dcsimg

Lifespan, longevity, and ageing

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Maximum longevity: 70 years (captivity) Observations: In the wild, these animals can live up to 26.8 years (http://bna.birds.cornell.edu/). There are anecdotal reports, which might be true, of animals living up to 70 years in captivity (John Terres 1980). IMR in the wild is probably less than 0.1. Breeding performance tends to decline after about age 12 (McCleery et al. 2008).
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Benefits

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Swans may attack people who approach their nests too closely. There are records of them knocking boaters off of jet skis. An adult swan can seriously injure children.

In addition, mute swans are thought to pose a threat to native wildlife as a result of competition for food, territories, and nesting areas.

Negative Impacts: injures humans

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Associations

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Mute swans are large and aggressive birds. As adults they are not often preyed on unless they are old or ill. Eggs and hatchlings are vulnerable to nest predation by raccoons, mink, and a wide variety of other medium to large-sized predators. But swan parents are typically present to protect their young.

Known Predators:

  • raccoons (Procyon lotor)
  • American minks (Neovison vison)
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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Alicia Ivory, University of Michigan-Ann Arbor
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Morphology

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Mute swans are large birds, measuring 144 to 158 cm. The wingspan is 2 to 2.5 meters. The two sexes are alike in appearance, except that males are generally larger than females. The plumage is white. They are best distinguished from North American swans by the knob at the base of the upper bill, and the color of the bill itself, which is orange, with the tip and base colored black. The head and neck may sometimes be stained brown from water and mud containing iron. (Reilly, 1968; Terres, 1980)

Range mass: 7600 to 14300 g.

Range length: 144 to 158 cm.

Range wingspan: 2 to 2.5 m.

Other Physical Features: endothermic ; homoiothermic; bilateral symmetry

Sexual Dimorphism: male larger

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Life Expectancy

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The greatest age recorded for a banded mute swan was 19 years. In captivity, they have lived 30 to 40 years.

Range lifespan
Status: wild:
19 (high) years.

Range lifespan
Status: captivity:
30-40 (high) years.

Average lifespan
Status: wild:
321 months.

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Habitat

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Mute swans are the most common swans in the wild, in parks or on country estates in their native range. In winter, they are more common on marine waters. They live in well-sheltered bays, open marshes, lakes, and ponds. (Reilly, 1968; Terres, 1980)

Habitat Regions: temperate ; saltwater or marine ; freshwater

Aquatic Biomes: lakes and ponds; rivers and streams; coastal

Wetlands: marsh

Other Habitat Features: estuarine

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Distribution

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Mute swans breed in the British Isles, north central Europe and north central Asia. They winter as far south as North Africa, the Near East, and to northwest India and Korea. They have been successfully introduced in North America, where they are a widespread species and permanent residents in many areas. (Reilly, 1968; Granlund, McPeek, and Adams, 1994)

Biogeographic Regions: nearctic (Introduced ); palearctic (Native ); ethiopian (Native )

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Alicia Ivory, University of Michigan-Ann Arbor
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Trophic Strategy

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The diet of mute swans consists of aquatic vegetation, and small proportions of aquatic insects, fish, and frogs. Mute swans do not dive, instead they plunge their head and long neck below the water's surface. Swans feed in deeper waters than ducks and other waterfowl that share their habitat and thus do not compete with them directly for food. Rather, food is made more readily available to other birds by swans because parts of the plants they consume float to the surface while the swans are feeding. However, mute swans compete with other swans for food because they feed in similar ways. (Reilly, 1968; Terres, 1980; http://aztec.inre.asu.edu/phxzoo/swanmute.html)

Animal Foods: insects; mollusks; aquatic or marine worms

Plant Foods: leaves; roots and tubers; algae

Primary Diet: herbivore (Folivore )

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Alicia Ivory, University of Michigan-Ann Arbor
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Associations

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Mute swans impact aquatic vegetation communities through their grazing.

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Alicia Ivory, University of Michigan-Ann Arbor
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Benefits

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Mute swans were domesticated for food in Britain. Markings on their feet indicated ownership. Eventual domestication saved the bird from becoming hunted to extinction there. Feathers were also used as quills for writing, the leathery web used for purses, and the wing bones for making whistles. ( http://www.airtime.co.uk.users/cygnus/muteswan/htm)

Positive Impacts: food ; body parts are source of valuable material

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Conservation Status

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The successful introduction and consequent expansion of mute swans into North America has begun to pose significant concerns to native wildlife. Common loons (threatened in Michigan) and recently re-introduced trumpeter swans are two species of primary concern. The North American population of mute swans has been increasing steadily since its introduction. These birds are aggressive, and have been known to drive off such stubborn and similarly sized species as Canada geese and trumpeter swans. Wildlife managers seek to control non-native mute swans in areas where native wildlife is being threatened. (Granlund, McPeek, and Adams, 1994)

There was a high incidence of lead poisoning in the mute swans of Great Britain, caused by the swans' ingestion of discarded lead shot that became entangled in aquatic vegetation. Since this problem was discovered, it is no longer a major threat to mute swan populations in Britain. ( http://www.airtime.co.uk/users/cygnus/muteswan.htm)

US Migratory Bird Act: no special status

US Federal List: no special status

CITES: no special status

State of Michigan List: no special status

IUCN Red List of Threatened Species: least concern

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Alicia Ivory, University of Michigan-Ann Arbor
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Behavior

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Mute swans have keen vision and hearing. Mute swans are usually silent, as the name suggests. Adults sometimes snort and make hissing noises or puppy-like barking notes or whistles, though the sounds are not far-reaching due to their straight trachea. Also, the sound of the wings during flight, which has been described as a musical throbbing or humming, is very audible. They also use visual displays as a form of communication, such as postures. For example, in an aggressive posture, males often arch their secondary wing feathers over the back.

Communication Channels: visual ; acoustic

Perception Channels: visual ; tactile ; acoustic ; chemical

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Reproduction

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Adults are not paired for life, contrary to the stereotype of the 'pining swan' who has lost its mate. In fact, some have been observed to have as many as four mates, or even 'divorce' one mate in favor of another. However, established pairs are more successful breeders than non-established pairs and mute swans do form monogamous pairs for at least a season.

Mating System: monogamous

Mute swans rarely nest in colonies. Nest sites are selected and breeding begins in March or early April. These swans either build a new nest or use a previously constructed mound, such as a muskrat house. The nest is large, made of aquatic vegetation, and lined with feathers and down. It is built well above the normal water level in swampy places near a pond or lake. It is possible for clutches of 5 to 12 to occur, but 5 to 7 is most common. The eggs are pale gray to pale blue-green. Incubation lasts 36 to 38 days. The chicks are brownish gray (gradually turning white within the next 12 months) and only remain in the nest for one day. The male may often take the first-hatched cygnet to the water while the female continues to incubate the remaining eggs. They are able to fly in about 60 days. Chicks can ride on the backs of their parents or under their wings. By the following breeding season the parents drive the young away. The cygnets then join flocks of other non-breeding swans, and during this time molt their feathers, becoming flightless for a short period of time. In the next two years, the cygnets begin to bond with a mate and begin to look for suitable breeding territory. Swans do not begin to breed until about their third year. (Granlund, McPeek and Adams, 1994; Reilly, 1968; Terres, 1980; http://www.airtime.co.uk/users/cygnus/muteswan.htm; http://www.aztec.inre.asu.edu/phxzoo/swanmute.html)

Breeding interval: Mute swans breed once yearly.

Breeding season: Breeding begins in March and April.

Range eggs per season: 5 to 12.

Average eggs per season: 5-7.

Range time to hatching: 36 to 38 days.

Average fledging age: 60 days.

Average time to independence: 12 months.

Average age at sexual or reproductive maturity (female): 3 years.

Average age at sexual or reproductive maturity (male): 3 years.

Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; oviparous

Average eggs per season: 5.

The sexes share incubation, though the female spends the majority of time sitting, and the male usually stands guard.

Even in semi-domestication, the nest is strongly defended; swans have been known to attack other waterfowl and even people. Blows from their powerful wings can be especially painful. They can be dangerous to children, and are capable of killing or maiming some of the larger predators.

Parental Investment: precocial ; pre-fertilization (Protecting: Female); pre-hatching/birth (Protecting: Male); pre-weaning/fledging (Provisioning: Male, Female, Protecting: Male, Female); pre-independence (Provisioning: Male, Female, Protecting: Male, Female)

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Ivory, A. 2002. "Cygnus olor" (On-line), Animal Diversity Web. Accessed April 27, 2013 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Cygnus_olor.html
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Alicia Ivory, University of Michigan-Ann Arbor
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Biology

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The mute swan feeds chiefly on submerged aquatic vegetation, which is obtained by upending (tipping head first into the water, so that the tail remains visible above the surface) (4). It also feeds in fields on young cereal crops (8), spilt grain (4), and on artificial food sources, such as bread given by the public (3). Territorial disputes may result in aggressive fights between males, in which they rush at one another and slide along the surface of the water (2). Pairs typically nest solitarily, although semi-domesticated birds may nest in large colonies (6) (notably at Abbotsbury in Dorset) (8). The cone-shaped nest is built at the edge of the water, and may be used in subsequent years by the same pair (4). After mid-April, between 5 and 7 (up to 12) whitish or pale blue eggs are laid. They are incubated, mainly by the female, for 35-42 days; the young, known as 'cygnets', leave the nest soon after hatching (4). Both parents take care of the cygnets for an extended period, often until the next breeding season (4).
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Conservation

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Strong lobbying to ban lead fishing weights has enabled mute swans to recover from the crash in numbers caused by lead poisoning. They will also have benefited from action carried out for other species of wildfowl, such as the creation and management of wetland nature reserves (8).
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Description

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The mute swan is Britain's largest bird (3), and one of the heaviest flying birds in the world (4); adults can weigh over 15 kg (3). The combination of their large size, very long neck, white plumage and orange-red bill with a black knob towards the top of the bill makes them easily to recognise (2). Males (cobs) and females (pens) are similar in appearance, although males are slightly larger and have a more prominent knob on the bill (4). Juveniles are greyish-brown with a grey bill, which lacks the knob seen in adults (2). Contrary to the name, the mute swan produces a range of vocalisations, including a rumbling 'heeorr', and an aggressive hissing noise when threatened (2). The wings make a loud 'clanking' in flight (8).
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Habitat

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Found in a wide range of water bodies, including rivers, lakes, ponds, reservoirs, flood waters, tidal estuaries, and sheltered coasts (4).
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Range

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Found throughout Britain, but absent from high ground and areas without fresh water (3). After 1960, the population began to decline as a result of poisoning from lead fishing weights (3). Since the mid-1980s and the banning of lead weights however, the population has increased (5). Outside of Britain, the mute swan is known throughout Europe and central Asia (6); it has also been introduced to New Zealand, Australia, South Africa, and North America (4).
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Status

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Widespread and common species, not listed under any conservation designations (3). Included in the Birds of Conservation Concern Amber List (medium conservation concern) (7).
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Threats

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Harsh winters and poisoning from lead fishing weights were responsible for the decline of the mute swan population in Britain. A series of mild winters combined with the banning of lead weights has resulted in the recovery of the numbers of this beautiful bird (5).
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Status in Egypt

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Winter visitor.

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Associations

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In Great Britain and/or Ireland:
Animal / parasite / endoparasite
fluke of Cotylurus cornutus endoparasitises small intestine of Cygnus olor

Animal / parasite / endoparasite
fluke of Filicollis anatis endoparasitises small intestine of Cygnus olor

Animal / parasite / endoparasite
tapeworm of Nematoparataenia southwelli endoparasitises small intestine of Cygnus olor

Animal / parasite / endoparasite
fluke of Polymorphus minutus endoparasitises small intestine of Cygnus olor

Animal / parasite / ectoparasite / blood sucker
Theromyzon tessulatum sucks the blood of nasal passage of Cygnus olor

Animal / parasite / endoparasite
fluke of Typhlocoelum sisowi endoparasitises trachea of Cygnus olor

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Mute swan

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Sound from wingbeats.

The mute swan (Cygnus olor) is a species of swan and a member of the waterfowl family Anatidae. It is native to much of Eurosiberia, and (as a rare winter visitor) the far north of Africa. It is an introduced species in North America, home to the largest populations outside of its native range, with additional smaller introductions in Australasia and southern Africa. The name 'mute' derives from it being less vocal than other swan species.[2][3][4] Measuring 125 to 160 cm (49 to 63 in) in length, this large swan is wholly white in plumage with an orange beak bordered with black. It is recognizable by its pronounced knob atop the beak, which is larger in males.

Taxonomy

The mute swan was first formally described by the German naturalist Johann Friedrich Gmelin as Anas olor in 1789, and was transferred by Johann Matthäus Bechstein to the new genus Cygnus in 1803. Both cygnus and olor mean "swan" in Latin; cygnus is a variant form of cycnus, a borrowing from Greek κύκνος kyknos, a word of the same meaning.[5][6][7][8]

Despite its Eurasian origin, its closest relatives are the black swan of Australia and the black-necked swan of South America, not the other Northern Hemisphere swans of the genus Cygnus.[2] The species is monotypic, with no living subspecies.[2][4]

Evolution

Mute swan subfossils, 6,000 years old, have been found in post-glacial peat beds of East Anglia, Great Britain.[9] They have been recorded from Ireland east to Portugal and Italy,[10] and from France, 13,000 BP (Desbrosse and Mourer-Chauvire 1972–1973). The paleosubspecies Cygnus olor bergmanni, which differed only in size from the living bird, is known from fossils found in Azerbaijan. A related paleospecies recorded from fossils and subfossils is the Giant swan, Cygnus falconeri, a flightless species which lived on the islands of Malta and Sicily during the Middle Pleistocene.

Fossils of swan ancestors more distantly allied to the mute swan have been found in four U.S. states: California, Arizona, Idaho and Oregon.[11] The timeline runs from the Miocene to the late Pleistocene, or 10,000 BP. The latest find was in Anza Borrego Desert, a state park in California.[11] Fossils from the Pleistocene include Cygnus paloregonus from Fossil Lake, Oregon, Froman's Ferry, Idaho, and Arizona, referred to by Howard in The Waterfowl of the World as "probably the mute type swan".[12]

Description

Adults of this large swan typically range from 140 to 160 cm (55 to 63 in) long, although can range in extreme cases from 125 to 170 cm (49 to 67 in), with a 200 to 240 cm (79 to 94 in) wingspan.[13][14] Males are larger than females and have a larger knob on their bill. On average, this is the second largest waterfowl species after the trumpeter swan, although male mute swans can easily match or even exceed a male trumpeter in mass.[4][15] Among standard measurements of the mute swan, the wing chord measures 53–62.3 cm (20.9–24.5 in), the tarsus is 10–11.8 cm (3.9–4.6 in) and the bill is 6.9–9 cm (2.7–3.5 in).[4] The plumage is white, while the legs are dark grey. The beak of the mute swan is bright orange, with black around the nostrils and a black nail.

The mute swan is one of the heaviest flying birds. In several studies from Great Britain, males (known as cobs) were found to average from about 10.6 to 11.87 kg (23.4 to 26.2 lb), with a weight range of 9.2–14.3 kg (20–32 lb) while the slightly smaller females (known as pens) averaged about 8.5 to 9.67 kg (18.7 to 21.3 lb), with a weight range of 7.6–10.6 kg (17–23 lb).[4][16][17][18][19] While the top normal weight for a big cob is roughly 15 kg (33 lb), one unusually big Polish cob weighed almost 23 kg (51 lb) and this counts as the largest weight ever verified for a flying bird, although it has been questioned whether this heavyweight could still take flight.[20]

Young birds, called cygnets, are not the bright white of mature adults, and their bill is dull greyish-black, not orange, for the first year. The down may range from pure white to grey to buff, with grey/buff the most common. The white cygnets have a leucistic gene. Cygnets grow quickly, reaching a size close to their adult size in approximately three months after hatching. Cygnets typically retain their grey feathers until they are at least one year old, with the down on their wings having been replaced by flight feathers earlier that year.

All mute swans are white at maturity, though the feathers (particularly on the head and neck) are often stained orange-brown by iron and tannins in the water.[21]

Polish swan

Two mute swan cygnets a few weeks old. The cygnet on the right is of the "Polish swan" colour morph, and carries a gene responsible for leucism.

The colour morph C. o. morpha immutabilis (immūtābilis is Latin for "immutable, unchangeable, unalterable"), also known as the "Polish swan", has pinkish (not dark grey) legs and dull white cygnets; as with white domestic geese, it is found only in populations with a history of domestication.[22][23] Polish swans carry a copy of a gene responsible for leucism.[23]

Behavior

Nest in Drilon, Pogradec, Albania. The cob (male) is patrolling the area close to the nest to protect his mate.

Mute swans nest on large mounds that they build with waterside vegetation in shallow water on islands in the middle or at the very edge of a lake. They are monogamous and often reuse the same nest each year, restoring or rebuilding it as needed. Male and female swans share the care of the nest, and once the cygnets are fledged it is not uncommon to see whole families looking for food. They feed on a wide range of vegetation, both submerged aquatic plants which they reach with their long necks, and by grazing on land. The food commonly includes agricultural crop plants such as oilseed rape and wheat, and feeding flocks in the winter may cause significant crop damage, often as much through trampling with their large webbed feet, as through direct consumption.[24] It will also feed on small proportions of aquatic insects, fish and frogs.[25]

Unlike black swans, mute swans are usually strongly territorial with just a single pair on smaller lakes, though in a few locations where a large area of suitable feeding habitat is found they can be colonial. The largest colonies have over 100 pairs, such as at the colony at Abbotsbury Swannery in southern England, and at the southern tip of Öland Island, Ottenby Preserve, in the coastal waters of the Baltic Sea, and can have nests spaced as little as 2 m (7 ft) apart.[22][26] Non-mated juveniles up to 3–4 years old commonly form larger flocks, which can total several hundred birds, often at regular traditional sites.[27] A notable flock of non-breeding birds is found on the River Tweed estuary at Berwick-upon-Tweed in northeastern England, with a maximum count of 787 birds.[28] A large population exists near the Swan Lifeline Station in Windsor, and live on the Thames in the shadow of Windsor Castle. Once the adults are mated they seek out their own territories and often live close to ducks and gulls, which may take advantage of the swan's ability to reach deep water weeds, which tend to spread out on the water surface.

The mute swan is less vocal than the noisy whooper and Bewick's swans; they do, however, make a variety of sounds, often described as "grunting, hoarse whistling, and snorting noises." During a courtship display, mute swans utter a rhythmic song. The song help synchronize the movements of their heads and necks. It could technically be employed to distinguish a bonded couple from two dating swans, as the rhythm of the song typically fails to match the pace of the head movements in two dating swans.[29] Mute swans usually hiss at competitors or intruders trying to enter their territory.[30] The most familiar sound associated with mute swans is the vibrant throbbing of the wings in flight which is unique to the species, and can be heard from a range of 1 to 2 km (0.6 to 1 mi), indicating its value as a contact sound between birds in flight.[22] Cygnets are especially vocal, and communicate through a variety of whistling and chirping sounds when content, as well as a harsh squawking noise when distressed or lost.

Nesting in spring, Cologne, Germany

Mute swans can be very aggressive in defence of their nests and are highly protective of their mate and offspring. Most defensive acts from a mute swan begin with a loud hiss and, if this is not sufficient to drive off the predator or intruder, are followed by a physical attack. Swans attack by striking at the threat with bony spurs in their wings, accompanied by biting with their large bill,[31] while smaller waterbirds such as ducks are normally grabbed with the swan's bill and dragged or thrown clear of the swan and its offspring. Swans will kill intruders into their territory, both other swans, and geese and ducks, by drowning, climbing onto and pecking the back of the head and forcing the other bird underwater.

The wings of the swan are very powerful, though not strong enough to break an adult man's leg, as is commonly misquoted.[32] Large waterfowl, such as Canada geese, (more likely out of competition than in response to potential predation) may be aggressively driven off, and mute swans regularly attack people who enter their territory.[33]

The cob is responsible for defending the cygnets while on the water, and will sometimes attack small watercraft, such as canoes, that it feels are a threat to its young. The cob will additionally try to chase the predator out of his family territory, and will keep animals such as foxes and raptors at bay. In New York (outside its native range), the most common predators of cygnets are common snapping turtles.[33] Healthy adults are rarely preyed upon, though canids such as coyotes, felids such as lynx, and bears can pose a threat to infirm ones (healthy adults can usually swim away from danger and nest defense is usually successful.) and there are a few cases of healthy adults falling prey to golden eagles.[34][35][36] In England, there has been an increased rate of attacks on swans by out-of-control dogs, especially in parks where the birds are less territorial. This is considered criminal in British law, and the birds are placed under the highest protection due to their association with the monarch.[37] Mute swans will readily attack dogs to protect themselves and their cygnets from an attack, and an adult swan is capable of overwhelming and drowning[38] even large dog breeds.[39]

The familiar pose with neck curved back and wings half raised, known as busking, is a threat display. Both feet are paddled in unison during this display, resulting in more jerky movement.[40] The swans may also use the busking posture for wind-assisted transportation over several hundred meters, so-called windsurfing.[41][42]

Like other swans, mute swans are known for their ability to grieve for a lost or dead mate or cygnet.[43][44] Swans will go through a mourning process, and in the case of the loss of their mate, may either stay where its counterpart lived, or fly off to join a flock.[45] Should one of the pair die while there are cygnets present, the remaining parent will take up their partner's duties in raising the clutch.

Breeding

Nest of a mute swan, Sweden
Cygnets captured one day after they hatched. Newburg Lake, Livonia, MI, U.S.
A three-day old cygnet

Mute swans lay from 4 to 10 eggs. The female broods for around 36 days, with cygnets normally hatching between the months of May and July.[46] The young swans do not achieve the ability to fly before about 120 to 150 days old. This limits the distribution of the species at the northern edge of its range as the cygnets need to learn to fly before the ponds and lakes freeze over.

Distribution and habitat

The mute swan is found naturally mainly in temperate areas of Europe then across the Palearctic as far east as Primorsky Krai, near Sidemi.[47]

It is partially migratory throughout northern latitudes in Europe and Asia, as far south as North Africa and the Mediterranean. It is known and recorded to have nested in Iceland and is a vagrant to that area as well as to Bermuda, according to the UN Environment Programme chart of international status chart of bird species, which places it in 70 countries, breeding in 49 countries, and vagrant in 16 countries. While most of the current population in Japan is introduced, mute swans are depicted on scrolls more than 1,000 years old, and wild birds from the mainland Asian population still occur rarely in winter. Natural migrants to Japan usually occur along with whooper and sometimes Bewick's swans.

The mute swan is protected in most of its range, but this has not prevented illegal hunting and poaching. It is often kept in captivity outside its natural range, as a decoration for parks and ponds, and escapes have happened. The descendants of such birds have become naturalised in the eastern United States and Great Lakes, much as the Canada goose has done in Europe.

World population

Native populations

Mute swans with cygnets in Wolvercote, Oxfordshire
An older mute swan cygnet in Haut-Rhin, France

The total native population of mute swans is about 500,000 birds at the end of the breeding season (adults plus young), of which up to 350,000 are in Russia.[2] The largest single breeding concentration is 11,000 pairs in the Volga Delta.[3]

The population in the United Kingdom is about 22,000 birds as of the 2006–2007 winter,[48] a slight decline from the peak of about 26,000–27,000 birds in 1990.[3] This includes about 5,300 breeding pairs, the remainder being immatures.[49] Other significant populations in Europe include 6,800–8,300 breeding pairs in Germany, 4,500 pairs in Denmark, 4,000–4,200 pairs in Poland, 3,000–4,000 pairs in the Netherlands, about 2,500 pairs in Ireland, and 1,200–1,700 pairs in Ukraine.[3]

For many centuries, mute swans in Great Britain were domesticated for food, with individuals being marked by nicks on their webs (feet) or beaks to indicate ownership. These marks were registered with the Crown and a Royal Swanherd was appointed. Any birds not so marked became Crown property, hence the swan becoming known as the "Royal Bird". It is quite possible that this domestication saved the mute swan from extirpation through overhunting in Great Britain.[50][51]

Populations in western Europe were largely exterminated by hunting pressure in the 13th–19th centuries, with the exception of semi-domesticated birds maintained as poultry by large landowners. Better protection in the late 19th and early 20th centuries allowed the species to expand and return to most or all of their former range.[52][53] More recently in the period from about 1960 up to the early 1980s, numbers declined significantly again in many areas in England,[54] primarily due to lead poisoning from birds swallowing lead shot from shooting and discarded fishing weights made from lead. After lead weights and shot were mostly replaced by other less toxic alternatives, mute swan numbers increased again rapidly.[3]

Introduced populations

Since being introduced into North America, the mute swan has increased greatly in number to the extent that it is considered as an invasive species there. Populations introduced into other areas remain small, with around 200 in Japan, fewer than 200 in New Zealand and Australia, and about 120 in South Africa.[2]

North America

The mute swan was introduced to North America in the late 19th century. Recently, it has been widely viewed as an invasive species because of its rapidly increasing numbers and its adverse effects on other waterfowl and native ecosystems. For example, a study of population sizes in the lower Great Lakes from 1971 to 2000 found that mute swan numbers were increasing at an average rate of at least 10% per year, doubling the population every seven to eight years.[55] Several studies have concluded that mute swans severely reduce densities of submerged vegetation where they occur.[56]

In 2003, the U.S. Fish and Wildlife Service proposed to "minimize environmental damages attributed to Mute Swans" by reducing their numbers in the Atlantic Flyway to pre-1986 levels, a 67% reduction at the time. According to a report published in the Federal Register of 2003[57] the proposal was supported by all thirteen state wildlife agencies which submitted comments, as well as by 43 bird conservation, wildlife conservation and wildlife management organisations. Ten animal rights organisations and the vast majority of comments from individuals were opposed. At this time mute swans were protected under the Migratory Bird Treaty Act due to a court order, but in 2005 the United States Department of the Interior officially declared them a non-native, unprotected species.[58] Mute swans are protected in some areas of the U.S. by local laws as, for example, in Connecticut.[59]

The status of the mute swan as an introduced species in North America is disputed by the interest group "Save the Mute Swans".[60] They assert that mute swans are native in the region and therefore deserving of protection. They claim that mute swans had origins from Russia and cite historical sightings and fossil records. These claims have been rejected as specious by the U.S. Department of the Interior.[58]

Oceania

The mute swan had absolute protection in New Zealand under the Wildlife Act 1953, but this was changed in June 2010 to a lower level of protection. It still has protection, but is now allowed to be killed or held in captivity at the discretion of the Minister of Conservation.[61]

A small feral population exists in the vicinity of Perth, Australia; however, it is believed to number less than 100 individuals.

In popular culture

The mute swan has been the national bird of Denmark since 1984. Prior to that, the skylark was considered Denmark's national bird (since 1960).[62]

The fairy tale "The Ugly Duckling" by Hans Christian Andersen tells the story of a cygnet ostracised by his fellow barnyard fowl because of his perceived unattractiveness. To his delight (and to the surprise of others), he matures into a graceful swan, the most beautiful bird of all.[62]

Today, the British Monarch retains the right to ownership of all unmarked mute swans in open water, but King Charles III exercises his ownership only on certain stretches of the Thames and its surrounding tributaries. This ownership is shared with the Vintners' and Dyers' Companies, who were granted rights of ownership by the Crown in the 15th century.[63][64]

The mute swans in the moat at the Bishops Palace at Wells Cathedral in Wells, England have for centuries been trained to ring bells via strings attached to them to beg for food. Two swans are still able to ring for lunch.[65]

The pair of swans in the Boston Public Garden are named Romeo and Juliet after the Shakespearean couple; however, it was found that both of them are females.[66]

Gallery

See also

References

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  2. ^ a b c d e del Hoyo, Josep; Elliott, Andrew; Sargatal, Jordi, eds. (1992). Ostrich to Ducks. Handbook of the Birds of the World. Vol. 1. Barcelona: Lynx Edicions. pp. 577–78. ISBN 978-84-87334-10-8.
  3. ^ a b c d e Snow, D. W.; Perrins, C. M. (1998). The Birds of the Western Palearctic (Concise ed.). Oxford: Oxford University Press. ISBN 978-0-19-854099-1.
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  5. ^ cycnus, olor. Charlton T. Lewis and Charles Short. A Latin Dictionary on Perseus Project.
  6. ^ κύκνος. Liddell, Henry George; Scott, Robert; A Greek–English Lexicon at the Perseus Project.
  7. ^ Simpson, D. P. (1979). Cassell's Latin Dictionary (5th ed.). London: Cassell. ISBN 978-0-304-52257-6.
  8. ^ Harper, Douglas. "cygnet". Online Etymology Dictionary.
  9. ^ Northcote, E. M. (1981). "Size difference between limb bones of recent and subfossil Mute Swans (Cygnus olor)". J. Archaeol. Sci. 8 (1): 89–98. doi:10.1016/0305-4403(81)90014-5.
  10. ^ Palmer, Ralph S., ed. (1976). Handbook of North American Birds. Vol. 2. New Haven: Yale University Press. ISBN 978-0300019025.
  11. ^ a b Jefferson, George T.; Lindsay, Lowell, eds. (2005). Fossil Treasures of the Anza-Borrego Desert. Sunbelt Publications. p. 153. ISBN 9780932653505.
  12. ^ The Waterfowl of the World. pp. 262–265.
  13. ^ Madge, Steve, Waterfowl: An Identification Guide to the Ducks, Geese, and Swans of the World. Houghton Mifflin Harcourt (1992), ISBN 978-0-395-46726-8
  14. ^ Mullarney, K., Svensson, L, Zetterstrom, D., & Grant, P.J. (1999) Collins Bird Guide. HarperCollins Publishers Ltd., London p. 14
  15. ^ del Hoyo, et al., Handbook of the Birds of the World. Volume 1: Ostrich to Ducks (Handbooks of the Birds of the World). Lynx Edicions (1992), ISBN 978-84-87334-10-8
  16. ^ CRC Handbook of Avian Body Masses by John B. Dunning Jr. (Editor). CRC Press (1992), ISBN 978-0-8493-4258-5.
  17. ^ Sears, J. (1989). Feeding activity and body condition of mute swans Cygnus olor in rural and urban areas of a lowland river system. Wildfowl, 40(40), 88–98.
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  29. ^ Xu, Gang (2021). Great Mute Swans: Habitat, Behavior, Faith, and Family. Boston. pp. 114–116. ISBN 978-1-7365193-0-1.
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Mute swan: Brief Summary

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Sound from wingbeats.

The mute swan (Cygnus olor) is a species of swan and a member of the waterfowl family Anatidae. It is native to much of Eurosiberia, and (as a rare winter visitor) the far north of Africa. It is an introduced species in North America, home to the largest populations outside of its native range, with additional smaller introductions in Australasia and southern Africa. The name 'mute' derives from it being less vocal than other swan species. Measuring 125 to 160 cm (49 to 63 in) in length, this large swan is wholly white in plumage with an orange beak bordered with black. It is recognizable by its pronounced knob atop the beak, which is larger in males.

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