Diagnosis
provided by World Register of Marine Species
Oral arms with small parabolic or orbicular appendages throughout; females with a central flattened appendage surrounded by a whorl of numerous similar appendages of different sizes often folded, and males with parabolic or only small linear appendages in the central disc; trapezoid rhopaliar pit; five marginal lappets per paramere.
Gamero-Mora, E., Collins, A.G., Boco, S.R., Geson III, S.M., Morandini, A.C. (2022). Revealing hidden diversity among upside-down jellyfishes (Cnidaria: Scyphozoa: Rhizostomeae: Cassiopea): distinct evidence allows the change of status of a neglected variety and the description of a new species. Invertebrate Systematics. 36(1): 63-89.
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