Ilex umbellulata (Wall.) Loes.

Shrubs or trees, evergreen, 4-15(-18) m tall; trunk to 40 cm d.b.h. Young branchlets brown or castaneous, glabrous, plicate, angular; older branchlets gray, terete, plicate, with raised leaf scars; terminal buds conical, small, glabrous. Stipules subulate-triangular, minute; petiole 1-1.2 cm, adaxially sulcate, glabrous; leaf blade olivaceous, opaque on both surfaces, oblong or ovate-oblong, 7-15 × (3.5-)5-6 cm, papery, membranous when young, midvein impressed adaxially, lateral veins 9-12 pairs, slightly raised abaxially, evident adaxially, anastomosing near margin, reticulate veins distinct on both surfaces, base rounded or obtuse, margin entire, narrowly recurved, apex abruptly acuminate or obtuse. Inflorescences: cymes of order 3-5, umbelliform, solitary, axillary on basal parts of current year’s branchlets, appearing paniculate when branchlets fail to develop. Male inflorescences: 6-20(-30)-flowered; peduncles 1.8-2.5 cm, puberulent; pedicels 3-5 mm, puberulent; bracteoles basal, deltoid, puberulent, apex acute; flowers white, 4- or 5-merous; calyx patelliform, ca. 2 mm in diam., puberulent, deeply 4- or 5-lobed, lobes ovate, ciliate; corolla rotate, ca. 5 mm in diam., petals oblong, ca. 2 × 1.5 mm, basally slightly connate; stamens equaling or slightly longer than petals, anthers ovoid-oblong; rudimentary ovary shorter than petals. Female flowers: sterile anthers sagittate; ovary ovoid, style present, stigma 4- or 5-lobed. Fruit red, globose, 5-7 mm in diam.; peduncles 1-2 cm, slightly stout, longitudinally angular and sulcate, puberulent; fruiting pedicel 3-5 mm, puberulent; persistent calyx explanate, ca. 4 mm in diam., puberulent, lobes semicircular, ciliate; persistent stigma thickly discoid, center depressed, rotate-lobed; pyrenes 6-10, oblong, ca. 4 mm, 1-1.5 mm in diam., abaxially longitudinally 2- or 3-striate and 1- or 2-sulcate, laterally smooth, endocarp woody. Fl. May-Sep, fr. Jul-Nov.

Yunnan [Bangladesh, India, Myanmar, Thailand, Vietnam].

Evergreen broad-leaved forests, sparse forests on slopes; 500-1700 m.

Ehretia umbellulata Wallich in Roxburgh, Fl. Ind. 2: 344. 1824; Ilex godajam (Colebrooke) J. D. Hooker var. sulcata (Wallich ex J. D. Hooker) Kurz; I. sulcata Wallich ex J. D. Hooker; I. umbellulata var. megalophylla Loesener; Pseudehretia umbellulata (Wallich) Turczaninow.