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Genus Overview ( Inglês )

fornecido por EOL authors

There are a number of characteristics which can be summarized regarding the Dudleya genus. Genus occurrences are restricted to southwest North America; in fact, according to Low, approximately 98 percent of genus occurrences are along or near the coasts of Baja California, (the states of Baja Sur and Baja Norte in Mexico), plus the northward extension into the USA in the states of California and southern Oregon. A small number of genus populations are found in Arizona and possibly Nevada. The center of genus diversity is the coastal region where Mexico and the USA meet. Dudleyas have a strong affinity for the coastal regions, but some species have ranges that extend into the nearby mountains and deserts.

Most species of Dudleya grow at elevations near sea level, but a few grow on or even atop mountains of Baja California, the high Sierra Nevada and in many other California mountain ranges. Many, such as D.cymosa, prefer canyon locations on vertical or near vertical rocky walls, often in the shade, surviving in very little soil in the crevices of almost solid rock. Sometimes the occurrences are in locations that receive no direct sunlight. Almost all Dudleya species have succulent leaves arranged into rosettes. Because of this rosette geometry many genus members are striking in their visual appeal. Many, including the Canyon Liveforever, have leaves that are glaucous (covered with a whitish or bluish waxy coating) or frosty looking.

The chromosome count of all Dudleya species is n=17, with about 35% of all populations consisting of polyploids (individuals with multiples of the base number). Remarkably, according to extensive crossing experiments conducted by Verity, all Dudleya taxa are capable of hybridizing in nature.

There are eight recognized subspecies of D. cymosa (flower color in parentheses):

  • D. cymosa subsp. agourensis K.M.Nakai: Canyon Liveforever (bright yellow)
  • D. cymosa subsp. crebrifolia K.M.Nakai & Verity: San Gabriel River dudleya (mustard yellow)
  • D. cymosa subsp. costafolia Bartel & Shevock: Pierpoint Springs Dudleya (bright yellow)
  • D. cymosa subsp. cymosa: Canyon Liveforever, the nominate subspecies (bright yellow, red or orange)
  • D. cymosa subspmarcescens Moran: Marescent Dudleya (bright yellow with orange or red markings)
  • D. cymosa subspovatifolia: (Britton) Moran Santa Monica Mountains Dudleya (bright yellow, rarely with red or orange marking)
  • D. cymosa subsp. paniculata: (Jeps.) K.M.Nakai (pale yellow-white to pale yellow-pink)
  • D. cymosa subsp. pumila: (Rose) K.M.Nakai (bright yellow or red)
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Dudleya ( Azerbaijano )

fornecido por wikipedia AZ

Dudleya (lat. Dudleya) — daşdələnçiçəklilər sırasının dovşankələmikimilər fəsiləsinə aid bitki cinsi.

Təbii yayılması

Botaniki təsviri

Ekologiyası

Azərbaycanda yayılması

İstifadəsi

Ədəbiyyat

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Dudleya: Brief Summary ( Azerbaijano )

fornecido por wikipedia AZ

Dudleya (lat. Dudleya) — daşdələnçiçəklilər sırasının dovşankələmikimilər fəsiləsinə aid bitki cinsi.

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Stenurt (Dudleya) ( Dinamarquês )

fornecido por wikipedia DA

Stenurt (Dudleya) er en slægt med ca. 45 arter, som er udbredt i det sydvestlige Nordamerika. Det er sukkulente stauder med en grundstillet roset af tykke, grå eller grågrønne blade. Blomsterne bæres på høje stængler, som ender i en stor stand. Der er 5 af både bægerblade og kronblade, og de er sammenvoksede forneden. Der er fem støvfang og 10 støvdragere.

Arter
  • Dudleya abramsii
  • Dudleya anthonyi – tidligere i Echeveria
  • Dudleya attenuata – tidligere i Echeveria
  • Dudleya blochmaniae
  • Dudleya caespitosa – tidligere Sedum cotyledon
  • Dudleya calcicola
  • Dudleya candelabrum
  • Dudleya candida – tidligere i Echeveria
  • Dudleya cedrosensis
  • Dudleya cultrata – tidligere i Echeveria
  • Dudleya cymosa – tidligere i Echeveria
  • Dudleya densiflora
  • Dudleya edulis – tidligere i Echeveria eller Sedum
  • Dudleya farinosa
  • Dudleya gnoma
  • Dudleya greenei
  • Dudleya guadalupensis
  • Dudleya hassei
  • Dudleya lanceolata
  • Dudleya linearis
  • Dudleya multicaulis
  • Dudleya nesiotica
  • Dudleya pachyphytum
  • Dudleya palmeri
  • Dudleya pulverulenta – tidligere Echeveria argentea eller E. pulverulenta
  • Dudleya saxosa – tidligere Echeveria collomiae
  • Dudleya setchellii
  • Dudleya stolonifera
  • Dudleya traskiae
  • Dudleya variegata
  • Dudleya verityi
  • Dudleya virens
  • Dudleya viscida
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Dudleya ( Alemão )

fornecido por wikipedia DE

Dudleya ist eine Pflanzengattung aus der Familie der Dickblattgewächse (Crassulaceae). Der botanische Name der Gattung ehrt den US-amerikanischen Botaniker William Russel Dudley (1849–1911).[1]

Beschreibung

Vegetative Merkmale

Die Arten der Gattung Dudleya sind ausdauernde, kahle Rosettenpflanzen mit meist faserigen Wurzeln und sukkulenten Blättern. Ihre Triebe sind nicht verzweigt oder verzweigt, dann meist dichotom. Sie sind an ihrer Basis meist mit mehr oder weniger vertrockneten Blättern oder Blattbasen bedeckt. Die Triebe sind meist kurz und etwas aufrecht und erreichen Durchmesser zwischen 10 und 40 Millimeter (selten 1,5 bis 90 Millimeter). Die Rosetten stehen endständig an den Trieben. Sie besitzen Durchmesser von 3 bis 25 Zentimetern (selten 1 bis 50 Zentimeter) und bestehen aus 20 bis 40, selten 3 bis 120 Blättern. Die Blätter sind in ihrer Form sehr variabel. Meist sind sie mehr oder weniger länglich bis länglich (verkehrt-)lanzettlich. Die Blattoberseite ist meistens mehr oder weniger flach oder leicht konkav bis rinnig. Ihre Unterseite ist meist etwas gerundet bis konkav. Sie sind mit einer breite Basis vollständig mit der Sprossachse verbunden und an der Basis oft verbreitert. Die Blattspitze ist stumpf-abgerundet bis scharf-spitz. Die glauken bis dicht bemehlten Blätter sind meist immergrün. Sie werden 1 bis 10 Zentimeter (selten 0,6 bis 40 Zentimeter) lang, 0,2 bis 3 Zentimeter (selten bis zu 10 Zentimeter) breit und sind 2 bis 6 Millimeter (selten 1 bis maximal 25 Millimeter) dick.

Blütenstände und Blüten

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Blüten von Dudleya cymosa
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Blüten von Dudleya nesiotica

Die einzelnen bis zahlreichen, meist mehr oder weniger aufrechten bis aufsteigenden Blütentriebe erscheinen jährlich aus den Blattachseln. Sie sind 5 bis 40 Zentimeter (bis maximal 100 Zentimeter) lang. Der zymöse Blütenstand besteht aus 3 bis 20 (selten 2 bis 30) Blüten. Die fünfzähligen (selten vierzähligen) duftlosen Blüten sind obdiplostemon. Der Blütenkelch ist 3 bis 8 Millimeter (selten 2 bis 9 Millimeter) lang. Seine Kelchblätter sind an der Basis miteinander verwachsen. Ihre freien Kelchzipfel sind 1,5 bis 6 Millimeter (selten 1 bis 8 Millimeter) lang. Die Blütenkrone hat Durchmesser von 3,5 bis 23 Millimeter und eine Länge von 1 bis 4 Millimetern (selten 0,5 bis 10 Millimetern). Ihre meist blass gelblichen bis gelben Kronblätter sind meist etwas elliptisch bis länglich oder eiförmig. Die Kronzipfel sind mehr oder weniger aufrecht oder von der Mitte bis zur Spitze etwas spreizend-ausgebreitet bis zurückgebogen.

Die zehn Staubblätter stehen in zwei Kreisen. Sie sind kürzer als Kronblätter. Die Staubbeutel sind meist gelb und 1 bis 2 Millimeter lang. Die gestutzten Nektarschüppchen sind 0,5 bis 2 Millimeter breit. Der Griffel ist schlank.

Früchte und Samen

Die Frucht ist eine mehr oder weniger braune oder bräunliche und nur selten rote Balgfrucht. Die darin enthaltenen mehr oder weniger braunen, schmal spindelförmigen Samen sind 0,5 bis 1,5 Millimeter groß. Ihre Samenschale ist gerippt.

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Untergattung Dudleya: Dudleya abramsii subsp. parva
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Untergattung Dudleya: Dudleya cespitosa
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Untergattung Dudleya: Dudleya candelabrum
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Untergattung Dudleya: Dudleya cymosa
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Untergattung Dudleya: Dudleya farinosa
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Untergattung Dudleya: Dudleya lanceolata
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Untergattung Dudleya: Dudleya pulverulenta
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Untergattung Dudleya: Dudleya saxosa
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Untergattung Dudleya: Dudleya verityi
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Untergattung Stylophyllum: Dudleya edulis
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Untergattung Stylophyllum: Dudleya virens subsp. hassei
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Untergattung Hasseanthus: Dudleya nesiotica

Systematik und Verbreitung

Die Gattung Dudleya ist im Südwesten Oregons, im Süden Nevadas, in Zentral- und West-Arizona und in Kalifornien in den Vereinigten Staaten sowie in den mexikanischen Bundesstaaten Sonora und Baja California verbreitet, wo die Pflanzen in meist küstennahen Gebieten vorwiegend an felsigen Stellen wachsen.

Die Erstbeschreibung durch Nathaniel Lord Britton und Joseph Nelson Rose wurde 1903 veröffentlicht.[2] Nach Joachim Thiede wird die Gattung Dudleya in drei Untergattungen mit folgenden Arten untergliedert:[3]

  • Untergattung Dudleya
    • Dudleya abramsii Rose: Mit den Unterarten:
      • Dudleya abramsii subsp. abramsii: Sie kommt in Kalifornien und Baja California in Höhenlagen zwischen 300 und 1900 Metern Meereshöhe vor.[4]
      • Dudleya abramsii subsp. affinis K.M. Nakai: Sie kommt in Kalifornien in Höhenlagen zwischen 1800 und 2600 Metern Meereshöhe vor.[4]
      • Dudleya abramsii subsp. bettinae (Hoover) Bartel: Sie kommt in Kalifornien in Höhenlagen zwischen 0 und 200 Metern Meereshöhe vor.[4]
      • Dudleya abramsii subsp. murina (Eastw.) Moran: Sie kommt in Kalifornien in Höhenlagen zwischen 100 und 600 Metern Meereshöhe vor.[4]
      • Dudleya abramsii subsp. parva (Rose & Davidson) Bartel: Sie wird von manchen Autoren auch als eigenständige Art angesehen: Dudleya parva Rose & Davidson. Sie kommt nur im Ventura County in Kalifornien in Höhenlagen zwischen 100 und 400 Metern Meereshöhe vor.[4]
    • Dudleya acuminata Rose (Syn.: Dudleya brevipes Rose): Sie kommt im nordwestlichen Mexiko vor.[5]
    • Dudleya albiflora Rose: Sie kommt in Niederkalifornien vor.[5]
    • Dudleya anthonyi Rose: Sie kommt in Niederkalifornien vor.[5]
    • Dudleya arizonica Rose: Sie kommt vom südöstlichen Kalifornien bis ins westliche Arizona, im nördlichen Baja California und im nordwestlichen mexikanischen Bundesstaat Sonora vor.[5]
    • Dudleya brittonii Johans.: Sie kommt im nördlichen Baja California vor.[5]
    • Dudleya cespitosa (Haw.) Britton & Rose; Heimat: Kalifornien; sie kommt in Höhenlagen zwischen 0 und 600 Metern Meereshöhe vor.[4]
    • Dudleya calcicola Bartel & Shevock. Sie wird auch von manchen Autoren als Unterart angesehen: Dudleya abramsii Rose subsp. calcicola (Bartel & Shevock) K. M. Nakai. Sie kommt in Kalifornien in Höhenlagen zwischen 500 und 2000 Metern Meereshöhe vor.[4]
    • Dudleya candelabrum Rose: Sie ist ein Endemit von Santa Cruz Island und Santa Rosa Island (Kalifornien) und kommt dort in Höhenlagen zwischen 0 und 600 Metern Meereshöhe vor.[4]
    • Dudleya candida Britton & Rose; Heimat: Mexiko (Baja California)
    • Dudleya cultrata Rose: Sie kommt im nordwestlichen Mexiko vor.[5]
    • Dudleya cymosa (Lem.) Britton & Rose; Heimat: Kalifornien und Oregon. Mit den Unterarten:
      • Dudleya cymosa subsp. agourensis K. M. Nakai: Sie kommt in Kalifornien in den Santa Monica Mountains in Höhenlagen zwischen 200 und 500 Metern Meereshöhe vor.[4]
      • Dudleya cymosa subsp. costatifolia Bartel & Shevock: Sie wird von manchen Autoren auch als Unterart Dudleya abramsii Rosesubsp. costatifolia (Bartel & Shevock) Moran zu Dudleya abramsii gestellt. Sie kommt in Kalifornien in Höhenlagen zwischen 1500 und 1700 Metern Meereshöhe vor.[4]
      • Dudleya cymosa subsp. crebrifolia K.M. Nakai & Verity: Sie ist ein Endemit der San Gabriel Mountains und kommt dort in Höhenlagen zwischen 300 und 600 Metern Meereshöhe vor.[4]
      • Dudleya cymosa subsp. cymosa: Sie kommt in Kalifornien in Höhenlagen zwischen 0 und 2700 Metern Meereshöhe vor.[4]
      • Dudleya cymosa subsp. marcescens Moran: Sie kommt in den Santa Monica Mountains, in Los Angeles und im Ventura County vor.[4]
      • Dudleya cymosa subsp. ovatifolia (Britton) Moran: Sie kommt in den Santa Monica Mountains und im Santa-Ana-Gebirge in Höhenlagen zwischen 200 und 400 Metern Meereshöhe vor.[4]
      • Dudleya cymosa subsp. paniculata (Jeps.) K.M. Nakai: Sie kommt in Kalifornien in Höhenlagen zwischen 30 und 1200 Metern Meereshöhe vor.[4]
      • Dudleya cymosa subsp. pumila (Rose) K.M. Nakai: Sie kommt in Kalifornien in Höhenlagen zwischen 100 und 2600 Metern Meereshöhe vor.[4]
    • Dudleya farinosa (Lindl.) Britton & Rose: Sie kommt in Kalifornien und in Oregon in Höhenlagen zwischen 0 und 600 Metern Meereshöhe vor.[4]
    • Dudleya gatesii Johans.: Sie kommt im nordwestlichen Mexiko vor.[5]
    • Dudleya gnoma S.W. McCabe: Sie kommt nur auf Santa Rosa Island vor.[4]
    • Dudleya greenei Rose: Sie kommt nur auf den kalifornischen Inseln Santa Cruz Island, Santa Catalina Island, San Miguel Island und Santa Rosa Island vor.[4]
    • Dudleya guadalupensis Moran: Sie kommt nur auf der Insel Guadalupe des mexikanischen Bundesstaates Baja California vor.[5]
    • Dudleya ingens Rose: Sie kommt in Baja California vor.[5]
    • Dudleya lanceolata (Nutt.) Britton & Rose: Sie kommt im mexikanischen Bundesstaat Baja California und in Kalifornien in Höhenlagen zwischen 0 und 1300 Metern Meereshöhe vor.[4]
    • Dudleya linearis (Greene) Britton & Rose: Sie kommt in Baja California vor.[5]
    • Dudleya nubigena (Brandegee) Britton & Rose (Syn.: Dudleya rigida Rose): Sie kommt im südlichen Baja California vor.[5] Mit den Unterarten:
      • Dudleya nubigena subsp. cerralvensis Moran
      • Dudleya nubigena subsp. nubigena
    • Dudleya pachyphytum Moran & M. Benedict: Sie kommt auf der Isla de Cedros vor.[5]
    • Dudleya palmeri (S. Watson) Britton & Rose: Sie kommt in Kalifornien von den Santa Lucia Mountains in Monterey County bis zu den Santa Monica Mountains in Los Angeles County in Höhenlagen zwischen 0 und 100 Metern Meereshöhe vor.[4]
    • Dudleya pauciflora Rose: Sie kommt in Baja California vor.[5]
    • Dudleya pulverulenta (Nutt.) Britton & Rose: Sie kommt vom südlichen Kalifornien bis zum nördlichen Baja California vor.[5]
    • Dudleya rigidiflora Rose: Sie kommt in Baja California vor.[5]
    • Dudleya rubens (Brandegee) Britton & Rose; Heimat: Mexiko (Baja California)[5]
    • Dudleya saxosa (M.E. Jones) Britton & Rose: Mit den Unterarten:
      • Dudleya saxosa subsp. aloides (Rose) Moran: Sie kommt in Kalifornien in Höhenlagen zwischen 200 und 1900 Metern Meereshöhe vor.[4]
      • Dudleya saxosa subsp. collomiae (Rose ex C.V. Morton) Moran: Sie kommt in Arizona in Höhenlagen zwischen 400 und 1800 Metern Meereshöhe vor.[4]
      • Dudleya saxosa subsp. saxosa: Sie kommt in Kalifornien in Höhenlagen zwischen 1100 und 2200 Metern Meereshöhe vor.[4]
    • Dudleya setchellii (Jeps.) Britton & Rose: Sie wird von manchen Autoren auch als Unterart Dudleya abramsii Rose subsp. setchellii (Jepson) Moran zu Dudleya abramsii gestellt. Sie kommt in Kalifornien in Höhenlagen zwischen 100 und 300 Metern Meereshöhe vor.[4]
    • Dudleya stolonifera Moran: Sie ist ein Endemit der San Joaquin Hills des Orange County in Kalifornien.[4]
    • Dudleya verityi K. Nakai: Sie kommt nur in den Santa Monica Mountains in Kalifornien vor.[4]
  • Untergattung Stylophyllum (Britton & Rose) Moran
    • Dudleya anomala (Davidson) Moran (Syn.: Stylophyllum anomalum Davidson): Sie kommt in Baja California vor.[5]>
    • Dudleya attenuata (S. Watson) Moran: Sie kommt in Kalifornien und im nördlichen Baja California vor.[5] Mit den Unterarten:
      • Dudleya attenuata subsp. attenuata (Syn.: Dudleya attenuata subsp. orcuttii (Rose) Moran): Sie kommt in Kalifornien und im mexikanischen Bundesstaaten Baja California in Höhenlagen zwischen 0 und 10 Metern Meereshöhe vor.[4]
      • Dudleya attenuata subsp. australis Moran: Sie kommt in Baja California vor.[4]
    • Dudleya campanulata Moran: Sie kommt im nördlichen Baja California vor.[5]
    • Dudleya densiflora (Rose) Moran; Heimat: Kalifornien, in Höhenlagen zwischen 100 und 600 Metern Meereshöhe.[4]
    • Dudleya edulis (Nutt.) Moran; Heimat: Südkalifornien, nördliches Baja California, in Höhenlagen zwischen 0 und 1300 Metern Meereshöhe.[4]
    • Dudleya formosa Moran: Sie kommt in Baja California Norte vor.[5]
    • Dudleya traskiae (Rose) Moran: Sie ist ein Endemit von Santa Barbara Island.[4]
    • Dudleya virens (Rose) Moran: Mit den Unterarten:
      • Dudleya virens subsp. extima Moran: Sie ist eine Endemit der Insel Guadalupe des mexikanischen Bundesstaates Baja California.[4]
      • Dudleya virens subsp. hassei (Rose) Moran: Sie ist ein Endemit von Santa Catalina Island.[4]
      • Dudleya virens subsp. insularis (Rose) Moran: Sie kommt in Kalifornien in Höhenlagen zwischen 0 und 300 Metern Meereshöhe vor.[4]
      • Dudleya virens subsp. virens: Sie ist eine Endemit von San Clemente Island.[4]
    • Dudleya viscida (S. Watson) Moran: Sie kommt in Kalifornien in Höhenlagen zwischen 0 und 500 Metern Meereshöhe vor.[4]
  • Untergattung Hasseanthus (Rose) Moran
    • Dudleya blochmaniae (Eastw.) Moran, mit den Unterarten:
      • Dudleya blochmaniae subsp. blochmaniae: Sie kommt in Kalifornien und in Baja California vor.[4]
      • Dudleya blochmaniae subsp. brevifolia (Moran) Moran: Sie wird von manchen Autoren auch als eigene Art angesehen: Dudleya brevifolia (Moran) Moran. Sie kommt in Kalifornien vor.[4]
      • Dudleya blochmaniae subsp. insularis (Moran) Moran: Sie ist eine Endemit von Santa Rosa Island (Kalifornien).[4]
    • Dudleya multicaulis(Rose) Moran: Sie kommt in Kalifornien in Höhenlagen zwischen 0 und 700 Metern Meereshöhe vor und ist eine Endemit der Küstenebene von Los Angeles und der angrenzenden Hügel.[4]
    • Dudleya nesiotica (Moran) Moran: Sie kommt in Kalifornien an der Küste in Höhenlagen zwischen 10 und 30 Metern Meereshöhe vor.[4]
    • Dudleya variegata (S. Watson) Moran; Heimat: Südkalifornien, Baja California, in Höhenlagen zwischen 0 und 500 Metern Meereshöhe.[4]

Zur Gattung gehören außerdem noch die Hybriden Dudleya × semiteres (Rose) Moran und Dudleya × sproulii P.H. Thomson.

Nachweise

Literatur

Einzelnachweise

  1. Lotte Burkhardt: Verzeichnis eponymischer Pflanzennamen – Erweiterte Edition. Teil I und II. Botanic Garden and Botanical Museum Berlin, Freie Universität Berlin, Berlin 2018, ISBN 978-3-946292-26-5 doi:10.3372/epolist2018.
  2. Bulletin of the New York Botanical Garden. Band 3, Nr. 9, S. 12. New York 1903
  3. Joachim Thiede: Dudleya. In: Urs Eggli: Sukkulenten-Lexikon Band 4. Crassulaceae (Dickblattgewächse). 2003, S. 87–105
  4. a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq Reid V. Moran : Dudleya Britton & Rose. In: Flora of North America, vol. 8. Crassulaceae. [1].
  5. a b c d e f g h i j k l m n o p q r s t Datenblatt Dudleya bei POWO = Plants of the World Online von Board of Trustees of the Royal Botanic Gardens, Kew: Kew Science.

Weblinks

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Dudleya: Brief Summary ( Alemão )

fornecido por wikipedia DE

Dudleya ist eine Pflanzengattung aus der Familie der Dickblattgewächse (Crassulaceae). Der botanische Name der Gattung ehrt den US-amerikanischen Botaniker William Russel Dudley (1849–1911).

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Dudleya ( Inglês )

fornecido por wikipedia EN

Dudleya, commonly known as liveforevers (Spanish: siemprevivas) is a genus of rosette-forming succulent plants in the stonecrop family, Crassulaceae, consisting of about 68 taxa in southwestern North America and Guadalupe Island. The species come in many forms, with some large and evergreen, others geophytic and deciduous. Yet, despite their dramatic variations in appearance, most species readily hybridize. The flowers of Dudleya have parts numbered in fives, with the petals arranged in tubular, star-shaped, and bell-shaped forms and, when fruiting, are filled with tiny, ovoid-crescent-shaped seeds.

The genus evolved as neoendemics, from ancestors in the stonecrop genus, Sedum. The ancestors radiated southward from Sedum during the creation of the dry summer climate, in the California region, five million years ago. Early botanists classified the larger species as Echeveria and Cotyledon, while the geophytic species were placed under Sedum. Taxonomic efforts, started by Joseph Nelson Rose and Nathaniel Lord Britton, created three genera; these initial genera were all eventually subsumed into Dudleya, proper, following Reid Moran's investigations into the genus. Phylogenetic research is still at an early stage in the genus, and is complicated by the fact that many species are becoming endangered and over-harvested (poached).

Dudleya is a relatively obscure genus, in comparison to other, more widely-cultivated succulents; converging interests, by succulent collectors, native plant enthusiasts and gardeners alike, have led to the wider cultivation of many species as ornamental plants. In the wild, many species of Dudleya are vulnerable, as land development and poachers often threaten particularly niche populations of plants. Poached plants are often shipped to East Asia, especially South Korea, where hybridisation and cultivation of succulents is very popular. Conservationists, nurseries and governments combat Dudleya poaching through propagation programs and protection laws.

Description

Characteristics and subgenera

This genus is the only taxon of the Sedeae tribe to have evolved sympodial branching.[2]

The genus is traditionally divided into three subgenera, two of which were formerly their own genera. The subgenera consist of Dudleya, Stylophyllum, and Hasseanthus.[3][4]

Epicuticular wax

Dudleya brittonii, which has the highest ultraviolet reflectivity ever discovered in a plant.

One of the most famous features of some Dudleya are their waxy coating. Numerous species have their foliage covered in an epicuticular wax, or farina.[5] This waxy coating is usually white, chalky, and mealy, and is the namesake in many epithets, like chalk dudleya, powdery liveforever, and giant chalk dudleya. The wax on the giant chalk dudleya is a surface with some of the highest ultraviolet reflectivity ever discovered in plants. When exposed to water, the wax may coat drops on leaves, preventing their evaporation.[10] The farina is mostly composed of pentacyclic triterpenoids.[11]

Other members, such as D. viscida and D. anomala, have a clear, sticky layer on their foliage.[12] This adhesive layer ultimately helps the plant protect its leaves from the sun, much like ones with "white" wax. It accomplishes this solar protection by allowing dust to adhere to the sticky layer on the leaves, which coats the foliage with dust and debris, blocking solar radiation.[13]

Taxonomy

Taxonomic history

Early history

The earliest species of the genus described was Cotyledon caespitosa, by Adrian Hardy Haworth in 1803, which would later be known as Dudleya caespitosa. The same species would later be described again as Sedum cotyledon by Joseph Franz von Jacquin in 1811, and in 1840 Thomas Nuttall described both Echeveria pulverulenta (Dudleya pulverulenta) and Echeveria lanceolata (Dudleya lanceolata). The differing generic placements of these early descriptions suggest that taxonomic disagreements over the genus had an early start, with some of these generic placements persisting even after the proper description of the genus by Britton and Rose.[3]

Nathaniel Lord Britton and Joseph Nelson Rose's revision of the North American Crassulaceae upended many of the early taxonomic classifications, with the newly-minted genus Dudleya containing 60 species, of which 41 were newly described by Britton and Rose. The two also defined the related genera Stylophyllum, which contained 12 species, and Hasseanthus, with 4 species. The primary differences between Dudleya and Stylophyllum were between the leaf shape, floral structure and petal orientation, whilst Hasseanthus was characterized by a different vegetative structure and more niche adaptations, primarily corm-like stems and deciduous leaves.[4]

In the 1930s, Alwin Berger revised the status of Dudleya and Stylophyllum into a sectional ranking within Echeveria, while merging the Hasseanthus genera into Sedum. This was in part due to a belief that Dudleya had evolved from Echeveria radiating northward from Mexico, evidenced by the similar tubular corollas, while Hasseanthus possessed aesthetic similarities to Sedum, primarily flowers with broadly-spreading petals. Stylophyllum was believed to occupy a transitory position between Dudleya and Hasseanthus. These conclusions were widely accepted by botanists for the first half of the 20th century, until proper molecular and phylogenetic analyses began to appear.[4]

Reid Moran, Charles H. Uhl and early phylogenetics

In 1942, Reid Moran, a botanist with a longstanding interest in the genera, offered a new revision of the taxa. Moran recognized Dudleya as being distinct from Echeveria, and merged Stylophyllum as a subgenera along with Eududleya (which was changed to just subgenus Dudleya). Moran still recognized Hasseanthus as a separate genus, but realized that it was closely related to Dudleya. With assistance from cytologist Charles H. Uhl, Moran came to the conclusion that Dudleya and Hasseanthus were related closer to each other than they were to Echeveria or Sedum.[4]

As Moran and Uhl conducted more cytological and taxonomic research on the genus, it became clear that Hasseanthus was also a subgenera of Dudleya, citing the karyological uniformity and the formation of hybrids between the genera.[4] In contrast, while Echeveria and Sedum freely hybridize within themselves and each other, all attempts at hybridizing Dudleya with Echeveria have failed. Uhl came to the conclusion that there are probably no intergeneric hybrids with Dudleya, believing that Dudleya had split from Echeveria far enough that intergeneric reproductive success was unlikely. Despite Uhl's research, he still believed that Dudleya had evolved northward from Mexico with Echeveria, probably some time before the formation of the summer-dry climate of California, making Dudleya a paleoendemic.[14]

Modern classifications and phylogenetic research

Around 1993, one of the only literary treatments of the genus was written, Paul H. Thomson's Dudleya and Hasseanthus Handbook.[6] In the book, Thompson made numerous changes and adjustments to species, and described several new species. The book included a large number of photographs, extensive descriptions, and horticultural practices for the care of Dudleya, derived from a long interest growing, visiting and collecting the plants. However, he failed to follow the International Code of Botanical Nomenclature guidelines for describing new taxa, forgoing designated type specimens, which invalidated his treatment.[15] His treatment also contains extensive pseudoscientific material, including references to the lost continents of Atlantis, Lemuria and Mu, a belief that cosmic rays and divine intervention directly induced the chromosome counts and evolution of Hasseanthus and Dudleya, and criticism of paleontologists and the scientifically accepted age of the Earth.[6]

Later research challenged the notion that Dudleya and Echeveria formed two distinct lineages within Echeverioideae. Joachim Thiede instead proposed that Dudleya evolved completely independently from Echeveria, evolving from relatives within the Leucosedum clade like Sedum moranii. This places Dudleya as a neoendemic that evolved recently with the formation of the summer-dry climate in California. This was evidenced by the similarities of seed-surface ornamentation between Dudleya and Sedum, which is unlike Echeveria, and the reproductive isolation between Echeveria and Dudleya.[3][16][17]

Research in the 21st century has supported the hypothesis of Dudleya's relation to Sedum. Phylogenetic analysis has given weight to this hypothesis, concluding that Dudleya is much closer to North American members of Sedoideae than to Echeverioideae. The plant Sedum spathulifolium, which co-exists with Dudleya in Oregon and California, has been show in multiple phylogenetic reconstructions to be a sister taxon to Dudleya, although it may not be the true sister taxon, as larger samplings are needed. Berger's hypothesis of a relationship between Hasseanthus and Sedum may still prove true, as in addition to morphological similarities, the Hasseanthus-type taxa could be basal to the genus. However, no truly conclusive evidence of significant divergence within Dudleya exists, and it remains unclear which groups are basal or derived.[18]

Selected species

Etymology

The genus is named after William Russel Dudley, the first head of the botany department at Stanford University.[5] The term liveforever may refer to the longevity and hardiness of some plants in the wild.[21]

The name of the subgenera Stylophyllum is an allusion to the pencil-shaped leaves of the type species,[22] Stylophyllum edule, now known as Dudleya edulis.[6]

The name of the subgenera Hasseanthus was in honor of Dr. Hermann E. Hasse, a surgeon and collector of California flora. Dudleya virens subsp. hassei is also named after him.[6]

Nomenclature

Reproductive biology

Pollinators and flower morphology

The pollinators of Dudleya are mostly hummingbirds and bees, although this has been inferred in some species only from flower morphology. The genus exhibits a wide array of diverse flower varieties, even within species complexes. The flowers are mostly hermaphroditic, although some individuals may have sterile anthers. Plants with long, tubular, reddish flowers, mostly in the subgenus Dudleya, are adapted towards pollination via hummingbirds, while short, spreading, yellow flowers favor pollination by insects. Despite the evolution of long flowers towards hummingbirds, the long flowers also show no performance disadvantage in pollination environments exclusively composed of insects. While the long flowers may provide an evolutionary advantage for more effective pollination, more energy is required to produce the larger amounts of floral tissue.[24]

The flower morphology of Dudleya also corresponds with the aforementioned groups of Hasseanthus, Stylophyllum, and Dudleya. The flowers in Hasseanthus and Stylophyllum are broad, white to yellow, and spreading, with bumblebees and bees as primary pollinators, and hummingbirds as occasional pollinators. As these species grade towards the subgenus Dudleya, the petals begin to fuse at the base, with the flowers becoming tubular and red, due to an association with hummingbird pollinators. This shift in pollinators corresponds to larger nectar volumes and higher energy contents in accordance with the demands of hummingbird pollinators.[3]

Seeds

Seed morphology also differs between Dudleya. Larger Dudleya have noticeably smaller seeds; this is likely a result of an evolutionary tradeoff. The investment in growing large allows plants to retain moisture in periods of drought, at the cost of energy placed into reproduction. Other reasons for smaller seeds may also be due to allocation of resources into vegetative reproduction via pup rosettes, instead of sexual reproduction. In contrast, the small, inconspicuous, Hasseanthus-type members produce the largest seeds in the genus. These larger seeds are not distributed very far, which contributes to the restricted ranges of the Hasseanthus subgenus.[7]

Distribution and habitat

Dudleya species are widespread and diverse in their range, but are typically found in rock outcroppings, cliff faces, or road cuts, where their leaves help them store water in a setting too dry for most types of plants. Most are small and inconspicuous when not in bloom.[5] The two predominant habitats where plants of this genera may be found are by the coast or mountains. Dudleya favor moderate temperatures, summer dormancy, winter precipitation, and rocky habitats, which means they may be found in diverse, disjunct locales from oceanic bluffs on the California coast to sky islands in Arizona.[3]

Dudleya gnoma, which grows on Miocene volcanic substrates on Santa Rosa Island

Regions where Dudleya can be found include The Californias, Arizona, coastal Sonora and Oregon, and southern Utah and Nevada.[3] The diversity of species of Dudleya is centered in Southern California and northern Baja California.[16]

Dudleya albiflora, found on the Baja California peninsula

Horticulture

Watering

In horticulture, Dudleya should be planted at an angle.[25] This allows accumulated water to drain from the nestlike center of the plant, thus preventing microbial decay.[26] Dudleya should not be watered from directly above, as this may damage their chalky coating known as farina, which is present on numerous species.[6][25]

Dudleya candida in cultivation

During the late fall to early spring period of growing, watering should be done frequently in order to promote good growth and flower development. Plants grown with exposure to rainfall, provided they are in a Mediterranean climate, they may obtain optimum moisture. In contrast, plants grown indoors or in greenhouses should be watered to ensure that the plant maximizes growth.[6]

During the dormant period, from late spring to fall, watering should be limited to bi-weekly intervals, but not completely cut out. In nature, plants may completely desiccate themselves during the dormant period.[25] Deciduous members of the genus in the Hasseanthus grouping should not be watered at all during dormancy.[6]

Growing medium

Growing mediums should attempt to imitate the preferred soil of each species in nature. The most essential element for the medium is good drainage.[25] Good drainage is important, as Dudleya are very susceptible to mold and fungi. How quickly the medium drains should be determined by the amount of shade in the location, which affects the rate of evaporation that the soil will go through. Thus, well-shaded Dudleya must have excellent drainage.[6]

Fertilizers may be used to maintain good color and growth, but they should be diluted.[6][27]

Clay is preferable to plastic pots when growing the plants due to the advantages with water drainage. Plastic pots may also kill the plants by overheating the roots.[6]

Mealybugs and aphids are main pests of Dudleya.[28][29] Powdery mildew has also been recorded in cultivated Dudleya.[30]

Shade

The amount of shade a species of Dudleya requires is dependent on the location. Plants growing in coastal regions may require little shade at all, whilst plants growing in the deserts, inland valleys and mountains will require shade. The majority of plants in the genus will appreciate north-facing sites and shade during the heat of the day. During the summer months, 50% shade may be beneficial for plants. If a cold-tolerant Dudleya is grown during a freeze or snow, it should be shaded as not to damage the plant, as a quick thaw may be detrimental.[6]

Propagation

Unlike their related genera Echeveria, many Dudleya cannot be propagated through leaf cuttings. Propagation is mostly achieved via offsets, germination by seed, or in nurseries, plant tissue culture.[6]

Dudleya variegata, one of the few species that can be propagated from the leaves

Sexual reproduction

Dudleya seeds are crescent shaped and near-microscopic, and can be collected by taking the dried flower buds and crushing the ovules inside, which can contain hundreds of seeds. Seeds can be sown into a soilless medium, such as pumice or vermiculite, after which germination can occur within a week to 10 days.[31]

Asexual reproduction

Vegetative reproduction may occur in multiple ways:[6]

  1. The rosette of the plant divides into two, eventually branching off to develop two separate rosettes.
  2. Plantlets will form in the axillary buds of old leaves around the primary rosettes, forming as many as 10 new rosettes.
  3. After a wildfire, when the plant has its main rosettes burned off, new ones will emerge from either the stem or the roots, forming a new plant within a single growing season.
  4. Rosettes emerge out of a long, horizontal root. This is commonplace when the plant has been decapitated on a cliff face, leaving only the long roots, which send out rosettes.
  5. Plants multiply via stoloniferous growth. Many Dudleya may send out stoloniferous rosettes, but they do not root. Only does the Laguna Beach Liveforever (Dudleya stolonifera) sprout roots out of stoloniferous plantlets, yet this can be difficult or impossible to induce in captivity.
  6. When the underground portion of the plant is lost to root rot, but the rosette and part of the caudex survive, the parts above the decayed roots can be removed cleanly, and planted provided they send out roots. Conversely, should the top of the rosette die but the lower stem still live, the deceased portion may be removed, a depression will remain in the caudex where the decayed rosette was removed, and at the edges new buds will sprout.
  7. Propagation via leaves. Only certain species, like those of the Hasseanthus subgenus and others like Dudleya parva, will root.[32] Most Dudleya will not reproduce from leaves, with the leaves simply dying after being severed.[6]
  8. Plant tissue culture methods are also available, and becoming more advanced. Currently, plant tissue culture is used for Dudleya propagation in commercial and conservation settings.[33]

Ethnobotany

Kumeyaay and Paipai

The indigenous peoples of the Kumeyaay and Paipai region utilized the genus for both medicinal and agricultural purposes. The tender, succulent leaves were chewed on to alleviate thirst, or used to treat calluses and corns. The budding inflorescences, in their early stages, were used as food, with a sweet flavor and juicy texture. The roots were pounded up and soaked in water, used as an astringent to "tighten the gums." The roots were also boiled whole as a decoction for asthma.[23][34]

Conservation

Several species of Dudleya are threatened by urban development in Coastal California and Mexico, and anthropogenic-induced wildfires. However, one of the most critical threats to Dudleya species is poaching, partially caused by a demand from East Asian succulent collectors paying lucrative prices for certain Dudleya species.[35]

Dudleya farinosa, which was threatened by poachers

The population of Verity's liveforever (Dudleya verityi), which was nearly wiped out during the 2013 Springs Fire, was targeted by poachers.[36][37]

The Cedros Island liveforever (Dudleya pachyphytum) is a rare and extremely specialized Dudleya endemic to Cedros Island. In 2016, Korean nationals began moving to Bahia Tortugas, a locality in Baja California Sur, to facilitate the poaching and shipping of the plants.[38] The species was seriously threatened after Mexican soldiers discovered poachers taking nearly 5,000 rosettes in a tractor-trailer.[39] It was suspected the poachers rappelled onto the succulent's location via helicopter, as D. pachyphytum occupies a nearly inaccessible habitat.[40] In 2019, the deaths and injuries of several fishermen from Bahía Tortugas who were on Cedros Island was allegedly the result of Dudleya trafficking, a conflict with the Sinaloa Cartel, or both.[41] In 2020, the Mexican Navy in the Second Naval Region revealed that two fishermen were killed after a dispute emerged over the trafficking of the rare plant.[42][43]

Bluff lettuce (Dudleya farinosa) was also targeted by poachers in numerous large-scale operations. Although not particularly rare, the size of the poaching operation pose a serious ecological threat. Starting in 2017, the U.S. Customs and Border Protection, along with the California Department of Fish and Wildlife, discovered large amounts of D. farinosa being shipped out of the country. South Korean and Chinese nationals have both been arrested in the smuggling of D. farinosa. According to nursery owners responsible for legally exporting Dudleya, buyers in Asia desired plants directly from the wild, owing to the aesthetic quality of their long caudices and weathered leaves.[44][45]

The candleholder dudleya (Dudleya candelabrum), native to the northern Channel Islands, was reported to have been poached, with the thieves shipping plants to South Korea.[46][47]

In response to the poaching of Dudleya, California State Assembly member Chris Ward proposed Bill AB-223, sponsored by the California Native Plant Society, which would make it illegal to poach Dudleya from state or private lands without a permit.[48] The California Native Plant Society and conservationists have also initiated propagation programs to oversaturate the market as a means to deter poachers.[31][33] On September 28, 2021, governor Gavin Newsom signed AB-223 into law.[49]

See also

References

  1. ^ Britton, N. L. & Rose, J. N.: Dudleya Britton & Rose, gen. nov.
  2. ^ Messerschmid, Thibaud; Klein, Johannes T. (September 2020). "Linnaeus's folly -phylogeny, evolution and classification of Sedum (Crassulaceae) and Crassulaceae subfamily Sempervivoideae". Taxon. 69 (45): 892–926. doi:10.1002/tax.12316. S2CID 225261669 – via ResearchGate.
  3. ^ a b c d e f g h i Thiede, J. (2004). The genus Dudleya Britton & Rose (Crassulaceae): its systematics and biology. Cactus and Succulent Journal (US), 76, 224-231.
  4. ^ a b c d e f g h Uhl, C. H., & Moran, R. (1953). The cytotaxonomy of Dudleya and Hasseanthus. American Journal of Botany, 492-502.
  5. ^ a b c d e f g McCabe, Stephen Ward. "Dudleya, in Jepson eFlora". Jepson Flora Project (eds.). Jepson Herbarium. Retrieved 8 June 2020.
  6. ^ a b c d e f g h i j k l m n o p Thomson, Paul H. (1993). Dudleya and Hasseanthus Handbook. Bonsall Publications. ISBN 978-0960206650.
  7. ^ a b Amoroso, D. M., & Wilson, P. (2018). Ten Cases of Divergence in the Seedling Ecology of Dudleya (Crassulaceae). Systematic Botany, 43(4), 889-900.
  8. ^ Platt, John R. (12 January 2017). "Meet the Endangered Plant Named after Rock Legend Jimi Hendrix". SCIENTIFIC AMERICAN. Retrieved 5 October 2021.
  9. ^ McCabe, Stephen W. (9 January 2017). "Two species of Dudleya were named in recent years" (PDF). International Crassulaceae Network. Archived (PDF) from the original on 28 January 2021. Retrieved 5 October 2021.
  10. ^ Spectral Properties of Heavily Glaucous and Non-Glaucous Leaves of a Succulent Rosette-Plant, Thomas W. Mulroy, Oecologia, 1979, [1]
  11. ^ Manheim Jr, B. S., & Mulroy, T. W. (1978). Triterpenoids in epicuticular waxes of Dudleya species. Phytochemistry, 17(10), 1799-1800.
  12. ^ Dudleya anomala (Davidson) Moran. Desert Plant Life. 14: 191. 1942.
  13. ^ Heiple, Paul. "Growing and Appreciating California Succulents in the Garden and in the Wild". YouTube. Archived from the original on 13 December 2021.
  14. ^ Uhl, Charles H. (1994). "INTERGENERIC HYBRIDS IN THE MEXICAN CRASSULACEAE: II. DUDLEYA (AND PLATE TECTONICS)". Cactus & Succulent Journal of America.
  15. ^ a b Rebman, J. P.; Gibson, J.; Rich, K. (2016). "Annotated checklist of the vascular plants of Baja California, Mexico" (PDF). San Diego Society of Natural History. 45. The green "form" of this species should likely be recognized as a separate species but the description by P.H. Thomson apparently lacks a designated type and may be invalidly published.
  16. ^ a b Thiede, J., & Eggli, U. (2007). Crassulaceae. In Flowering Plants· Eudicots (pp. 83-118). Springer, Berlin, Heidelberg. Chicago
  17. ^ Hart, H. T., & Eggli, U. (1995). Evolution and systematics of the Crassulaceae. Backhuys.
  18. ^ Yost, J. M.; Bontrager, M.; McCabe, S. W.; Burton, D.; Simpson, M. G.; Kay, K. M.; Ritter, M. (2013). "Phylogenetic relationships and evolution in Dudleya (Crassulaceae)" (PDF). Systematic Botany. 38 (4): 1096–1104. doi:10.1600/036364413X674760. S2CID 15715233.
  19. ^ Dodero, M. W. and M. G. Simpson. (2012). "Dudleya crassifolia (Crassulaceae), a new species from northern Baja California, Mexico". Madroño 59(4) 223–229.
  20. ^ Said to have been published in the December 2016 issue of Madroño.
  21. ^ Griffith, Caroline (28 June 2021). "Proposed Bill Would Stop Succulent Poaching". the Northcoast Environmental Center. Retrieved 18 October 2021. Dudleyas are also known as "liveforevers" due to their individual longevity and hardiness, a somewhat deceptive name given the rarity of many species.
  22. ^ Britton, N. L. & Rose, J. N.: Stylophyllum Britton & Rose, gen. nov.
  23. ^ a b c Wilken, Michael A. (2012) An Ethnobotany of Baja California's Kumeyaay Indians. Retrieved 13 October 2021
  24. ^ Aigner, P. A. (2005). "Variation in pollination performance gradients in a Dudleya species complex: can generalization promote floral divergence?". Functional Ecology. 19 (4): 681–689. doi:10.1111/j.1365-2435.2005.01009.x.
  25. ^ a b c d Zane, Nadia (9 August 2015). "Native Succulents for Water-Wise Gardens". San Joaquin UCCE Master Gardeners. Archived from the original on 27 September 2015.
  26. ^ Wilson, Bert; Wilson, Celeste (9 July 2013). "Dudleya pulverulenta". Las Pilitas Nursery. Archived from the original on 16 March 2015.
  27. ^ Kasten, Roy (2004). "Dudleya". Henry Shaw Cactus and Succulent Society. Archived from the original on 30 September 2021.
  28. ^ Hodgkiss, R.J. (5 February 2021). "Dudleya - Crassulaceae". The Succulent Plant Page. Archived from the original on 25 October 2007.
  29. ^ Bornstein, Carol; Fross, David; O'Brien, Bart (2005). California Native Plants for the Garden. Los Olivos, California: Cachuma Press. ISBN 0-9628505-8-6.
  30. ^ Henricot, B. (2008). Occurrence of powdery mildew (Erysiphe sp.) on Echeveria spp., Crassula spp., Cotyledon and Dudleya in the UK. Plant Pathology, 57(4).
  31. ^ a b Chesnut, John (October 2018). "Defeat Dudleya Poaching Through Propagation". CNPS-SLO. California Native Plant Society. Archived from the original on 17 August 2021.
  32. ^ Dudleya abramsii ssp. parva ( = Dudleya parva )(Conejo Dudleya) 5-Year Review: Summary and Evaluation (PDF). Ventura Fish and Wildlife Office, Ventura, California: U.S. Fish and Wildlife Service. March 2015. In a recent greenhouse study, conducted in Santa Cruz, California, by Stephen McCabe and Rachel Ormes, D. parva was rooted from the leaves of the plant, making it the only known species in the Dudleya subgenus to exhibit this behavior (McCabe in litt. 2008a).
  33. ^ a b Alison, Kevin (25 June 2020). "In-vitro cultivation of liveforevers (genus Dudleya) to reduce poaching and extinction". Orange County Chapter of the California Native Plant Society. doi:10.7280/D1NH4W – via Zenodo.
  34. ^ Hedges, K. (1986). Santa Ysabel Ethnobotany. San Diego Museum of Man.
  35. ^ Margulies, J. D. (2020). Korean ‘Housewives’ and ‘Hipsters’ Are Not Driving a New Illicit Plant Trade: Complicating Consumer Motivations Behind an Emergent Wildlife Trade in Dudleya farinosa. Frontiers in Ecology and Evolution, 8, 367.
  36. ^ Carlson, Cheri (April 18, 2019). "First a wildfire, then poachers. How a rare Ventura County plant has been nearly wiped out". Ventura County Star. August 16, 2021.
  37. ^ Townsend, Peggy (25 April 2019). "Succulent savior". UC Santa Cruz Newscenter. University of California, Santa Cruz. Archived from the original on 22 May 2019.
  38. ^ Vargas, Elizabeth (9 December 2019). "Planta endemica: movil sobre supuestos pescadros desparecidos". Ensenada.net. Archived from the original on 10 December 2019. Retrieved 2 January 2022.
  39. ^ "Militares decomisan plantas endemicas". MAS Noticias BCS. 28 May 2017.
  40. ^ McCabe, S.W. (September 24, 2019)
  41. ^ "CJNG en desaparición de pescadores". Investigaciones Zeta. 9 December 2019. Retrieved 21 August 2021.
  42. ^ "Reconoce Marina extracción y tráfico de "siempre viva" en Isla de Cedros". Investigaciones Zeta. 20 January 2021. Retrieved 21 August 2021.
  43. ^ Uhler, M. (2018). At the edge of California and the edge of existence: How siemprevive de Isla de Cedros (Dudleya pachyphytum [Crassulaceae]) was poached ever closer to extinction. CNPS 2018 Conservation Conference: Horticulture as a Part of Conservation. California Native Plant Society
  44. ^ Callahan, Mary (9 March 2019). "Plant smugglers take 'massive toll' on California's Dudleya farinosa succulent species". The Press Democrat. Retrieved 16 August 2021.
  45. ^ Robertson, Michelle (20 March 2019). "A poaching conspiracy is playing out on Northern California's coastline". SFGATE. Retrieved 16 August 2021.
  46. ^ Bradford, Steven (2021). "AB 223 Bill Analysis" (PDF). California State Senate Public Safety Committee – via California Senate. In 2020, posts on social media pointed to shipments of Dudleya candelabrum (candleholder liveforever) harvested from California and destined for sale in Korea. Candleholder liveforever is restricted to San Miguel, Santa Rosa, and Santa Cruz Islands off the coast of Ventura County.
  47. ^ Jensen, Nick (Spring 2021). "New Protections for Dudleya?" (PDF). Flora. 4: 30 – via California Native Plant Society. In 2020, for example, posts on social media suggested that shipments of candleholder dudleya (Dudleya candelabrum), a species restricted to San Miguel, Santa Rosa, and Santa Cruz Islands off the coast of Ventura County, were destined for sale overseas.
  48. ^ Jensen, Nick; Foy, Patrick (23 July 2021). "Plant poachers threaten California's biodiversity". CalMatters. Retrieved 16 August 2021.
  49. ^ Bryant, David (28 September 2021). "Governor Newsom signs CNPS-sponsored AB 223 into law, providing important new protections for California's imperiled dudleya". California Native Plant Society. Retrieved 25 October 2021.
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Dudleya: Brief Summary ( Inglês )

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Dudleya, commonly known as liveforevers (Spanish: siemprevivas) is a genus of rosette-forming succulent plants in the stonecrop family, Crassulaceae, consisting of about 68 taxa in southwestern North America and Guadalupe Island. The species come in many forms, with some large and evergreen, others geophytic and deciduous. Yet, despite their dramatic variations in appearance, most species readily hybridize. The flowers of Dudleya have parts numbered in fives, with the petals arranged in tubular, star-shaped, and bell-shaped forms and, when fruiting, are filled with tiny, ovoid-crescent-shaped seeds.

The genus evolved as neoendemics, from ancestors in the stonecrop genus, Sedum. The ancestors radiated southward from Sedum during the creation of the dry summer climate, in the California region, five million years ago. Early botanists classified the larger species as Echeveria and Cotyledon, while the geophytic species were placed under Sedum. Taxonomic efforts, started by Joseph Nelson Rose and Nathaniel Lord Britton, created three genera; these initial genera were all eventually subsumed into Dudleya, proper, following Reid Moran's investigations into the genus. Phylogenetic research is still at an early stage in the genus, and is complicated by the fact that many species are becoming endangered and over-harvested (poached).

Dudleya is a relatively obscure genus, in comparison to other, more widely-cultivated succulents; converging interests, by succulent collectors, native plant enthusiasts and gardeners alike, have led to the wider cultivation of many species as ornamental plants. In the wild, many species of Dudleya are vulnerable, as land development and poachers often threaten particularly niche populations of plants. Poached plants are often shipped to East Asia, especially South Korea, where hybridisation and cultivation of succulents is very popular. Conservationists, nurseries and governments combat Dudleya poaching through propagation programs and protection laws.

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Dudleya ( Espanhol; Castelhano )

fornecido por wikipedia ES

Dudleya es un género de plantas suculentas perteneciente a la familia Crassulaceae y que tiene unas 45 especies aceptadas.[1]​ Son nativas del sudoeste de Norteamérica.

Descripción

Son plantas herbáceas con hojas carnosas y glaucas que se desarrollan en una roseta basal, los colores varían del verde al gris. Las inflorescencias se encuentran en tallos verticales o inclinados a un metro de altura sobre la cima con brácteas como hojas alternas. Tiene pequeñas flores con cinco sépalos y cinco tépalos fusionados debajo, cinco pistilos, también fusionados y 10 estambres en su alrededor.

Las especies de Dudleya se encuentran ampliamente distribuidas en roquedales, acantilados, bordes de carreteras, donde sus hojas ayudan a almacenar el agua en lugares demasiados secos para otras plantas.

Taxonomía

El género fue descrito por Britton & Rose y publicado en New or Noteworthy North American Crassulaceae 12–28. 1903.[2][3]

Etimología

Es género fue nombrado en honor de William Russell Dudley, el primer director del departamento de botánica de la Universidad de Stanford.

Especies seleccionadas

Referencias

 title=
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Dudleya: Brief Summary ( Espanhol; Castelhano )

fornecido por wikipedia ES

Dudleya es un género de plantas suculentas perteneciente a la familia Crassulaceae y que tiene unas 45 especies aceptadas.​ Son nativas del sudoeste de Norteamérica.

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Hopearuusukkeet ( Finlandês )

fornecido por wikipedia FI

Hopearuusukkeet (Dudleya) on maksaruohokasveihin kuuluva mehikasvisuku, jossa on 40 lajia. Niitä esiintyy Yhdysvaltojen etelä- ja lounaisosissa seka Meksikon pohjois- ja luoteisosissa. Hopearuusukkeet ovat tyvestä lehtiruusukkeellisia monivuotisia sukkulenttikasveja. Niiden kukat ovat putkimaisia, kellon- tai tähdenmuotoisia, keltaisia, valkoisia tai punaisia.[1] Joitain hopearuusukelajeja viljellään huonekasvina.[2]

Lajeja

Lähteet

  1. Brickell, Christopher & Cathey, H. Marc (editors): A to Z Encyclopedia of Garden Plants (The American Horticultural Society). DK Publishing 2004. ISBN 0-7566-0616-0
  2. a b Räty, Ella (toim.): Viljelykasvien nimistö. Puutarhaliiton julkaisuja nro 363. Helsinki 2012. ISBN 978-951-8942-92-7
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Hopearuusukkeet: Brief Summary ( Finlandês )

fornecido por wikipedia FI

Hopearuusukkeet (Dudleya) on maksaruohokasveihin kuuluva mehikasvisuku, jossa on 40 lajia. Niitä esiintyy Yhdysvaltojen etelä- ja lounaisosissa seka Meksikon pohjois- ja luoteisosissa. Hopearuusukkeet ovat tyvestä lehtiruusukkeellisia monivuotisia sukkulenttikasveja. Niiden kukat ovat putkimaisia, kellon- tai tähdenmuotoisia, keltaisia, valkoisia tai punaisia. Joitain hopearuusukelajeja viljellään huonekasvina.

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Dudleya ( Croato )

fornecido por wikipedia hr Croatian

Dudleya je rod od oko 40 vrsta koje dolaze iz jugozapada SAD-a i Meksika. Neko vrijeme bile su smještene u rod Cotyledon, pa nakon toga u rod Echeveria dok na kraju nisu dobile svoj vlastiti rod. Ovaj rod je dobio ime po profesoru Williamu Dudleyu.

Dudleya su sukulentne rozete. Starije biljke se mogu rezati radi presađivanja. Nakon što smo odrezali dio biljke, taj dio treba ostaviti 3 dana na suhom da vidimo je li biljka zdrava, a nakon toga biljku treba presaditi u novu zemlju. Najbolje vrijeme za rezanje biljaka je kasno proljeće i ljeto. Ove biljke zahtijevaju puno sunca.

Ove vrste rastu zimi i ne podnose jako zalijevanje tokom ljetnih dana. Cvjetaju u proljeće.

Većina vrsta može podnijeti smrzavanje.

Vrste:

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Dudleya ( Neerlandês; Flamengo )

fornecido por wikipedia NL

Dudleya is een geslacht van succulenten uit de vetplantenfamilie (Crassulaceae). De soorten komen voor in het zuidwesten van Noord-Amerika.

Soorten

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Dudleya: Brief Summary ( Neerlandês; Flamengo )

fornecido por wikipedia NL

Dudleya is een geslacht van succulenten uit de vetplantenfamilie (Crassulaceae). De soorten komen voor in het zuidwesten van Noord-Amerika.

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Dudleya ( Polonês )

fornecido por wikipedia POL
Commons Multimedia w Wikimedia Commons

Dudleyarodzaj sukulentów występujących w płd.-zach. części Ameryki Północnej, obejmujący około 45 gatunków.

Systematyka

Pozycja systematyczna według Angiosperm Phylogeny Website (2001...)[1]

Rodzaj Dudleya należy do podrodziny Sempervivoideae, rodziny gruboszowatych (Crassualaceae), do rzędu skalnicowców (Saxifragales) i wraz z nim do okrytonasiennych.

Pozycja rodzaju w systemu Reveala (1993-1999)

Gromada okrytonasienne (Magnoliophyta Cronquist), podgromada Magnoliophytina Frohne & U. Jensen ex Reveal, klasa Rosopsida Batsch, podklasa różowe (Rosidae Takht.), nadrząd Saxifraganae Reveala, rząd skalnicowce (Saxifragales Dumort.), rodzina gruboszowate (Crassulaceae DC. in Lam. & DC.), rodzaj Dudleya[2].

Wybrane gatunki[3]

Przypisy

  1. a b P.F. Stevens: Angiosperm Phylogeny Website (ang.). 2001–. [dostęp 2009-07-11].
  2. James L. Reveal System of Classification. PBIO 250 Lecture Notes: Plant Taxonomy. Department of Plant Biology, University of Maryland, 1999. Systematyka rodzaju Dudleya według Reveala
  3. Germplasm Resources Information Network (GRIN). [dostęp 2010-01-25].
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Dudleya: Brief Summary ( Polonês )

fornecido por wikipedia POL

Dudleya – rodzaj sukulentów występujących w płd.-zach. części Ameryki Północnej, obejmujący około 45 gatunków.

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Dudleya ( Português )

fornecido por wikipedia PT

Dudleya é um género botânico pertencente à família Crassulaceae[1].

  1. «Dudleya — World Flora Online». www.worldfloraonline.org. Consultado em 19 de agosto de 2020
 title=
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Dudleya: Brief Summary ( Português )

fornecido por wikipedia PT

Dudleya é um género botânico pertencente à família Crassulaceae.

«Dudleya — World Flora Online». www.worldfloraonline.org. Consultado em 19 de agosto de 2020  title=
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Дудлея ( Ucraniano )

fornecido por wikipedia UK

Етимологія

Рід отримав назву від імені професора Стенфордського університету Вільяма Дадлі.

 src=
Dudleya blochmaniae subsp. brevifolia
 src=
Цвіте Dudleya caespitosa

Загальна біоморфологічна характеристика

Багаторічні сукулентні рослини з укороченим стеблом. Деякі види утворюють невеликий каудекс. Більшість видів — розеткові рослини, але деякі — чагарнички до 40 см заввишки. Стебла — одиночні або вилкоподібно розгалужені, низькі, часто дуже вкорочені. Листки — м'ясисті, соковиті, переважно ланцетні, видовжено-ланцетні, лінійні, дуже різноманітні за формою, сизі або зелені, нерідко мають вапняно-білий наліт. На верхівках пагонів листя по 50-100 штук зібрані в розетку діаметром від 2 до 50 см. Квітки — 5-членні, дзвоникоподібні, 0,7 — 1,5 см завдовжки, зібрані у верхівкові метельчаті або щитковидні суцвіття, мають жовтавий, червоний, рожевий або білий колір. Квітконоси із сильно редукованим листям виходять з пазух листків.

Дудлеї знаходяться у близькому рідстві з Ечеверіями і зовнішньо дуже схожі. В природі вони нерідко зростають поруч, однак дудлеї ростуть повільніше.

Гібриди серед мексиканських товстолистих дуже рідкісні в дикій природі, ймовірно, тому що ті деякі види, які зустрічаються разом і цвітуть, в той же час обслуговуються різними запильниками. Тим не менше, більшість видів і родів легко схрещуються у вирощуванні. Якобсен і Роулі у 1958 запропонували спеціальну назву для міжродових гібридів Dudleya з Echeveriaх Dudleveria.

Поширення

Рослини цього роду поширені на південному заході США і півночі Мексиці.

Охорона в природі

Види Dudleya traskiae і Dudleya stolonifera знаходяться на грані зникнення і їх продаж заборонений без спеціального дозволу.

Утримання в культурі

В культурі вирощують переважно розеточні дудлеї. В колекціях зустрічаються багато видів. Рослини утримують в світлих приміщеннях, на яскравому сонці злегка притінюють. З весни до осені регулярно поливають по мірі висихання ґрунту. Взимку утримують сухо при температурі 5 — 10 °C. Землесуміш — повітро і вологопроникна, складається з рівних частин лтистової землі і піску.

Розмножують переважно насінням, або підсушеними протягом декількох днів листовими живцями.

Колекція Ботанічного саду імені академіка Олександра Фоміна в Києві представлена вісьмома видами.

Види

За різними джерелами рід Дудлея налічує від 40 до 50 видів:

Види роду Dudleya за даними сайту «The Plant List»[2]

Примітки

Джерела

Література

  • Flora of North America Editorial Committee. 2009. Magnoliophyta: Paeoniaceae to Ericaceae. Fl. N. Amer. 8: i-xxiv, 1-585.

Посилання

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Dudleya ( Vietnamita )

fornecido por wikipedia VI

Dudleya là một chi thực vật có hoa trong họ Crassulaceae.[1]

Các loài

Chú thích

  1. ^ The Plant List (2010). Dudleya. Truy cập ngày 25 tháng 8 năm 2013.

Liên kết ngoài


Hình tượng sơ khai Bài viết chủ đề họ Lá bỏng này vẫn còn sơ khai. Bạn có thể giúp Wikipedia bằng cách mở rộng nội dung để bài được hoàn chỉnh hơn.
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Dudleya: Brief Summary ( Vietnamita )

fornecido por wikipedia VI

Dudleya là một chi thực vật có hoa trong họ Crassulaceae.

licença
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original
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wikipedia VI