Eunice fimbriata Grube 1870

Eunice fimbriata Grube, 1870

Eunice fimbriata Grube, 1870b:55–56.

MATERIAL EXAMINED.—Holotype, ZMH V-793; paratype ZMB Q2262, Viti Islands (Fiji) Mus. Godeffroy.

COMMENTS ON MATERIAL EXAMINED.—The holotype was originally described as consisting of 322 segments and being 230 mm long. The Hamburg specimen is complete with 362 setigers and is 230 mm long and is here considered the holotype. The Berlin specimen is incomplete and consists of 72 setigers.

DESCRIPTION.—Holotype of unknown sex, with 362 setigers, total length 230 mm; maximal width 7 mm at setiger 30; length through setiger 10, 11 mm; width at setiger 10, 6 mm. Paratype incomplete with 72 setigers; length 42 mm; maximal width 7 mm around setiger 30; length through setiger 10, 10 mm. Body cylindrical, dorsally covered with rugosities, mostly as short, longitudinal folds.

Prostomium (Figure 45h) distinctly shorter than peristomium, nearly as wide as peristomium, as deep as of peristomium. Prostomial lobes frontally rounded, dorsally excavate with thickened rim; median sulcus shallow. Antennae in transverse row, evenly spaced, similar in thickness, emerging from shallow depressions on prostomial face. Ceratophores ring-shaped in all antennae, without articulations. Ceratostyles short, digitiform, with 5 distinct articulations. All antennae to front edge of setiger 1. Peristomium cylindrical, with distinct muscular lower lip. Separation between rings distinct on all sides; anterior ring ∼ of total peristomial length. Peristomial cirri to middle of anterior peristomial ring, digitiform, with 3 articulations.

Maxillary formula of paratype 1+1, 4+4, 7+0, 5+8, and 1+1.

Branchiae (Figure 45i) present, pectinate, distinctly longer than notopodial cirri, not reduced in mid-body region, flexible. Branchiae from setiger 9 to setiger 122, thus absent on posterior half of body, present on less than 55% of total number of setigers. All branchiae except last 10–12 pairs with 2 or more filaments, maximum 5 filaments at about setiger 15. Branchial stems pliable, shorter than notopodial cirri. Filaments digitiform, much longer than notopodial cirri. Number of filaments reduced to 3 from about setiger 30, retaining same length and shape in all remaining branchial setigers.

Neuropodial acicular lobes asymmetrically rounded with high point above midline; aciculae emerging above midline. Presetal lobes low, transverse folds. Anterior postsetal lobes truncate folds, becoming symmetrically rounded in median and posterior setigers, outreaching acicular lobes. First 4 to 5 ventral cirri thick, tapering. Ventral cirri basally inflated in branchial region. Inflated bases transverse welts; narrow tips short and button-shaped. Postbranchial ventral cirri short, nearly tubercular, nearly invisible in last 100 setigers. Anterior notopodial cirri thick, digitiform, with 3 indistinct articulations. Articulations lost in branchial region. Notopodia decreasing in size towards posterior end of body, retaining similar shape.

Limbate setae slender, nearly straight; limbation marginally fraying in most setae. Pectinate setae (Figure 45n) short, with slender, cylindrical shafts. Blades flaring, flat. Both marginal teeth longer than other teeth, with ∼15 teeth. Shafts of compound falcigers (Figure 45j) long, slender, tapering, marginally smooth; beaks indistinct. Appendages (Figure 45j,m) short; heads distinct, bidentate. Proximal and distal teeth similar in length. Proximal teeth slender, tapering, directed obliquely distally, upturned near tips. Distal teeth curved, directed obliquely distally, nearly claw-like. Guards symmetrically rounded, marginally finely serrated; mucros absent, Pseudocompound falcigers and compound spinigers absent. Aciculae (Figure 45l) single, black, distally sharply bent ventrally; cross-section round. Separation between core and sheath distinct in both aciculae and subacicular hooks. Subacicular hooks (Figure 45k) black, bidentate. Hooks first present from setiger 26, present in all setigers thereafter, always single (except for replacements). Hooks tapering towards large heads. Proximal teeth larger than distal teeth; both teeth directed distally.

UNKNOWN MORPHOLOGICAL FEATURES.—Relationship between Mx III and left Mx IV; pygidium and anal cirri.

EXPECTED STATES OF SELECTED UNKNOWN FEATURES.—Mx III short, forming part of distal arc with left Mx IV.

CHARACTERS USED IN PREPARATION OF KEY NOT SCORED.—Inappropriate Characters: 56, 58, 59. Unknown Characters: 13, 14.

ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—None.