the Linnean Society of London, 10(1), S. 197 ohne Typusart erstveröffentlicht. Im International Code of Botanical Nomenclature (I.C.B.N.), Berlin, 1988 wurde durch Werner Greuter: Regnum Vegetabile 118: 191 ein Lectotypus: Telopea speciosissima (Sm.) R.Br. festgelegt und der Gattungsname Telopea konserviert, wobei der 1809 veröffentlichte Name Hylogyne Salisb. ex Knight zurückgewiesen wurde.[2]

Die fünf Telopea-Arten haben Verbreitungsgebiete, die sich nicht oder nur minimal überlappen; von Norden nach Süden sind das:

Kultivierung

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Eine weiße Sorte.

Die New South Wales Waratah (Telopea speciosissima) stammt aus der Gegend des Sydneybeckens, der Central Coast, der Südküste von New South Wales und den Blue Mountains; sie erreicht eine Wuchshöhe von bis zu 4 Meter. Telopea speciosissima wächst üblicherweise auf sandigen Lehmböden entlang der Ränder von Plateaus. Diese Telopea-Art ist in New South Wales heimisch; sie ist aber wegen ihrer Popularität als Zierpflanze auch in Victoria und Tasmanien verbreitet.

Telopea-Sorten sind beliebt als Zierpflanzen in australischen Gärten, allerdings etwas schwierig in der Pflege. Verschiedene Hybride und Cultivare wurden gezüchtet, darunter auch einige creme-weiße und rosa Blumen sowie mit natürlichen Rot. Weiße Sorten der Telopea speciosissima heißen 'Telopea Wirrimbirra White' und 'T. Shady Lady White'. Dagegen sind 'T. Shady Lady Pink' und 'T. Shady Lady Red' Hybride von Telopea speciosissima mit Telopea oreades.

Kulturelle Bedeutung

Der im Englischen verwendete Trivialname „Waratah“ stammt von Aborigines vom Stamm der Eora, den ursprünglichen Bewohnern der Gegend von Sydney. In der Mythologie der Aborigines des Südwesten hat die Waratah vielfachen Eingang gefunden und wird auch als Totem gepflegt.[4]

Die botanische Zeitschrift Telopea ist nach dieser Gattung benannt, ebenso der Vorort Sydneys mit dem Namen Telopea (Sydney).

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Die Staatsblume auf einem Schild am Ortseingang von Mulwala, einer Grenzstadt im Süden New South Wales’.

Telopea speciosissima ist die Staatsblume des Bundesstaats New South Wales und verschiedener Organisationen im Bundesstaat, wie zum Beispiel des Rugbyteams New South Wales Waratahs.[5]

Literatur

  • M. D. Crisp & P. H. Weston: Telopea - New South Wales Flora Online. (Abschnitt Beschreibung) zugegriffen am 12. Juni 2010
  • M.D.Crisp & P.H.Weston: Telopea. In: McCarthy, Patrick (Hrsg.): Flora of Australia: Volume 16: Eleagnaceae, Proteaceae 1. CSIRO Publishing / Australian Biological Resources Study, 1995, ISBN 0-643-05693-9, S. 382–390.

Einzelnachweise

  1. Thomas H. Everett: The New York Botanical Garden illustrated encyclopedia of horticulture, 1982, Seite 3310: The New York Botanical Garden illustrated encyclopedia of horticulture – Google-Book Online.
  2. Australian Plant Name Index (APNI).
  3. Telopea truncata - Flora of Tasmania (PDF; 62 kB).
  4. Sue Wesson (2005) Murni Dhugang Jirrar: Living in the Illawarra (PDF; 7,2 MB) Department of Environment, Climate Change, and Water, State Government of New South Wales, Seite 7, zugegriffen am 22. Februar 2010
  5. Waratah (Telopea speciosissima) - Floral Emblems of Australia von Australian National Botanic Gardens.
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Autoren und Herausgeber von Wikipedia
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wikipedia DE

Telopea: Brief Summary ( Alemão )

fornecido por wikipedia DE

Telopea ist eine Pflanzengattung in der Familie der Silberbaumgewächse (Proteaceae) mit fünf Arten.

licença
cc-by-sa-3.0
direitos autorais
Autoren und Herausgeber von Wikipedia
original
visite a fonte
site do parceiro
wikipedia DE

Waratah ( Inglês )

fornecido por wikipedia EN

Waratah (Telopea) is an Australian-endemic genus of five species of large shrubs or small trees, native to the southeastern parts of Australia (New South Wales, Victoria, and Tasmania). The best-known species in this genus is Telopea speciosissima, which has bright red flowers and is the New South Wales (NSW) state emblem. The waratah is a member of the family Proteaceae, flowering plants distributed in the Southern Hemisphere. The key diagnostic feature of Proteaceae is the inflorescence, which is often very large, brightly coloured and showy, consisting of many small flowers densely packed into a compact head or spike. Species of waratah boast such inflorescences ranging from 6–15 cm in diameter with a basal ring of coloured bracts. The leaves are spirally arranged, 10–20 cm long and 2–3 cm broad with entire or serrated margins. The name waratah comes from the Eora Aboriginal people, the pre-European inhabitants of the Sydney area.

Taxonomy

The genus Telopea belongs to the plant family Proteaceae. Within the Proteaceae, their closest relatives appear to be the genera Alloxylon (tree waratahs), Oreocallis and Embothrium, a group of generally terminal red-flowering plants that skirt the southern edges of the Pacific Rim. Together they make up the subtribe Embothriinae.[1][2] The genus was first described by Robert Brown in 1810 from the type species Telopea speciosissima. There are five species of plant within the genus, all of which readily hybridize in cultivation.[3] There are two main branches, with one being the species pair of T. speciosissima and T. aspera, with the other lineage giving rise to T. truncata first, then T. oreades and T. mongaensis.[4] The speciosissima-aspera lineage (clade) has two synapomorphies—distinguishing common characteristics presumed not present in ancestors—leaves with toothed margins, and large red involucral bracts. The truncata-oreades-mongaensis lineage has flowers that open from the centre to the edge of the inflorescence (basitonic) rather than the reverse (acrotonic), which is a feature of the speciosissima-aspera clade and more distant relatives.[3]

Species

The genus Telopea contains five species:

Habitat

Species grow as either large shrubs or small trees with spirally arranged leaves with either entire or serrated margins. They prefer sandy loam soils and are a pyrogenic flowering species, meaning that they rely on post-fire flowering followed by production and dispersal of non-dormant seeds to take advantage of favourable growing conditions in the altered environment following a fire.

Distribution

The natural distributions of the five species of Telopea are all confined to east coast regions from northern New South Wales to Tasmania. Each of the species has its own distinctive natural distribution with minimal or no overlap. Climatic changes may have restricted the expansion of species distribution or led to its isolation in a particular region. However, waratahs can also grow outside of these natural distribution areas. Cultivation mostly occurs north of Sydney and in the Dandenong Ranges, Victoria. T. speciosissima has also been grown successfully in areas not on the map. In Australia these areas include south-west Western Australia, the Queensland coast and also Toowoomba. Overseas, New Zealand, the USA, South Africa, and Israel are all also able to grow waratah with varying degrees of success. It was introduced to England in 1789 but cannot survive English winters out of doors except in the south-west coastal regions, and it rarely flowers in glasshouses.

Cultivation

For some time the waratah has had a reputation as a difficult plant. It has a complex culture and for many years there have been cases of people trying to establish the plant only to have the attempt fail. This can be the effect of unsuitable soil conditions, aspect or climate. The waratah is also a slow plant to mature with a flowering period that is short, unpredictable and unreliable. Early issues with cultivation meant that approximately 90% of all waratahs sold at Sydney’s Flemington markets in the early 90s were bush picked. Some progress has been made in the 20 years since then with several cultivars being commercially grown mostly in areas to the North and South of Sydney and in the Dandenong Ranges in Victoria. Issues with cultivation are still present however.[5]

Propagation

Waratah seed pods, Blue Mountains, Sydney, 2022

The propagation of waratah for commercial production is relatively easy in comparison to other stages of the growth cycle. Plants are usually propagated from cuttings, fresh seed or stored seed. Fresh seed has a good germination rate but deteriorates fairly rapidly unless stored at low temperature and low humidity. Dry seed will last a few years in refrigerated storage but should be treated with a general purpose fungicide prior to storage and at propagation to ensure good germination rates and healthy seedlings. The best time to take cuttings is when the plant is experiencing a flush of growth. The cuttings are taken from firm wood from the last twelve months growth. If plant material is scarce, single nodes can be used for cutting propagation.

Other methods of propagation that are successful but not widely used for commercial production include grafting and tissue culture. Stock for grafting is vigorous and as such is labour-intensive in constant control of stock regrowth from the lignotuber. As such, it is not a recommended method but is useful for the fast growth of limited material. Rootstock and scion combinations are used for many woody perennials to provide the necessary mix of floral or fruit characteristics and cultural requirements.

Tissue culture is very labour-intensive and would likely only be used in the case of rapidly increasing the number of plants from limited or valuable material. The process is complex though as different clones require different optimal culture conditions requiring different developmental work for each clone. There also exists problems with hardening off, with roots and leaves produced in vivo dying off on planting out.

For Telopea plants propagated from seed, the transition from seedling to flower takes about 5 years. Cuttings may take only 2 years. The most common form of propagation is from seed, however, certain varieties and cultivars must be propagated from cuttings if the grower wishes the plants to remain true to form. Fresh seed has a higher viability than cuttings and will germinate 2–3 weeks after sewing.

Commercial cultivation

The overall cultivation of the waratah as a single process is a tricky one as flowering time, number and quality are easily affected by changes in the plant environment. These factors must be considered as early as prior to buying land for production. North facing aspect ensures a maximum of sun exposure. A combination of northerly, easterly and westerly aspects will spread the flowering time with the western slope flowering slightly later on. As the location approaches the equator, flowering time will be earlier. Elevation also has to be considered as it will affect temperature, a major influence on flowering time. More flowers will be produced in full sunlight although better quality flowers are found in the shade. Paul Nixon (1997) in his book ‘The Waratah’ claims that ‘the ideal situation is to have rich, well drained, deep soil with a north-easterly aspect giving the plants full sun until the flowers buds have initiated and then put shade cloth over the bushes until they have flowered’.

A primary consideration for cultivation is water drainage. The waratah naturally grows in poor, sandy soils where it thrives due to the soils excellent water draining properties. Drainage properties can be linked to aspect and as a result plants grown on a north easterly aspect will generally flower 1–2 weeks earlier than a westerly aspect at the same location. Watering systems must also be carefully considered to coincide with correct drainage. As an Australian native the waratah is a sturdy plant well adapted to coping with harsh environments and low rainfall. Watering systems are still necessary for cultivation in order to produce a reliable crop and a quality bloom.

In the wild the waratah has become adapted to growing in nutrient-poor soils leading many people to believe that this is what is necessary for the development of the plant. Fertilisers are not necessary for the development of the plant but the waratah has been shown to be receptive to some treatments. Many native plants have been known to have poor establishment in soils with high levels of nitrogen and phosphorus. This coincides with the observation that fertiliser application immediately after transplant of waratahs in the field often leads to high mortalities. Other studies have reported a strong growth response to high nutrient levels, particularly phosphorus. As yet, the details of this response are not yet clear and suggest a complex nitrogen-phosphorus relationship. Earlier work (1963) on related species, had indicated that fertiliser application may hasten maturation and give early flowering. If fertiliser is applied, drainage properties of the soil mean that nutrients are quickly leached and so the best application method is multiple applications at critical stages in development such as flush periods.

Pruning is a very important consideration for the commercial growth of waratahs in the effort to design a shape for the plant that will encourage the maximum production of saleable blooms. The aim is to get as many growing tips as possible as it is on these that the flowers will develop.

With the correct mix of factors for cultivation it is possible to produce up to sixty blooms per plant per year. This could translate to up to 20,000 to 50,000 blooms per ha. Waratah inflorescences are harvested when 0–50% of flowers are open, although inflorescences with 0–5% of flower open have the longest vase life and least opportunity for bract damage in the field.[5]

Cultivars

Cultivar 'Wirrimbirra White'
Cultivar 'Braidwood Brilliant'

A number of selected forms of T. speciosissima and hybrids with other Telopea species are being brought into cultivation. These exhibit variations in the colour of the flowers and/or the bracts. Some examples include:

  • 'Fire 'n' Ice' – red with white tips
  • 'Songlines' – pink in bud, opening flame red
  • 'Dreaming' – styles open white and mature to pink
  • 'Shade of Pale' – cream tinged with pink
  • 'Brimstone Blush' – red with a pink blush
  • 'Braidwood Brilliant' – red (T. speciosissima x T. mongaensis)
  • 'Wirrimbirra White' – almost pure white
  • 'Shady Lady' – blood red (T. speciosissima x T. oreades)

Issues of cultivation

Issues with cultivation occur throughout the growth cycle of Telopea spp. with the first issues being encountered at the seedling stage. Waratah seedlings are often associated with a common plant condition known as damping off. Damping off is a condition that causes the death of seeds or seedlings facilitated by a number of different fungal pathogens. A given seed can become infected with a fungus, often causing it to darken and soften, killing the seedling before it emerges or causing it to emerge in a pre-weakened state. Seedlings can also be infected after emergence resulting in the stem thinning until it eventually rots and the seedling topples over. The problem is often associated with and encouraged by excessively wet conditions.

Another major issue for production is bract browning – a discolouration of the showy floral bracts of the floral head occurring prior to harvest. Bract browning has been a major restraint to financial returns due to reduced cut flower quality. Browning is usually a result of sun damage but can also be associated with wind burn. The addition of shade cloths to crop management strategies has been shown to reduce levels of excessive light and has significantly minimised financial losses due to the reduction of occurrence of bract browning.

In the natural state, the waratah does not compete very well with surrounding shrubs and is at its best after fire when competition is removed. This means that weed presence during cultivation has a profound effect on growth, particularly when the waratahs are small. Weeds should be seriously tended to in the early stages of growth. Once plants have reached waist height cases should be analysed individually to determine the best method of weed control.

In New South Wales the most destructive pest to waratah crops is the macadamia twig girdler (Neodrepta luteotactella). The damage is caused by the larvae and damage generally first shows at a branch fork or leaf. The condition is generally confined to young shrubs or trees. The leaves are skeletonised and the larvae web them together into a shelter that incorporates larval faeces. Larvae can also burrow into the developing flower head, obliterating a crop entirely if left uncontrolled. Biological control methods include encouragement of bird species. The borer is present throughout the year and so pesticide treatments with a strong persistent spray should be carried out monthly. What is used for the borer should also control lesser pests such as white scale, which is common in the natural state, and Macadamia leafminer (Acocercops chionosema). Leafminer poses no threat to the life of the plant but it is preferred that stems for the market still have leaves attached. Thus damaged leaves lower the value of the crop. Chemicals with zylene or toluene should not be used as the will not be tolerated by the plant.

There are also issues of fungal infection. Fungal species cause stem rot (Phytophthora spp.) and root rot (Rhizoctonia spp.) that can ultimately cause plant death but can usually be controlled by adequate drainage.

A problem for production that emanates from within the plant is the high amount of genetic and therefore morphological variability present in the flowers produced. The need to lift the quality and consistency of cut flower product has been repeatedly highlighted by industry reviews. Inconsistency of product is a key impediment to further industry growth.

Floral morphology

Telopea speciosissima floral morphology

In many genera of Proteaceae the inflorescence is very large and showy, often in bright colours, consisting of many small flowers densely packed into a compact head or spike. The individual flowers within the inflorescence also give Proteaceae species a unique look. Telopea species are long-lived, perennial plants that re-sprout from lignotubers after fire. After a few years of fire, re-sprouting stems produce the terminal flowers which continue the flowering cycle annually. The Telopea 'flower' is in fact an inflorescence that comprises from as few as 10 to as many as 240 individual flowers, depending upon the species concerned. The style is thickened at the distal end to form a 'pollen-presenter'. Pollen presenters have an area on the style end that presents the pollen to the pollinator. The stigma is initially trapped within the perianth and as the style grows it becomes bent until it splits the perianth and the pistil is released to spring upright. An open inflorescence usually contains functionally male and female flowers at any one time. Inflorescences range from 6–15 cm in diameter with a basal ring of coloured bracts.

Flowering processes

Before a flower can be produced the plant must undergo floral induction and initiation. Floral induction involves physiological processes in the plant that result in the shoot apical meristem becoming competent to develop flowers. It involves biochemical changes at the apex, particularly those caused by cytokinins and the processes can be reversed. Floral initiation is the morphological transformation of an induced growing point from a vegetative to a floral primordium and involves the plant hormone florigen. Florigen is produced in the leaves in reproductively favourable conditions and acts in buds and growing tips to induce a number of different physiological and morphological changes. Once this process begins, in most plants, it cannot be reversed and the stems develop flowers, even if the initial start of the flower formation event was dependent of some environmental cue. Once the process begins, even if that cue is removed the stem will continue to develop a flower. Flower induction and initiation can simply occur when a plant has reached a mature enough age. However, in many plant species floral process occur in response to a number of environmental signals, or alternatively, are repressed by environmental signals.

Floral initiation in T. speciosissima has been observed from mid-December, with floral buds developing more rapidly on older shoots and floral primordia emerging from mid-January to February. The floral primordia initiate over a 6–8 week period after primary flush growth from November to January. After the primordia initiation there may be another vegetative flush of growth on the plants. The flower develops in bud form for seven to eight months. The date of flowering is highly variable as waratah flowering is sensitive to its environment.

Royal Doulton "Waratah" pattern, after 1922, a local flower to appeal to the Australian market.

Flowering tends to vary with geographic location and climatic differences, occurring from early August in coastal Queensland and up to December in Tasmania. Waratahs have a short flowering period in one location, although varieties can be selected to ensure a reasonable spread. It has been shown that in a population of 1000 seedlings where the total flowering time was five weeks, the spread was so that 10% flowered in week one and 10% in week five. Much variation was accounted for by varietal differences with plants of the one variety flowering at much the same time. Generally, waratahs flower over a 4–6 week period in spring (September–October) in the Sydney region, but later in cooler areas.

Cultural references

Indigenous Tharawal peoples from around the Cronulla region of southern Sydney use the waratah medicinally. Placing the flowers into a bowl of water, so that the nectar be soaked out, the flower water is then drunk for pleasure, for its strengthening effect and for curing illnesses in children and elderly.

The botanical journal Telopea is named after the genus, as is the western Sydney suburb of Telopea, New South Wales.

Neptune Oil Company used the waratah as a brand for its Motor spirit from the late 1910's until being phased out in the 40's

Telopea speciosissima the floral emblem of the state of New South Wales and several organisations in the state, including the New South Wales Waratahs rugby team and Grace Bros. (now Myer). Waratah is also the name of the Sydney Trains A set, a class of electric multiple unit trains operated by Sydney Trains in Sydney, New South Wales, Australia. In 2009, the Premier of New South Wales, Nathan Rees, commissioned a state logo based on the floral emblem. The resultant logo design has been criticised as resembling a lotus rather than the New South Wales waratah.[6]

Further reading

  • Atwell BJ, Kriedemann PE, Turnbull C (1999) The processes of floral induction and initiation. In ‘Plants in Action: Adaptation in Nature, Performance in Cultivation’. pp. 8.3.2 – 8.4. (Macmillan Education Australia Pty Ltd.: Melbourne, Australia).
  • Ausin I, Alonso-Blanco C, Martinez-Zapater JM (2005) Environmental regulation of flowering. Int. J. Dev. Biol. 49:689–705.
  • Boden A (1985) Waratah – Telopea speciosissima – Floral Emblem of New South Wales, Australian National Botanic Gardens. Available from URL: http://www.anbg.gov.au/emblems/nsw.emblem.html [accessed 20 August 2012].
  • Burnett J (1993) Will the waratah ever fulfil its potential?. In ‘SGAP 17th Biennial Seminar, Sydney, NSW, 27 Sep – 1 Oct’. Available at URL: http://anpsa.org.au/APOL14/jun99-5.html [accessed 20 August 2012].
  • Collins BG, Walsh M, Grey J (2008). "Floral development and breeding systems of Dryandra sessilis and Grevillea wilsonii (Proteaceae)". Australian Journal of Botany. 56 (2): 119–130. doi:10.1071/bt07147.
  • Crisp MD, Weston PH (1993). "Geographic and Ontogenic Variation in Morphology of Australian Waratahs (Telopea, Proteaceae)". Systematic Biology. 42: 49–76. doi:10.1093/sysbio/42.1.49.
  • Crisp, MD & Weston, PH (1995). "Telopea". In McCarthy, Patrick (ed.). Flora of Australia: Volume 16: Eleagnaceae, Proteaceae 1. CSIRO Publishing / Australian Biological Resources Study. pp. 382–390. ISBN 0-643-05693-9.
  • Cruden RW (2009). "Pollen grain size, stigma depth, and style length: the relationships revisited". Plant Systematics and Evolution. 278 (3–4): 223–238. doi:10.1007/s00606-008-0142-8. S2CID 22688500.
  • Denham AJ (2008). "Seed predation limits post-fire recruitment in the waratah (Telopea speciosissima)". Plant Ecology. 199: 9–19. doi:10.1007/s11258-008-9407-0. S2CID 9479180.
  • Denham AJ, Auld TD (2002). "Flowering, seed dispersal, seed predation and seedling recruitment in two pyrogenic flowering resprouters". Australian Journal of Botany. 50 (5): 545–557. doi:10.1071/bt02009.
  • Dupee SA, Goodwin PB (1990a). "Effect of Temperature, Daylength and Growth-regulators on Flowering in Protea, Telopea and Leucospermum". Acta Horticulturae. 264 (264): 79–86. doi:10.17660/actahortic.1990.264.9.
  • Dupee SA, Goodwin PB (1990b). "Flower Initiation in Protea and Telopea". Acta Horticulturae. 264 (264): 71–77. doi:10.17660/ActaHortic.1990.264.8.
  • Faragher JD (1986). "Effects of Cold-storage Methods on Vase life and Physiology of Cut Waratah Inflorescences (Telopea speciosissima, Proteaceae)". Scientia Horticulturae. 29 (1–2): 163–171. doi:10.1016/0304-4238(86)90043-9.
  • Gerber AI, Theron KI, Jacobs G (2001b). "Manipulation of flowering time by pruning of Protea cv. Sylvia (P-eximia x P-susannae)". HortScience. 36: 909–912. doi:10.21273/HORTSCI.36.5.909.
  • Goodwin PB (1983). "Australian Natives – Fertilizing Container-Grown Plants". Australian Horticulture. 81: 57–65.
  • Jarillo JA, del Olmo I, Gomez-Zambrano A, Lazaro A, Lopez-Gonzalez L, Miguel E, Narro-Diego L, Saez D, Pineiro M (2008). "Photoperiodic control of flowering time". Spanish Journal of Agricultural Research. 6: 221–244. doi:10.5424/sjar/200806s1-391.
  • Jarillo JA, Pineiro M (2011). "Timing is everything in plant development. The central role of floral repressors". Plant Science. 181 (4): 364–378. doi:10.1016/j.plantsci.2011.06.011. PMID 21889042.
  • Lyne A (2011) Growing Waratahs. Available at URL: http://www.anbg.gov.au/telopea/index.html [accessed 20 August 2012]
  • Martyn AJ, Larkum AW, McConchie R, Offord CA (2008). "Photoinhibition and changes in pigments associated with bract browning in waratahs (Telopea spp., Proteaceae)". Journal of Horticultural Science & Biotechnology. 83 (3): 367–373. doi:10.1080/14620316.2008.11512393. S2CID 88004468.
  • Martyn A, McConchie R, Offord C (2006) Effect of Shade on Bract Browning of Waratahs(Telopea spp.). ‘Proceedings of the VIIth International Protea Research Symposium, International Society Horticultural Science’. (Ed A Gerber) pp. 117–126.
  • McConchie R, Delaporte K, Ekman J, Gollnow B, Lidbetter J, Srhoj J, Seaton K, Worral R (2008) Standing out from the bunch: an Australian update on improving quality. In ‘IX International Protea Research Symposium, Stellenbosch, South Africa, 3–6 Sep 2008’. pp. 191–196 (International Society for Horticultural Science (ISHS)).
  • Mercure PS (1998) Damping off, University of Connecticut. Available at URL:http://www.hort.uconn.edu/ipm/greenhs/htms/dampofgh.htm [accessed 20 August 2012].
  • Nichols DG, Beardsell DV (1981). "The response of phosphorus sensitive plants to slow-release fertilisers in soil-less potting mixtures". Scientia Horticulturae. 15 (4): 301–309. doi:10.1016/0304-4238(81)90084-4.
  • Nixon P (1997) The Waratah. 2nd ed. (Kangaroo Press, East Roseville, Sydney, Australia). NSW Government, "Symbols & Emblems of New South Wales". URL: http://www.nsw.gov.au/symbols-emblems-nsw [accessed 19 July 2011].
  • Offord CA (1996) Waratahs. In ‘Horticulture of Australian Plants’. (UNSW Press, Sydney, Australia).
  • Offord CA, Campbell LC (1994). "The Waratah: Taming of an Emblem". Australian Horticulture: 52–57.
  • Offord CA, Goodwin PB, Nixon P (1990). "Clonal selection and micropropagation of waratah". Second International Protea Research Symposium. 264: 49–52.
  • QLD Gov. (2003) Macadamia Problem Solver and Bug Identifier, Queensland Government. Available at URL: http://era.deedi.qld.gov.au/1964/10/mac-problemsolver_Part2.pdf [accessed 20 August 2012].
  • Rossetto M, Thurlby KAG, Offord CA, Allen CB, Weston PH (2011) The impact of distance and a shifting temperature gradient on genetic connectivity across a heterogeneous landscape. Bmc Evolutionary Biology 11.
  • Specht RL (1963). "Dark island heath (Ninety-Mile-Plain, South Australia). VII. The effect of fertilizers on composition and growth, 1950–60". Australian Journal of Botany. 11: 67–94. doi:10.1071/bt9630067.
  • Specht RL (1978). "Conditions for the cultivation of Australian plants". Australian Plants. 9: 312–313.
  • Tooke F, Ordidge M, Chiurugwi T, Battey N (2005). "Mechanisms and function of flower and inflorescence reversion". Journal of Experimental Botany. 56 (420): 2587–2599. doi:10.1093/jxb/eri254. PMID 16131510.
  • Whelan RJ, Goldingay RL (1989). "Factors affecting fruit-set in Telopea speciosissima (Proteaceae) – The importance of pollen limitation". Journal of Ecology. 77 (4): 1123–1134. doi:10.2307/2260827. JSTOR 2260827.
  • Worrall RJ (1985) Thesis submitted for the degree of Master of Science in Agriculture. Unpublished, University of Sydney.
  • Worall RJ (1993) Market-focussed plan for the development of the waratah cut flower industry in NSW. (NSW Agriculture).

References

  1. ^ Johnson, L. A. S.; Briggs, Barbara G. (1975). "On the Proteaceae: the evolution and classification of a southern family". Botanical Journal of the Linnean Society. 70 (2): 83–182. doi:10.1111/j.1095-8339.1975.tb01644.x.
  2. ^ Weston, Peter H.; Barker, Nigel P. (2006). "A new suprageneric classification of the Proteaceae, with an annotated checklist of genera". Telopea. 11 (3): 314–44. doi:10.7751/telopea20065733.
  3. ^ a b Rossetto, Maurizio; Allen, Chris B.; Thurlby, Katie A.G.; Weston, Peter H.; Milner, Melita L. (2012). "Genetic structure and bio-climatic modeling support allopatric over parapatric speciation along a latitudinal gradient". BMC Evolutionary Biology. 12. 149. doi:10.1186/1471-2148-12-149. PMC 3495659. PMID 22906180.
  4. ^ Weston, Peter H.; Crisp, Michael D. (1994). "Cladistic biogeography of waratahs (Proteaceae, Embothrieae) and their allies across the pacific". Australian Systematic Botany. 7 (3): 225–49. doi:10.1071/SB9940225.
  5. ^ a b Joyce, Burnett. "Will the waratah ever fulfil its potential?". Australian native plant society (Australia). Retrieved 7 March 2015.
  6. ^ Benson, Simon (16 July 2009). "NSW Premier accidentally replaces waratah with lotus". Herald-Sun. News Ltd. Retrieved 27 February 2015.

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Waratah: Brief Summary ( Inglês )

fornecido por wikipedia EN

Waratah (Telopea) is an Australian-endemic genus of five species of large shrubs or small trees, native to the southeastern parts of Australia (New South Wales, Victoria, and Tasmania). The best-known species in this genus is Telopea speciosissima, which has bright red flowers and is the New South Wales (NSW) state emblem. The waratah is a member of the family Proteaceae, flowering plants distributed in the Southern Hemisphere. The key diagnostic feature of Proteaceae is the inflorescence, which is often very large, brightly coloured and showy, consisting of many small flowers densely packed into a compact head or spike. Species of waratah boast such inflorescences ranging from 6–15 cm in diameter with a basal ring of coloured bracts. The leaves are spirally arranged, 10–20 cm long and 2–3 cm broad with entire or serrated margins. The name waratah comes from the Eora Aboriginal people, the pre-European inhabitants of the Sydney area.

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Telopeo ( Esperanto )

fornecido por wikipedia EO
 src=
Telopea speciosissima, simbolo de Nov-Sud-Kimrio

Telopeo (Telopea, angle Waratah) estas aŭstralia genro de kvin specioj de arbustoj kaj arbetoj, apartenanta al la familio de Proteacoj. Ĝi indiĝenas en la aŭstraliaj subŝtatoj Nov-Sud-Kimrio, Viktorio, kaj Tasmanio. La plej konata specio estas Telopea speciosissima, kies grandaj ruĝaj floroj estas ŝtata simbolo de Nov-Sud-Kimrio.

Specioj

  • Telopea aspera - nordorienta parto de Nov-Sud-Kimrio
  • Telopea mongaensis - sudorienta parto de Nov-Sud-Kimrio
  • Telopea oreades - orienta parto de Viktorio ĝis ekstreme sudorienta Nov-Sud-Kimrio
  • Telopea speciosissima - orienta parto de Nov-Sud-Kimrio
  • Telopea truncata - Tasmanio

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Telopeo: Brief Summary ( Esperanto )

fornecido por wikipedia EO
 src= Telopea speciosissima, simbolo de Nov-Sud-Kimrio

Telopeo (Telopea, angle Waratah) estas aŭstralia genro de kvin specioj de arbustoj kaj arbetoj, apartenanta al la familio de Proteacoj. Ĝi indiĝenas en la aŭstraliaj subŝtatoj Nov-Sud-Kimrio, Viktorio, kaj Tasmanio. La plej konata specio estas Telopea speciosissima, kies grandaj ruĝaj floroj estas ŝtata simbolo de Nov-Sud-Kimrio.

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Telopea ( Espanhol; Castelhano )

fornecido por wikipedia ES

Waratah (Telopea) es un género de cinco especies de grandes arbustos o pequeños árboles de las proteaceae, nativas del sureste de Australia, de Nueva Gales del Sur, Victoria y Tasmania.

Descripción

Tienen las hojas dispuestas en espiral 10-20 cm de largo y 2-3 cm de ancho con márgenes enteros o serrados y grandes y densos capítulos de 6-15 cm diámetro con numerosas pequeñas flores rojas y un anillo basal de brácteas rojas. El nombre waratah viene de los pueblos aborígenes de los habitantes alrededor del área de Sídney.

Taxonomía

Dentro de las proteaceae, sus más parientes más cercanos parecen ser Alloxylon (Árboles Waratahs), Oreocallis y Embothrium, un grupo de plantas con capullos de flores rojas que bordean los confines del sur del anillo del Pacífico. Todos ellos juntos componen la subtribu Embothriinae dentro de la familia.

Especies

Existen cinco especies que ocupan distintos rangos con mínimos o ningún traslape; listadas de norte a sur:

El waratah de Nueva Gales del Sur es nativo de áreas de la cuenca geológica de Sídney, distritos central y sur de la costa, y de las Montañas Azules; crece aproximadamente 4 m de alto. Se desarrolla típicamente en suelos con margas arenosas en cadenas montañosas y mesetas. Este waratah es endémico de Nueva Gales del Sur, pero se ha extendido gracias a su popularidad, a Victoria y aún Tasmania.

Cultivo

 src=
Un cultivar blanco de Waratah.
 src=
Waratah de Nueva Gales del Sur.

Los Waratahs son populares, aunque algo difícil de cultivar. Son plantas ornamentales en los jardín en Australia; algunos híbridos y cultivares se han desarrollado, incluyendo algunos con flores cremosa-blancas y rosas así como también el rojo natural. Las formas blancas de Telopea speciossisima son nombradas Telopea "Blanco Wirrimbirra" y T. "dama blanca de sombra", mientras T. "dama blanca de sombra" y T. "dama roja de sombra" son en realidad híbridos entre Telopea speciosissima y Telopea oreades.

Cultura popular

The revista científica botánica Telopea debe su nombre debido al género, así como el suburbio occidental de Telopea en Sídney.

Telopea speciosissima es el emblema floral de Nueva Gales del Sur y varios organizaciones en el estado, incluyendo, el equipo de rugby de los Waratahs y Grace Bros. (ahora Myer).

Referencias

  • Crisp, MD & Weston,PH (1995). «Telopea». En McCarthy, Patrick (ed.), ed. Flora of Australia: Volume 16: Eleagnaceae, Proteaceae 1. CSIRO Publishing / Australian Biological Resources Study. pp. 382-390. ISBN 0-643-05693-9.

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Telopea: Brief Summary ( Espanhol; Castelhano )

fornecido por wikipedia ES

Waratah (Telopea) es un género de cinco especies de grandes arbustos o pequeños árboles de las proteaceae, nativas del sureste de Australia, de Nueva Gales del Sur, Victoria y Tasmania.

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Telopea ( Francês )

fornecido por wikipedia FR
 src=
Cultivar de Telopea blanc
 src=
Cultivar de telopea; photo: Suellen Harris

Telopea est un genre de cinq espèces de grands buissons ou petits arbres de la famille des Proteaceae, originaires du sud-est de l'Australie, en Nouvelle-Galles du Sud, au Victoria et en Tasmanie. Ils ont des feuilles verticillées de 10 à 20 centimètres de long et de 2 à 3 de large, des grappes de nombreuses petites fleurs rouges formant des boules de 6 à 15 cm de diamètre avec une base de bractées rouges.

Classification

Dans les Proteaceae, les genres les plus proches sont les Alloxylon, les Oreocallis et les Embothrium, un groupe de plantes à fleurs rouges terminales qui poussent dans les pays de l'hémisphère sud bordant l'Océan Pacifique. Ces quatre genres sont regroupés dans la sous famille des Embothriinae.

Espèces

Les cinq espèces de telopea occupent toutes un territoire différent avec parfois un très faible chevauchement.

Telopea speciosissima est originaire de la région de Sydney et de ses alentours y compris les Montagnes Bleues. Ce sont des arbustes pouvant atteindre quatre mètres de haut qui poussent dans les sols limoneux des montagnes et des plateaux. Endémique en Nouvelle Galles du Sud, on la trouve maintenant au Victoria et en Tasmanie à cause de sa popularité.

Culture

Les telopeas sont des plantes populaires, quoique parfois difficiles à faire pousser, dans les jardins australiens. On a développé plusieurs hybrides et cultivars avec des fleurs blanc-crème, rose ou, comme dans la nature, rouge. Les deux principales variétés blanches s'appellent Telopea "Wirrimbirra White" et T. "Shady Lady White". T. "Shady Lady Pink" et T. "Shady Lady Red" sont des hybrides de Telopea speciosissima et Telopea oreades.

L'école Telopea Park School est située à Canberra en Australie et son emblème est la fleur Telopea Speciossissima.

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Telopea: Brief Summary ( Francês )

fornecido por wikipedia FR
 src= Cultivar de Telopea blanc  src= Cultivar de telopea; photo: Suellen Harris

Telopea est un genre de cinq espèces de grands buissons ou petits arbres de la famille des Proteaceae, originaires du sud-est de l'Australie, en Nouvelle-Galles du Sud, au Victoria et en Tasmanie. Ils ont des feuilles verticillées de 10 à 20 centimètres de long et de 2 à 3 de large, des grappes de nombreuses petites fleurs rouges formant des boules de 6 à 15 cm de diamètre avec une base de bractées rouges.

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wikipedia FR

Telopėja ( Lituano )

fornecido por wikipedia LT

Telopėja (lot. Telopea) – protėjinių (Proteaceae) augalų šeimai priklausanti visžalių augalų gentis. Paplitusi pietryčių Australijoje (Naujasis Pietų Velsas, Tasmanija, Viktorija).

Lapai išsidėstę spirališkai, 10-20 cm ilgio, 2-3 cm pločio. Žiedynas 6-15 cm skersmens.

Rūšys

Gentį sudaro 5 stambių krūmų ir nedidelių medžių rūšys.


Vikiteka

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Telopėja: Brief Summary ( Lituano )

fornecido por wikipedia LT

Telopėja (lot. Telopea) – protėjinių (Proteaceae) augalų šeimai priklausanti visžalių augalų gentis. Paplitusi pietryčių Australijoje (Naujasis Pietų Velsas, Tasmanija, Viktorija).

Lapai išsidėstę spirališkai, 10-20 cm ilgio, 2-3 cm pločio. Žiedynas 6-15 cm skersmens.

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Telopea ( Português )

fornecido por wikipedia PT

Telopea é um género botânico pertencente à família Proteaceae[1]. É endémico da Austrália.

Referências

  1. «Telopea — World Flora Online». www.worldfloraonline.org. Consultado em 19 de agosto de 2020
 title=
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Telopea: Brief Summary ( Português )

fornecido por wikipedia PT

Telopea é um género botânico pertencente à família Proteaceae. É endémico da Austrália.

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Телопея ( Russo )

fornecido por wikipedia русскую Википедию
Царство: Растения
Подцарство: Зелёные растения
Отдел: Цветковые
Надпорядок: Proteanae Takht., 1967
Порядок: Протеецветные
Семейство: Протейные
Род: Телопея
Международное научное название

Telopea (Sm.) R.Br.

Виды Wikispecies-logo.svg
Систематика
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NCBI 54954GRIN g:11927IPNI ???FW 317967

Телопея (лат. Telopea) — род растений семейства Протейные.

Распространен в юго-восточной части Австралии (Новый Южный Уэльс и Виктория) и Тасмании.

Описание

Листья — плотные, кожистые.

Цветы — собраны в крупные соцветия 6—15 см в диаметре, естественная окраска цветов — красная.

Виды

Род состоит из 5 видов крупных кустарников или небольших деревьев:

Применения

 src=
Telopea speciosissima, белая форма

Является популярным декоративным растением в садах Австралии. Было выведено несколько новых гибридов и сортов, с цветовой окраской белого и розового цвета, в отличие от естественного красного.

Культурное значение

Telopea speciosissima — цветочная эмблема штата Новый Южный Уэльс, а также нескольких организаций.

Примечания

  1. Об условности указания класса двудольных в качестве вышестоящего таксона для описываемой в данной статье группы растений см. раздел «Системы APG» статьи «Двудольные».


Дубовый лист Это заготовка статьи по ботанике. Вы можете помочь проекту, дополнив её.  title=
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Телопея: Brief Summary ( Russo )

fornecido por wikipedia русскую Википедию

Телопея (лат. Telopea) — род растений семейства Протейные.

Распространен в юго-восточной части Австралии (Новый Южный Уэльс и Виктория) и Тасмании.

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