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Eucladium Moss

Eucladium verticillatum Bruch & W. P. Schimper ex B. S. G. 1846

Comments

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According to P.-C. Chen (1941) Brotherus’s report (1929) of Eucladium verticillatum from Yunnan province was based on misidentification of a specimen collected by Handel-Mazzetti (no. 359, H). The specimen is Gymnostomum rupestre Schwaegr. (= G. aeruginosum Sm.).
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Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Moss Flora of China Vol. 2: 173 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Moss Flora of China @ eFloras.org
editor
Gao Chien & Marshall R. Crosby
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eFloras.org
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Comments

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Eucladium verticillatum is associated with year‑round seepage, possibly the reason that, although the stem may be reduced, the leaves are seldom reduced as they frequently are in drought-tolerant species of the family. The plants are typically a yellow-green, and there is usually a vivid contrast between the pellucid green laminal cells and the clear basal ones. Stem hyalodermis cells are frequently attached to the costa base, and form decurrencies of long and thin-walled cells in the leaf angles when leaves are removed. Gymnostomum and Molendoa are similar plants but differ by blunt apices and subpercurrent costae. Both these genera possess a stem central strand, no hyalodermis, and occasional 2-stratose areas in the lamina. Eucladium may be separated from these taxa and others that may be mistaken for it, such as Hymenostylium and Anoectangium by its highly differentiated bulging or lax, thin-walled basal cells. Assurances in the literature to the contrary, many other taxa also possess serrulate or denticulate leaf-shoulder margins, especially Hymenostylium, while Eucladium may rarely lack such serrulations. Specimens of Eucladium in which the lamina is highly reduced so that the leaves seem entirely costate are included without special rank in the range of variation of the species in North America. The laminal papillae of Hymenostylium are clear, sharp, well-defined; those of Eucladium are low, amorphous or scablike on the lumen surface.
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cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 27: 487, 560, 578 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
editor
Flora of North America Editorial Committee
project
eFloras.org
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eFloras

Description

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Plants 5–20 mm high, dark green to yellowish green or brownish, in dense tufts. Stems erect, usually repeatedly branched. Leaves 1.0–2.5 mm long, slightly contorted when dry, erect-spreading when moist, oblong-ovate at the base, gradually narrowed toward the apex, narrowly lanceolate, rounded or blunt at the apex; margins plane, entire at the leaf base, serrate above the base; costa stout, percurrent; upper leaf cells small, green, rounded-quadrate, 8–10 µm wide, thick-walled, papillose; basal cells larger, irregularly rectangular, thin-walled, smooth, hyaline, marginal cells linear. Setae slender, straight, 7–11 mm long; capsules erect, elliptic-cylindrical; annuli poorly developed, of one row of cells; peristome teeth yellowish brown, slightly twisted counterclockwise, 2–3-divided to the middle, often irregularly perforate, densely papillose; opercula conic, obliquely rostrate. Calyptrae cucullate, smooth. Spores 8–13 µm in diameter, pale yellowish, smooth.
license
cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Moss Flora of China Vol. 2: 173 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Moss Flora of China @ eFloras.org
editor
Gao Chien & Marshall R. Crosby
project
eFloras.org
original
visit source
partner site
eFloras

Description

provided by eFloras
Stems erect, 0.7-6 cm, leaves often branched in whorls in successive innovations. Leaves 1.2-2.5 mm, flat to channeled distally or at mid leaf, in transverse section at mid leaf often tapering from costa to leaf margin because of decreasing size of cells, mucro occasionally ending in a sharp, clear cell; marginal serrulations projecting from the distal end of cells of the hyaline cells of the leaf base; costa to 1/3 or more the width of the leaf near the base, basal cells 12-15 µm wide, 2-5:1; median and distal laminal cells 8-10 µm wide, 1(-2):1, walls rather thick, often irregularly so, large juxtacostally and decreasing in size to the leaf margin, irregular in shape from quadrate to rectangular, 1-2(-3):1, sometimes transversely elongate 2:1, occasionally with transverse walls oblique, marginal cells narrower in places and 2-3:1 occasionally approximating a border; papillae variously scattered or centered. Seta yellow to red‑brown, not or little twisted. Capsule 0.8-1.8 mm, red-brown when old; operculum 0.5-0.8 mm; peristome rudimentary or to 300 µm, yellow to orange, with a low, papillose basal membrane. Calyptra ca. 2.5 mm. Spores pale.
license
cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 27: 487, 560, 578 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
editor
Flora of North America Editorial Committee
project
eFloras.org
original
visit source
partner site
eFloras

Distribution

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Distribution: China, Japan, India, northern Africa, Russia, Europe, and North America.
license
cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Moss Flora of China Vol. 2: 173 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Moss Flora of China @ eFloras.org
editor
Gao Chien & Marshall R. Crosby
project
eFloras.org
original
visit source
partner site
eFloras

Habitat

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Habitat: on rocks or thin soil over rocks.
license
cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Moss Flora of China Vol. 2: 173 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Moss Flora of China @ eFloras.org
editor
Gao Chien & Marshall R. Crosby
project
eFloras.org
original
visit source
partner site
eFloras

Synonym

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Weissia verticillata Bridel, J. Bot. (Schrader) 1800(1): 283. 1801
license
cc-by-nc-sa-3.0
copyright
Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA
bibliographic citation
Flora of North America Vol. 27: 487, 560, 578 in eFloras.org, Missouri Botanical Garden. Accessed Nov 12, 2008.
source
Flora of North America @ eFloras.org
editor
Flora of North America Editorial Committee
project
eFloras.org
original
visit source
partner site
eFloras